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<document id="02262CDE49BF9976D17766F38DAFA418" ID-DOI="10.11646/phytotaxa.428.3.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="13875873" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727821504607" checkinUser="felipe" docAuthor="Tennakoon, Danushka S., Kuo, Chang-Hsin &amp; Hyde, Kevin D." docDate="2020" docId="038C043AF96A8304FF3C64C3FE36CA7B" docLanguage="en" docName="phytotaxa.428.3.5.pdf" docOrigin="Phytotaxa 428 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.428.3.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Phaeodothis tricuspidis Syd. &amp; P. Syd., Annls" docType="treatment" docVersion="2" lastPageNumber="248" masterDocId="FFB57C42F96D830CFFB46179FF86CB17" masterDocTitle="Multi-locus phylogeny reveals Phaeodothis mori sp. nov. (Didymosphaeriaceae, Pleosporales) from dead leaves of Morus australis" masterLastPageNumber="254" masterPageNumber="241" pageNumber="248" updateTime="1727825652866" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="A3C242188F35512432A201B666D085A3">
<mods:title id="A2951D841126C4DBED4BEBD22B3504EA">Multi-locus phylogeny reveals Phaeodothis mori sp. nov. (Didymosphaeriaceae, Pleosporales) from dead leaves of Morus australis</mods:title>
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<mods:name id="4B9D58176CE5700CF8991E9F637586C7" type="personal">
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<mods:roleTerm id="5022BBEA71B76D56F9DAF9C6E3500D36">Author</mods:roleTerm>
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<mods:namePart id="A158A35CB93090A87EE2A4B66A8E906F">Tennakoon, Danushka S.</mods:namePart>
<mods:affiliation id="DE46AC238601076BDE1997D8034E15C1">Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan. &amp; School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand. &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai, 57100, Thailand. &amp; Key Laboratory for Plant Biodiversity and Biogeography of East Asia (KLPB), Kunming Institute of Botany, Chinese Academy of Science, Kunming 650201, Yunnan China.</mods:affiliation>
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</mods:name>
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<mods:roleTerm id="922FA60A5B548891DBA5D1AE568558AE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C41616C75934CA31D56F394B46CC5871">Kuo, Chang-Hsin</mods:namePart>
<mods:affiliation id="88675000BB20BEAD31F84C1A6351D655">Department of Plant Medicine, National Chiayi University, 300 Syuefu Road, Chiayi City 60004, Taiwan.</mods:affiliation>
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</mods:name>
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</mods:role>
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<mods:namePart id="46039B9916670F2271A0FB06FBDF68B7">Hyde, Kevin D.</mods:namePart>
<mods:affiliation id="C74638043CE1698B6C8CCC607F0DE4F9">School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand. &amp; Center of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai, 57100, Thailand. &amp; Key Laboratory for Plant Biodiversity and Biogeography of East Asia (KLPB), Kunming Institute of Botany, Chinese Academy of Science, Kunming 650201, Yunnan China. &amp; Department of Biology, Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="561F9CDD86A92C11FFB1F08A45796015">text</mods:typeOfResource>
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<mods:title id="223D5F6F875FB2BA69449A9AD8338D71">Phytotaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="AA6A294016B116FFFFF2E210C55F0B8D">
<mods:date id="406E950E044AD7B3EED01C7C82CBFC2A">2020</mods:date>
<mods:detail id="C08E5AAED7B50CCE0AE0E8E4735D424C" type="pubDate">
<mods:number id="86A43DF190DAB1E88027C63DE07A0DDB">2020-01-14</mods:number>
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</mods:detail>
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<mods:detail id="6550544AE746F0302DB279826CB95312" type="volume">
<mods:number id="C5166CD89AA101582E52CFE461F991E8">428</mods:number>
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</mods:detail>
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<mods:detail id="C21CBE31245C55F3C0106D37CB2185AD" type="issue">
<mods:number id="67BF409B3D00BEFC48858DD6500B7E1D">3</mods:number>
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</mods:detail>
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<mods:extent id="3929869CD1C2D7810887828958563491" unit="page">
<mods:start id="0E9F790C3940D0AE2DDC343E1034E921">241</mods:start>
<mods:end id="EF3E1714B0800FAD55EB2C97885774D4">254</mods:end>
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<mods:location id="F7433DC46B98C2204097779F509EC73A">
<mods:url id="8B388622CFACAB1B66B532299B31F702">http://dx.doi.org/10.11646/phytotaxa.428.3.5</mods:url>
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</mods:location>
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<mods:classification id="10877BECBE09EFEE63DABFA32C17A66E">journal article</mods:classification>
<mods:identifier id="2454CA6A32A19930A7637172177CFDBF" type="DOI">10.11646/phytotaxa.428.3.5</mods:identifier>
<mods:identifier id="D1A77FF7DA29D1BCACF6E112973EE6EB" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="983F3440AE96AAA810E55D8B0E6C8741" type="Zenodo-Dep">13875873</mods:identifier>
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</mods:mods>
<treatment id="038C043AF96A8304FF3C64C3FE36CA7B" LSID="urn:lsid:plazi:treatment:038C043AF96A8304FF3C64C3FE36CA7B" httpUri="http://treatment.plazi.org/id/038C043AF96A8304FF3C64C3FE36CA7B" lastPageId="8" lastPageNumber="248" pageId="7" pageNumber="248">
<subSubSection id="C33FE6A7F96A830BFF3C64C3FC29CEEF" pageId="7" pageNumber="248" type="nomenclature">
<paragraph id="8B9AB52CF96A830BFF3C64C3FC22CEC2" blockId="7.[136,932,1466,1493]" box="[136,932,1466,1493]" pageId="7" pageNumber="248">
<heading id="D0D20240F96A830BFF3C64C3FC22CEC2" box="[136,932,1466,1493]" fontSize="11" level="2" pageId="7" pageNumber="248" reason="3">
<taxonomicName id="4C25CEAFF96A830BFF3C64C3FD16CEC2" ID-CoL="4FMYY" authority="Syd. &amp; P. Syd., Annls" authorityName="Syd. &amp; P. Syd., Annls" box="[136,656,1466,1493]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF96A830BFF3C64C3FE10CEC2" bold="true" box="[136,406,1466,1493]" italics="true" pageId="7" pageNumber="248">Phaeodothis tricuspidis</emphasis>
Syd. &amp; P. Syd., Annls
</taxonomicName>
mycol. 2(2): 166 (1904)
</heading>
</paragraph>
<paragraph id="8B9AB52CF96A830BFF3C649BFC29CEEF" blockId="7.[136,943,1506,1528]" box="[136,943,1506,1528]" pageId="7" pageNumber="248">
Index Fungorum Number: IF 185110; Facesoffungi number: FoF 00060,
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<figureCitation id="131EA9A9F96A830BFCF3649AFC29CEEF" box="[839,943,1507,1528]" captionStart="FIGURE 3" captionStartId="8.[136,229,1439,1461]" captionTargetBox="[154,1433,389,1408]" captionTargetId="figure-114@8.[151,1436,386,1411]" captionTargetPageId="8" captionText="FIGURE 3. Phaeodothis tricuspidis (SF125876, holotype) a. Herbarium packet and specimen. b. Ascomata on host. c. Close up of ascoma. d, e. Sections through ascomata. f. Peridium comprising hyaline compressed cells. g–j. Mature and immature asci surrounded by pseudoparaphyses. k–n. ascospores. Scale bars: c = 1000 µm, d, e = 100 µm, f = 20 µm, g–j = 30 µm, k–n = 5 µm." figureDoi="http://doi.org/10.5281/zenodo.13875879" httpUri="https://zenodo.org/record/13875879/files/figure.png" pageId="7" pageNumber="248">FIGURE 3</figureCitation>
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</paragraph>
</subSubSection>
<subSubSection id="C33FE6A7F96A8304FF3C675EFE36CA7B" lastPageId="8" lastPageNumber="249" pageId="7" pageNumber="248" type="description">
<paragraph id="8B9AB52CF96A830BFF3C675EFB47CC76" blockId="7.[136,1452,1573,2069]" pageId="7" pageNumber="248">
<emphasis id="B951693EF96A830BFF3C675EFF6ACD56" box="[136,236,1575,1601]" italics="true" pageId="7" pageNumber="248">Saprobic</emphasis>
on dead wood or grasses.
<emphasis id="B951693EF96A830BFD80675FFD5DCD56" bold="true" box="[564,731,1574,1601]" italics="true" pageId="7" pageNumber="248">Sexual morph</emphasis>
:
<emphasis id="B951693EF96A830BFD44675EFCD8CD56" box="[752,862,1575,1601]" italics="true" pageId="7" pageNumber="248">Ascomata</emphasis>
220–300×120−180 μm (
<emphasis id="B951693EF96A830BFB3E675CFB11CD2C" box="[1162,1175,1573,1595]" italics="true" pageId="7" pageNumber="248">x</emphasis>
= 255×137 μm, n=10), immersed to nearly superficial under clypeus, scattered to gregarious, subglobose to hemispherical, dark brown to black, papillate.
<emphasis id="B951693EF96A830BFEF16716FE11CD9E" box="[325,407,1647,1673]" italics="true" pageId="7" pageNumber="248">Papilla</emphasis>
black, with pore-like ostioles, ostiolar canal filled with periphyses.
<taxonomicName id="4C25CEAFF96A830BFB106716FA8ACD9E" box="[1188,1292,1647,1673]" class="Magnoliopsida" family="Peraceae" genus="Peridium" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="genus">
<emphasis id="B951693EF96A830BFB106716FA8ACD9E" box="[1188,1292,1647,1673]" italics="true" pageId="7" pageNumber="248">Peridium</emphasis>
</taxonomicName>
15−25μm (
<emphasis id="B951693EF96A830BFA2C6714FA23CD94" box="[1432,1445,1645,1667]" italics="true" pageId="7" pageNumber="248">x</emphasis>
=23μm, n=20) wide, comprising several layers of hyaline to light brown, compressed cells of
<emphasis id="B951693EF96A830BFB2367EDFADECDBA" box="[1175,1368,1683,1709]" italics="true" pageId="7" pageNumber="248">
textura
<taxonomicName id="4C25CEAFF96A830BFB5967EAFADECDBA" box="[1261,1368,1683,1709]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Paraphaeosphaeria" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="angularis">angularis</taxonomicName>
</emphasis>
, fusing and indistinguishable from the host tissues.
<emphasis id="B951693EF96A830BFD3167CEFCA4CDC6" box="[645,802,1719,1745]" italics="true" pageId="7" pageNumber="248">Hamathecium</emphasis>
of 2−4 μm wide (
<emphasis id="B951693EF96A830BFC4A67CCFB8DCDDC" box="[1022,1035,1717,1739]" italics="true" pageId="7" pageNumber="248">x</emphasis>
= 2.6 μm, n=20), aseptate, cellular pseudoparaphyses often embedded in a gelatinous matrix.
<emphasis id="B951693EF96A830BFCBB67A2FCB9CDE2" box="[783,831,1755,1781]" italics="true" pageId="7" pageNumber="248">Asci</emphasis>
50 − 80 × 7−12 μm (
<emphasis id="B951693EF96A830BFB9867A0FBBFCDF8" box="[1068,1081,1753,1775]" italics="true" pageId="7" pageNumber="248">x</emphasis>
= 69 × 10 μm, n = 20), 8-spored, bitunicate, fissitunicate, cylindrical, short pedicellate, apically rounded with a minute ocular chamber.
<emphasis id="B951693EF96A830BFA996786FA2ACC0E" box="[1325,1452,1791,1817]" italics="true" pageId="7" pageNumber="248">Ascospores</emphasis>
14−17 × 4−5μm (
<emphasis id="B951693EF96A830BFEE16658FEE4CC20" box="[341,354,1825,1847]" italics="true" pageId="7" pageNumber="248">x</emphasis>
= 15.8 × 4.5 μm, n=30), unit to biseriate and partially overlapping, fusiform with rounded ends, olivaceous brown, 1-septate, constricted at the septa, smooth-walled.
<emphasis id="B951693EF96A830BFC26663FFBC4CC76" bold="true" box="[914,1090,1862,1889]" italics="true" pageId="7" pageNumber="248">Asexual morph</emphasis>
: unknown.
</paragraph>
<paragraph id="8B9AB52CF96A830BFF096612FD49CCBE" blockId="7.[136,1452,1573,2069]" pageId="7" pageNumber="248">
Material examined:
<emphasis id="B951693EF96A830BFE2A6612FE3CCC92" bold="true" box="[414,442,1899,1925]" pageId="7" pageNumber="248">—</emphasis>
<collectingCountry id="F332F5BCF96A830BFE0E6612FDE6CC92" box="[442,608,1899,1925]" name="Argentina" pageId="7" pageNumber="248">ARGENTINA</collectingCountry>
, San José,
<collectingRegion id="49E17BCEF96A830BFD5F6612FCA0CC92" box="[747,806,1899,1925]" country="Argentina" name="Salta" pageId="7" pageNumber="248">Salta</collectingRegion>
, on leaves of
<taxonomicName id="4C25CEAFF96A830BFC626615FA87CC92" authority="Griseb." authorityName="Griseb." box="[982,1281,1899,1925]" class="Magnoliopsida" family="Elaeocarpaceae" genus="Tricuspis" kingdom="Plantae" order="Oxalidales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B951693EF96A830BFC626615FB25CC92" box="[982,1187,1899,1925]" italics="true" pageId="7" pageNumber="248">Tricuspis latifolia</emphasis>
Griseb.
</taxonomicName>
(
<taxonomicName id="4C25CEAFF96A830BFAA16612FAF1CC92" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1301,1399,1899,1925]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
),
<date id="FF9B93ECF96A830BFA3B6612FEA9CCBE" pageId="7" pageNumber="248" value="1873-02-12">12 February 1873</date>
, P.G. Lorentz (SF125876,
<emphasis id="B951693EF96A830BFDEC66F6FD39CCBE" bold="true" box="[600,703,1935,1961]" pageId="7" pageNumber="248">
<typeStatus id="549E0B8EF96A830BFDEC66F6FD39CCBE" box="[600,703,1935,1961]" pageId="7" pageNumber="248" type="holotype">holotype</typeStatus>
</emphasis>
).
</paragraph>
<paragraph id="8B9AB52CF96A8304FF0966CAFE36CA7B" blockId="7.[136,1452,1573,2069]" lastBlockId="8.[136,1452,158,365]" lastPageId="8" lastPageNumber="249" pageId="7" pageNumber="248">
Notes:—
<taxonomicName id="4C25CEAFF96A830BFE9666CAFDA8CCDA" box="[290,558,1971,1997]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF96A830BFE9666CAFDA8CCDA" box="[290,558,1971,1997]" italics="true" pageId="7" pageNumber="248">Phaeodothis tricuspidis</emphasis>
</taxonomicName>
was introduced by
<bibRefCitation id="EFB4C8DDF96A830BFCA566CAFBBBCCDA" author="Sydow, H. &amp; Sydow, P." box="[785,1085,1971,1997]" pageId="7" pageNumber="248" pagination="162 - 174" refId="ref10677" refString="Sydow, H. &amp; Sydow, P. (1904) Novae fungorum species. Ann. Mycol. 2: 162 - 174." type="journal article" year="1904">Sydow and Sydow (1904)</bibRefCitation>
, from living leaves of
<taxonomicName id="4C25CEAFF96A830BFAF166CDFEB8CCE6" authority="Griseb." authorityName="Griseb." class="Magnoliopsida" family="Elaeocarpaceae" genus="Tricuspis" kingdom="Plantae" order="Oxalidales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B951693EF96A830BFAF166CDFF67CCE6" italics="true" pageId="7" pageNumber="248">Tricuspis latifolia</emphasis>
Griseb.
</taxonomicName>
(
<taxonomicName id="4C25CEAFF96A830BFEFB66AEFE28CCE6" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[335,430,2007,2033]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="7" pageNumber="248" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) and they assumed it to be very close to those of
<taxonomicName id="4C25CEAFF96A830BFBB666AEFBFCCCE6" box="[1026,1146,2007,2033]" class="Dothideomycetes" family="Polystomellaceae" genus="Dothidella" kingdom="Fungi" order="Asterinales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="genus">
<emphasis id="B951693EF96A830BFBB666AEFBFCCCE6" box="[1026,1146,2007,2033]" italics="true" pageId="7" pageNumber="248">Dothidella</emphasis>
</taxonomicName>
, but differ in having dark ascospores (
<bibRefCitation id="EFB4C8DDF96A830BFEA76682FDAAC302" author="Malloch, D. &amp; Mallik, A." box="[275,556,2043,2069]" pageId="7" pageNumber="248" pagination="1265 - 1275" refId="ref8890" refString="Malloch, D. &amp; Mallik, A. (1998) Taxonomy of Orphnodactylis kalmiae gen. et sp. nov. influenced by the hyperparasite Didymosphaeria kalmiae. Canadian journal of botany 76: 1265 - 1275. https: // doi. org / 10.1139 / b 98 - 147" type="journal article" year="1998">Malloch &amp; Mallik 1998</bibRefCitation>
).
<taxonomicName id="4C25CEAFF96A830BFDF66682FC8BC302" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[578,781,2043,2069]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF96A830BFDF66682FC8BC302" box="[578,781,2043,2069]" italics="true" pageId="7" pageNumber="248">Phaeodothis mori</emphasis>
</taxonomicName>
can be distinguished from the
<typeStatus id="549E0B8EF96A830BFBC36682FB21C302" box="[1143,1191,2043,2069]" pageId="7" pageNumber="248">type</typeStatus>
species,
<taxonomicName id="4C25CEAFF96A830BFAA76685FA2DC302" box="[1299,1451,2043,2069]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="7" pageNumber="248" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF96A830BFAA76685FAA2C302" box="[1299,1316,2044,2069]" italics="true" pageId="7" pageNumber="248">P</emphasis>
.
<emphasis id="B951693EF96A830BFA816682FA2DC302" box="[1333,1451,2043,2069]" italics="true" pageId="7" pageNumber="248">tricuspidis</emphasis>
</taxonomicName>
in having smaller ascomata (100–150 × 80–130 μm) and yellowish brown, ascospores (10.5–12.5 × 3–4 μm) with mucilaginous sheath, whereas
<taxonomicName id="4C25CEAFF9658304FE5861BAFD03CBCB" box="[492,645,194,221]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="8" pageNumber="249" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF9658304FE5861BAFE7BCBCB" box="[492,509,195,220]" italics="true" pageId="8" pageNumber="249">P</emphasis>
.
<emphasis id="B951693EF9658304FDBB61BBFD03CBCB" box="[527,645,194,220]" italics="true" pageId="8" pageNumber="249">tricuspidis</emphasis>
</taxonomicName>
has larger ascomata (220–300×120−180 μm) and olivaceous brown, ascospores (14−17 × 4−5μm) lacking a mucilaginous sheath. As well as,
<taxonomicName id="4C25CEAFF9658304FC7F619FFB53CA17" box="[971,1237,230,256]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="8" pageNumber="249" phylum="Ascomycota" rank="species" species="tricuspidis">
<emphasis id="B951693EF9658304FC7F619FFB53CA17" box="[971,1237,230,256]" italics="true" pageId="8" pageNumber="249">Phaeodothis tricuspidis</emphasis>
</taxonomicName>
has been recorded from dead grass stems (
<taxonomicName id="4C25CEAFF9658304FE2E6073FE7ACA33" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[410,508,266,292]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="249" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
), which is a monocot; whereas
<taxonomicName id="4C25CEAFF9658304FCDE6072FC47CA33" authorityName="Tennakoon, C. H. Kuo &amp; K. D. Hyde" authorityYear="2020" box="[874,961,267,293]" class="Dothideomycetes" family="Didymosphaeriaceae" genus="Phaeodothis" kingdom="Fungi" order="Pleosporales" pageId="8" pageNumber="249" phylum="Ascomycota" rank="species" species="mori">
<emphasis id="B951693EF9658304FCDE6072FCFDCA33" box="[874,891,267,292]" italics="true" pageId="8" pageNumber="249">P</emphasis>
.
<emphasis id="B951693EF9658304FC386072FC47CA33" box="[908,961,267,292]" italics="true" pageId="8" pageNumber="249">mori</emphasis>
</taxonomicName>
was recorded from the decaying leaves of
<taxonomicName id="4C25CEAFF9658304FF3C6056FEBCCA5F" box="[136,314,302,328]" class="Magnoliopsida" family="Moraceae" genus="Morus" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="249" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B951693EF9658304FF3C6056FEBCCA5F" box="[136,314,302,328]" italics="true" pageId="8" pageNumber="249">Morus australis</emphasis>
</taxonomicName>
(
<taxonomicName id="4C25CEAFF9658304FEFD6057FE38CA5F" box="[329,446,302,328]" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="249" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
), which is a dicot. However, molecular data are essentially needed to clarify phylogenetic placement of
<typeStatus id="549E0B8EF9658304FE94602AFED6CA7A" box="[288,336,339,365]" pageId="8" pageNumber="249">type</typeStatus>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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