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<document id="E0CD2969CACEB74119581CA56DDE293E" ID-DOI="10.5281/zenodo.171388" ID-GBIF-Dataset="164b0a72-f861-4de7-a939-a3ad42b5c71d" ID-ISSN="1175­5326" ID-Zenodo-Dep="171388" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459721013543" checkinUser="plazi" docAuthor="Boucher, Stéphanie" docDate="2005" docId="03839D7E230AFFECFEBEFEC3FCEC207A" docLanguage="en" docName="zt00993.pdf" docOrigin="Zootaxa 993" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Cerodontha (Dizygomyza) nigrihalterata Boucher, 2005, sp. nov." docType="treatment" docVersion="10" lastPageNumber="4" masterDocId="FFBAE5062308FFE8FFB6FFDFFFDB222B" masterDocTitle="Description of an unusual new Costa Rican species of Cerodontha (Dizygomyza) with additional notes on Neotropical species of Dizygomyza (Diptera: Agromyzidae)" masterLastPageNumber="8" masterPageNumber="1" pageNumber="3" updateTime="1698209392311" updateUser="plazi">
<mods:mods id="547C4CA23CF9A8A0328A9F5D105CA441" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2592464C0B1E5D1D0946B90A72F32B6A">
<mods:title id="0FCA5C9FFAF249E89A355CBD1F37DF2B">Description of an unusual new Costa Rican species of Cerodontha (Dizygomyza) with additional notes on Neotropical species of Dizygomyza (Diptera: Agromyzidae)</mods:title>
</mods:titleInfo>
<mods:name id="DD15DA6B14D2D355273F3F8AE1A05960" type="personal">
<mods:role id="BDBC617DA65AA7F14F090A98B1094CC6">
<mods:roleTerm id="64426A7DF2CC91AA24B0FD45489038A0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="815D6099361D30B40FC5C0DB3D44FC3A">Boucher, Stéphanie</mods:namePart>
</mods:name>
<mods:typeOfResource id="ADA07F8F92E3261081BE2B8140850C53">text</mods:typeOfResource>
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<mods:title id="ED57A0761366D4F89FA0FA0126713961">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6EE0D1A8D6E55C0AB6F6C1F60F342EB4">
<mods:date id="297A00A7A204ED1AAEB42E06B427DA35">2005</mods:date>
<mods:detail id="2B7653176AB46166255612527CFC518E" type="volume">
<mods:number id="DB18D040E7104873E7CE0A71853C8427">993</mods:number>
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<mods:start id="559D7E31E65DD56880BFB3A77347402D">1</mods:start>
<mods:end id="4E696A89958DCA2454750A3DC8821DF3">8</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="89D4B02148F346919CFCC56C2E691FDC">journal article</mods:classification>
<mods:identifier id="42F6844EE9F29F54E952A5A578560F31" type="DOI">10.5281/zenodo.171388</mods:identifier>
<mods:identifier id="A8251FC58691133C9F3C68C5F5F36E9F" type="GBIF-Dataset">164b0a72-f861-4de7-a939-a3ad42b5c71d</mods:identifier>
<mods:identifier id="F4827BBB7C8A461225F29C9665862916" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="1931B1F3BFAAD14B62B1FBED14CBEE9E" type="Zenodo-Dep">171388</mods:identifier>
</mods:mods>
<treatment id="03839D7E230AFFECFEBEFEC3FCEC207A" ID-DOI="http://doi.org/10.5281/zenodo.6265861" ID-GBIF-Taxon="119335632" ID-Zenodo-Dep="6265861" LSID="urn:lsid:plazi:treatment:03839D7E230AFFECFEBEFEC3FCEC207A" httpUri="http://treatment.plazi.org/id/03839D7E230AFFECFEBEFEC3FCEC207A" lastPageId="4" lastPageNumber="4" pageId="2" pageNumber="3">
<subSubSection id="C3307FE3230AFFEAFEBEFEC3FEA72372" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="8B952C68230AFFEAFEBEFEC3FC8C231E" blockId="2.[264,855,283,345]" box="[264,855,283,310]" pageId="2" pageNumber="3">
<heading id="D0DD9B04230AFFEAFEBEFEC3FC8C231E" bold="true" box="[264,855,283,310]" fontSize="11" level="1" pageId="2" pageNumber="3" reason="1">
<emphasis id="B95EF07A230AFFEAFEBEFEC3FC8C231E" bold="true" box="[264,855,283,310]" pageId="2" pageNumber="3">
<taxonomicName id="4C2A57EB230AFFEAFEBEFEC3FD2A231D" ID-CoL="8BF25" box="[264,753,284,310]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nigrihalterata" status="sp. nov." subGenus="Dizygomyza">
<emphasis id="B95EF07A230AFFEAFEBEFEC3FD2A231D" bold="true" box="[264,753,284,310]" italics="true" pageId="2" pageNumber="3">Cerodontha (Dizygomyza) nigrihalterata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D4D01230AFFEAFD4FFEC4FC8C231E" box="[761,855,283,309]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B952C68230AFFEAFEBEFEE0FEA72372" blockId="2.[264,855,283,345]" box="[264,380,319,345]" pageId="2" pageNumber="3">
(
<figureCitation id="131130ED230AFFEAFEA7FEE0FEAF2372" box="[273,372,319,345]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,383,1387,1411]" captionTargetBox="[393,1219,615,1361]" captionTargetId="figure@3.[364,1250,602,1382]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Cerodontha (Dizygomyza) nigrihalterata sp. nov. 1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing." httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="2" pageNumber="3">Fig. 1–7</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3307FE3230AFFECFEBEFE50FCEC207A" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="8B952C68230AFFEAFEBEFE50FC4623D2" blockId="2.[264,1324,399,1870]" pageId="2" pageNumber="3">
<typeStatus id="549192CA230AFFEAFEBEFE50FE992382" box="[264,322,399,425]" pageId="2" pageNumber="3">Type</typeStatus>
material:
<typeStatus id="549192CA230AFFEAFE77FE50FDE92382" box="[449,562,399,425]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
. Male.
<collectingCountry id="F33D6CF8230AFFEAFD27FE50FC9D2382" box="[657,838,399,425]" name="Costa Rica" pageId="2" pageNumber="3">COSTA RICA</collectingCountry>
: Puntarenas: Golfito, R.F. Golfo Dulce, Estación Agujas, Sendera Ajo,
<quantity id="4CD2818D230AFFEAFD01FE68FCE323FA" box="[695,824,439,465]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="2" pageNumber="3" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">
250–
<specimenCount id="9D2CE7E1230AFFEAFD42FE68FCE323FA" box="[756,824,439,465]" pageId="2" pageNumber="3" type="generic">350m</specimenCount>
</quantity>
.
<date id="FF940AA8230AFFEAFCE5FE68FBDA23FA" box="[851,1025,439,465]" pageId="2" pageNumber="3" value="1998-11-02">2 Nov 1998</date>
. M. Umana. Manual. L_S_276750_526550 #56934, INB0003308651(
<collectionCode id="ED3BB4AD230AFFEAFCF5FE00FC4E23D2" box="[835,917,479,505]" pageId="2" pageNumber="3">INBIO</collectionCode>
)
</paragraph>
<paragraph id="8B952C68230AFFEAFE8EFDD8FC91214A" blockId="2.[264,1324,399,1870]" pageId="2" pageNumber="3">
<typeStatus id="549192CA230AFFEAFE8EFDD8FE75200A" box="[312,430,519,545]" pageId="2" pageNumber="3" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="F33D6CF8230AFFEAFE0EFDD8FDB3200A" box="[440,616,519,545]" name="Costa Rica" pageId="2" pageNumber="3">COSTA RICA</collectingCountry>
: Puntarenas: Est. Agujas. Send. Elvia.
<specimenCount id="9D2CE7E1230AFFEAFB98FDD8FBA3200A" box="[1070,1144,519,545]" pageId="2" pageNumber="3" type="generic">
<quantity id="4CD2818D230AFFEAFB98FDD8FBAF200A" box="[1070,1140,519,545]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="3" unit="m" value="300.0">300m</quantity>
.
</specimenCount>
<date id="FF940AA8230AFFEAFBC9FDD8FAD6200A" box="[1151,1293,519,545]" pageId="2" pageNumber="3" value="1998-11-03">3 Nov 1998</date>
. J. Matarrita, I. Lopez, J. Gutierrez. L_S_276750_526550 #51810 INB0003021049 (1M,
<collectionCode id="ED3BB4AD230AFFEAFEBEFD88FE85205A" box="[264,350,599,625]" pageId="2" pageNumber="3">INBIO</collectionCode>
); Golfito, R.F. Golfo Dulce, Estación Agujas, Sendera Ajo,
<quantity id="4CD2818D230AFFEAFBFAFD88FB16205A" box="[1100,1229,599,625]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="2" pageNumber="3" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">
250–
<specimenCount id="9D2CE7E1230AFFEAFB3EFD88FB16205A" box="[1160,1229,599,625]" pageId="2" pageNumber="3" type="generic">350m</specimenCount>
</quantity>
.
<date id="FF940AA8230AFFEAFB6BFD88FE9D20B2" pageId="2" pageNumber="3" value="1998-11-02">2 Nov 1998</date>
. M. Umana. Manual. L_S_276750_526550 #56934, INB0003308650 (1F,
<collectionCode id="ED3BB4AD230AFFEAFB71FDA0FAC620B2" box="[1223,1309,639,665]" pageId="2" pageNumber="3">INBIO</collectionCode>
); same except INB0003308658 (1M,
<collectionCode id="ED3BB4AD230AFFEAFD1BFD78FCD820EA" box="[685,771,679,705]" pageId="2" pageNumber="3">INBIO</collectionCode>
);
<quantity id="4CD2818D230AFFEAFCAEFD78FC8820EA" box="[792,851,679,705]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="3" unit="km" value="3.0">3 km</quantity>
SW of Rincon,
<geoCoordinate id="EE1E4AAF230AFFEAFBA4FD78FBB120EA" box="[1042,1130,679,705]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="9.916667">9°55’N</geoCoordinate>
,
<geoCoordinate id="EE1E4AAF230AFFEAFBC1FD78FB3920EA" box="[1143,1250,679,705]" direction="west" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="-84.21667">84°13’W</geoCoordinate>
,
<quantity id="4CD2818D230AFFEAFB59FD78FAFD20EA" box="[1263,1318,679,705]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="10.0">10m</quantity>
,
<date id="FF940AA8230AFFEAFEBEFD10FE6C20C2" box="[264,439,719,745]" pageId="2" pageNumber="3" value="1990-10" valueMax="1990-12-31" valueMin="1990-10-00">Oct–Dec 1990</date>
, malaise trap, P. Hanson (1F,
<collectionCode id="ED3BB4AD230AFFEAFC97FD10FCA020C2" LSID="urn:lsid:biocol.org:col:34871" box="[801,891,719,745]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">USNM</collectionCode>
); R.F. Golfo Dulce,
<quantity id="4CD2818D230AFFEAFBC7FD10FB7E20C2" box="[1137,1189,719,745]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="3" unit="km" value="5.0">5km</quantity>
W Piedras Blancas,
<quantity id="4CD2818D230AFFEAFEC3FD28FE61213A" box="[373,442,759,785]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="100.0">100m</quantity>
, malaise trap,
<date id="FF940AA8230AFFEAFDD0FD28FD62213A" box="[614,697,759,785]" pageId="2" pageNumber="3" value="1991-10">x.1991</date>
, P. Hanson (1F, LEM); Peninsula Osa,
<quantity id="4CD2818D230AFFEAFB3DFD28FB0D213A" box="[1163,1238,759,785]" metricMagnitude="4" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="3" unit="km" value="23.0">23 km</quantity>
N Pto. Jimenez, La Palma,
<quantity id="4CD2818D230AFFEAFE4CFCC0FDEA2112" box="[506,561,799,825]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="m" value="10.0">10m</quantity>
, Malaise trap,
<date id="FF940AA8230AFFEAFD54FCC0FC802112" box="[738,859,799,825]" pageId="2" pageNumber="3" value="1990-10" valueMax="1990-11-31" valueMin="1990-10-00">x–xi.1990</date>
, P. Hanson (1F, LEM); Cartago: Turrialba,
<date id="FF940AA8230AFFEAFEF1FC98FE56214A" box="[327,397,839,865]" pageId="2" pageNumber="3" value="1915-07">vii­15</date>
­19.1965, P.J. Spangler (1M,
<collectionCode id="ED3BB4AD230AFFEAFD52FC98FCE5214A" LSID="urn:lsid:biocol.org:col:34871" box="[740,830,839,865]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="2" pageNumber="3">USNM</collectionCode>
).
</paragraph>
<paragraph id="8B952C68230AFFEAFE8EFCB0FC3321F2" blockId="2.[264,1324,399,1870]" pageId="2" pageNumber="3">
Diagnosis. This species can be differentiated from all other
<taxonomicName id="4C2A57EB230AFFEAFC43FCB0FAF121A3" authority="(Dizygomyza)" baseAuthorityName="Dizygomyza" box="[1013,1322,879,904]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B95EF07A230AFFEAFC43FCB0FAF121A3" box="[1013,1322,879,904]" italics="true" pageId="2" pageNumber="3">Cerodontha (Dizygomyza)</emphasis>
</taxonomicName>
species by the combination of the following characters: black halter, the presence of only one
<emphasis id="B95EF07A230AFFEAFE8CFC60FE8421F3" box="[314,351,959,984]" italics="true" pageId="2" pageNumber="3">ors</emphasis>
and wings conspicuously infuscated in cell r1 and r2+3.
</paragraph>
<paragraph id="8B952C68230AFFEAFE8EFC33FDD52655" blockId="2.[264,1324,399,1870]" pageId="2" pageNumber="3">Description. Head including antenna, brown, gena paler, yellowish­brown; epistoma and facial ridge yellowish. Thorax uniformly brown, calypter including margin white, fringe brown. Halter completely brown. Legs brown, except foreknee and tarsi pale yellow. Abdomen brown.</paragraph>
<paragraph id="8B952C68230AFFEBFE8EFB53FC9D201D" blockId="2.[264,1324,399,1870]" lastBlockId="3.[264,1323,284,566]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
Frons width
<quantity id="4CD2818D230AFFEAFE62FB53FD5E268D" box="[468,645,1164,1190]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="2" pageNumber="3" unit="mm" value="0.375" valueMax="0.4" valueMin="0.35">0.35–0.40 mm</quantity>
; ratio of frons width to eye width 1.6–1.8; orbit broad, 0.22–0.27 width of frons at midpoint; orbit widening anteriorly (
<figureCitation id="131130ED230AFFEAFBBCFB6BFB8F26E5" box="[1034,1108,1204,1230]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,383,1387,1411]" captionTargetBox="[393,1219,615,1361]" captionTargetId="figure@3.[364,1250,602,1382]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Cerodontha (Dizygomyza) nigrihalterata sp. nov. 1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing." httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="2" pageNumber="3">Fig. 6</figureCitation>
); frons projecting above and in front of eye in profile, orbit forming only narrow ring below eye; eye slightly pubescent; lunule and anterior margin of orbit with patch of silvery pubescence best seen in posterodorsal view; orbit and ocellar triangle shining; frons with microtomentum; usually 3
<emphasis id="B95EF07A230AFFEAFEE7FA8BFEA82746" box="[337,371,1364,1389]" italics="true" pageId="2" pageNumber="3">ori</emphasis>
and 1
<emphasis id="B95EF07A230AFFEAFE77FA8BFE3C2746" box="[449,487,1364,1389]" italics="true" pageId="2" pageNumber="3">ors</emphasis>
(one aberrant specimen with 3
<emphasis id="B95EF07A230AFFEAFCE1FA8BFCA72746" box="[855,892,1364,1389]" italics="true" pageId="2" pageNumber="3">ors</emphasis>
, including one displaced medially at margin of orbit and frons) anterior
<emphasis id="B95EF07A230AFFEAFD03FAA3FD0C27BE" box="[693,727,1404,1429]" italics="true" pageId="2" pageNumber="3">ori</emphasis>
a little shorter than other
<emphasis id="B95EF07A230AFFEAFBACFAA3FBE727BE" box="[1050,1084,1404,1429]" italics="true" pageId="2" pageNumber="3">ori</emphasis>
; first 2 anterior
<emphasis id="B95EF07A230AFFEAFABEFAA3FAF127BE" box="[1288,1322,1404,1429]" italics="true" pageId="2" pageNumber="3">ori</emphasis>
inclinate; posterior
<emphasis id="B95EF07A230AFFEAFE59FA7BFDCA2796" box="[495,529,1444,1469]" italics="true" pageId="2" pageNumber="3">ori</emphasis>
reclinate and slightly lateroclinate;
<emphasis id="B95EF07A230AFFEAFC0BFA7BFC392796" box="[957,994,1444,1469]" italics="true" pageId="2" pageNumber="3">ors</emphasis>
reclinate and slightly inclinate; 4–5 reclinate and/or inclinate orbital setulae, short, normal, in one row, present anteriorly (not reaching
<emphasis id="B95EF07A230AFFEAFDA0FA2BFDE02426" box="[534,571,1524,1549]" italics="true" pageId="2" pageNumber="3">ors</emphasis>
); first flagellomere slightly enlarged in male with white pubescence on lateral surface and black pubescence on medial surface, extent of white pubescence variable; first flagellomere in female with black pubescence apically and medially; arista longer than eye height with long pubescence; arista white basally in some specimens, not obviously swollen at base; gena with uniform width; gena height 0.20–0.25 times eye height; lunule broad, in the form of a semicircle, or a little higher, upper margin reaching level of first or second
<emphasis id="B95EF07A230AFFEAFD35F93BFD7E24D6" box="[643,677,1764,1789]" italics="true" pageId="2" pageNumber="3">ori</emphasis>
; bases of antennae widely separated. Three postsutural dc, anterior postsutural 0.57–0.70 times length of second postsutural; presutural dc absent; acrostichals irregular in 7–8 rows; prsc absent; one postsutural intra­alar; 1 presutural and two postsutural supra­alar; notopleuron with 2 bristles on ventral side. Midtibia with strong apico­ventral bristle, almost half as long as first tarsomere; no other bristles on fore or mid tibiae. Wing length
<quantity id="4CD2818D230BFFEBFDF2FEB3FD0A23AD" box="[580,721,364,390]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="3" pageNumber="4" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9–2.1 mm</quantity>
in males and 1.8–2.0 mm in females; R4+5 ending a little closer to wing tip than M1+2; costa extending to M1+2; wing conspicuously infuscated in cell r1 and r2+3 (
<figureCitation id="131130ED230BFFEBFE5FFE1BFDEF23F5" box="[489,564,452,478]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,383,1387,1411]" captionTargetBox="[393,1219,615,1361]" captionTargetId="figure@3.[364,1250,602,1382]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Cerodontha (Dizygomyza) nigrihalterata sp. nov. 1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing." httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
); cross­vein dm­cu present; second costal sector 3.8–4.7 times length of fourth; last section of CuA1 short: 0.48–0.56 times length of penultimate. Crossvein r­m located at 0.3–0.4 the length of cell dm.
</paragraph>
<caption id="DF557CE0230BFFEBFEBEFAB4FAD927E3" httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="3" pageNumber="4" targetBox="[393,1219,615,1361]" targetPageId="3">
<paragraph id="8B952C68230BFFEBFEBEFAB4FAD927E3" blockId="3.[264,1322,1387,1480]" pageId="3" pageNumber="4">
<emphasis id="B95EF07A230BFFEBFEBEFAB4FE6827A8" bold="true" box="[264,435,1387,1411]" pageId="3" pageNumber="4">FIGURES 1–7.</emphasis>
<taxonomicName id="4C2A57EB230BFFEBFE0FFAB3FCB427A8" box="[441,879,1388,1411]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nigrihalterata" status="sp. nov." subGenus="Dizygomyza">
<emphasis id="B95EF07A230BFFEBFE0FFAB3FCB427A8" box="[441,879,1388,1411]" italics="true" pageId="3" pageNumber="4">Cerodontha (Dizygomyza) nigrihalterata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26D4D01230BFFEBFCC2FAB3FC1D27A8" box="[884,966,1388,1411]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing.
</paragraph>
</caption>
<paragraph id="8B952C68230BFFEBFE8EFA23FBA224A5" blockId="3.[264,1325,1532,1884]" pageId="3" pageNumber="4">
Male genitalia. Distiphallus with tubules mostly parallel sided, slightly dilated at apex (
<figureCitation id="131130ED230BFFEBFEA7F9FBFEAD2415" box="[273,374,1572,1598]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,383,1387,1411]" captionTargetBox="[393,1219,615,1361]" captionTargetId="figure@3.[364,1250,602,1382]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Cerodontha (Dizygomyza) nigrihalterata sp. nov. 1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing." httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="3" pageNumber="4">Fig. 1, 2</figureCitation>
). Hypandrium broad and rounded (
<figureCitation id="131130ED230BFFEBFCA5F9FBFC832415" box="[787,856,1572,1598]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,383,1387,1411]" captionTargetBox="[393,1219,615,1361]" captionTargetId="figure@3.[364,1250,602,1382]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Cerodontha (Dizygomyza) nigrihalterata sp. nov. 1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing." httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
). Surstylus with patch of strong bristles at medioventral margin (
<figureCitation id="131130ED230BFFEBFD87F993FDA0244D" box="[561,635,1612,1638]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,383,1387,1411]" captionTargetBox="[393,1219,615,1361]" captionTargetId="figure@3.[364,1250,602,1382]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Cerodontha (Dizygomyza) nigrihalterata sp. nov. 1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing." httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
). Ejaculatory apodeme with narrow base and moderately enlarged at apex (
<figureCitation id="131130ED230BFFEBFE6EF9ABFDFA24A5" box="[472,545,1652,1678]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,383,1387,1411]" captionTargetBox="[393,1219,615,1361]" captionTargetId="figure@3.[364,1250,602,1382]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Cerodontha (Dizygomyza) nigrihalterata sp. nov. 1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing." httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
). Anal projection of epandrium prominent (
<figureCitation id="131130ED230BFFEBFB94F9ABFBB124A5" box="[1058,1130,1652,1678]" captionStart="FIGURES 1 – 7" captionStartId="3.[264,383,1387,1411]" captionTargetBox="[393,1219,615,1361]" captionTargetId="figure@3.[364,1250,602,1382]" captionTargetPageId="3" captionText="FIGURES 1 – 7. Cerodontha (Dizygomyza) nigrihalterata sp. nov. 1, phallus with hypandrium, ventral view; 2, phallus lateral view; 3, subepandrial sclerite and surstylus (ventral, left); 4, ejaculatory apodeme; 5, epandrium lateral view showing anal projection (ap); 6, head dorsal view; 7, wing." httpUri="https://zenodo.org/record/171389/files/figure.png" pageId="3" pageNumber="4">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="8B952C68230BFFEBFE8EF943FEE624F5" blockId="3.[264,1325,1532,1884]" pageId="3" pageNumber="4">Derivation of Species Name. The species name refers to the black halteres of this species.</paragraph>
<paragraph id="8B952C68230BFFECFE8EF933FCEC207A" blockId="3.[264,1325,1532,1884]" lastBlockId="4.[264,1323,284,593]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
Comments. The conspicuously infuscated cells r1 and r3+
<quantity id="4CD2818D230BFFEBFC69F926FBD3252D" box="[991,1032,1772,1802]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="4" unit="in" value="4.0">4 in</quantity>
the wings and the presence of only one
<emphasis id="B95EF07A230BFFEBFE63F8C5FE202518" box="[469,507,1818,1843]" italics="true" pageId="3" pageNumber="4">ors</emphasis>
are unique characters among New World
<taxonomicName id="4C2A57EB230BFFEBFC58F8C5FBA62518" box="[1006,1149,1818,1843]" class="Insecta" family="Agromyzidae" genus="Dizygomyza" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B95EF07A230BFFEBFC58F8C5FBA62518" box="[1006,1149,1818,1843]" italics="true" pageId="3" pageNumber="4">Dizygomyza</emphasis>
</taxonomicName>
. However, the dark halter is also present in
<taxonomicName id="4C2A57EB230BFFEBFDDDF89DFC1E2577" authority="Spencer" authorityName="Spencer" box="[619,965,1858,1884]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="inepta">
<emphasis id="B95EF07A230BFFEBFDDDF89DFC922570" box="[619,841,1858,1883]" italics="true" pageId="3" pageNumber="4">Cerodontha inepta</emphasis>
(Spencer)
</taxonomicName>
a species known from
<collectingCountry id="F33D6CF8230BFFEBFB55F89DFAF02577" box="[1251,1323,1858,1884]" name="Brazil" pageId="3" pageNumber="4">Brazil</collectingCountry>
(
<bibRefCitation id="EFBB5199230CFFECFEA6FEC3FE6D231D" author="Spencer" box="[272,438,284,310]" pageId="4" pageNumber="8" refString="Spencer, K. A. (1963) A synopsis of the Neotropical Agromyzidae (Diptera). Transactions of the Royal Entomological Society of London, 115, 291 - 389." type="journal article" year="1963">Spencer 1963</bibRefCitation>
). That species was originally placed in the subgenus
<emphasis id="B95EF07A230CFFECFB96FEC3FB51231E" box="[1056,1162,284,309]" italics="true" pageId="4" pageNumber="5">Poemyza</emphasis>
and was later transferred to
<taxonomicName id="4C2A57EB230CFFECFE19FE9BFCDA2375" authority="Spencer 1966" authorityName="Spencer" authorityYear="1966" box="[431,769,324,350]" class="Insecta" family="Agromyzidae" genus="Dizygomyza" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B95EF07A230CFFECFE19FE9BFDE52376" box="[431,574,324,349]" italics="true" pageId="4" pageNumber="5">Dizygomyza</emphasis>
(
<bibRefCitation id="EFBB5199230CFFECFDF9FE9BFD2C2375" author="Spencer" box="[591,759,324,350]" pageId="4" pageNumber="8" refString="Spencer, K. A. (1966) Notes on Neotropical Agromyzidae. Papeis Avulsos do Departamento de Zoologia. Secretaria da Agricultura. Sao Paulo, Brasil, 19, 141 - 150." type="journal article" year="1966">Spencer 1966</bibRefCitation>
)
</taxonomicName>
. But according to the description of the lunule (high and narrow, upper margin above level of upper
<emphasis id="B95EF07A230CFFECFC3BFEB3FC7423AE" box="[909,943,364,389]" italics="true" pageId="4" pageNumber="5">ori</emphasis>
) it seems that this species does not belong in the subgenus
<taxonomicName id="4C2A57EB230CFFECFDF1FE4BFD0D2386" box="[583,726,404,429]" class="Insecta" family="Agromyzidae" genus="Dizygomyza" higherTaxonomySource="CoL" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B95EF07A230CFFECFDF1FE4BFD0D2386" box="[583,726,404,429]" italics="true" pageId="4" pageNumber="5">Dizygomyza</emphasis>
</taxonomicName>
. Unfortunately the species is known from a female specimen only. Other than the higher lunule,
<taxonomicName id="4C2A57EB230CFFECFC87FE63FBCB23FE" box="[817,1040,444,469]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="inepta">
<emphasis id="B95EF07A230CFFECFC87FE63FBCB23FE" box="[817,1040,444,469]" italics="true" pageId="4" pageNumber="5">Cerodontha inepta</emphasis>
</taxonomicName>
differs externally from
<taxonomicName id="4C2A57EB230CFFECFEBEFE3BFDA223D6" box="[264,633,484,509]" class="Insecta" family="Agromyzidae" genus="Cerodontha" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nigrihalterata" subGenus="Dizygomyza">
<emphasis id="B95EF07A230CFFECFEBEFE3BFDA223D6" box="[264,633,484,509]" italics="true" pageId="4" pageNumber="5">Cerodontha (D.) nigrihalterata</emphasis>
</taxonomicName>
by having 2
<emphasis id="B95EF07A230CFFECFCAFFE3BFCE023D6" box="[793,827,484,509]" italics="true" pageId="4" pageNumber="5">ori</emphasis>
and 3
<emphasis id="B95EF07A230CFFECFC39FE3BFC6723D6" box="[911,956,484,509]" italics="true" pageId="4" pageNumber="5">ors,</emphasis>
orbital setulae unusually long, last section of CuA1 only slightly shorter than penultimate in ratio 20:22, wing only slightly brownish, and stalk of halter yellowish.
</paragraph>
</subSubSection>
</treatment>
</document>
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