<documentid="DAB263100739C9D3BAAEC98C41E362F8"ID-CLB-Dataset="48132"ID-DOI="10.5281/zenodo.207073"ID-GBIF-Dataset="a3815507-6100-464e-aaf6-60d6820a1eec"ID-ISSN="1175-5326"ID-Zenodo-Dep="207073"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.tables_requiresApprovalFor="existingObjects,plazi"IM.taxonomicNames_approvedBy="felipe"checkinTime="1460228776049"checkinUser="plazi"docAuthor="Torres-Carvajal, Omar, Etheridge, Richard & Queiroz, Kevin De"docDate="2011"docId="038287FDFFAD605F69838D73D3A0F9CC"docLanguage="en"docName="zt02752p044.pdf"docOrigin="Zootaxa 2752"docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article"docStyleId="890A69B780ED73D6DB8551B71C8AC79E"docStyleName="Zootaxa.2009-2012.journal_article"docStyleVersion="4"docTitle="Enyalioides oshaughnessyi Boulenger 1881"docType="treatment"docVersion="11"lastPageNumber="25"masterDocId="FFBBFF85FFBB604769148B6ED174FF93"masterDocTitle="A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)"masterLastPageNumber="44"masterPageNumber="1"pageNumber="23"updateTime="1720202091178"updateUser="ExternalLinkService">
<mods:titleid="5646BFA0E2EB68063B55B6B0A9AB29BA">A systematic revision of Neotropical lizards in the clade Hoplocercinae (Squamata: Iguania)</mods:title>
<bibRefCitationid="EFBA4B1AFFAD605168F88D73D3C6F9A4"author="Boulenger"box="[492,690,1565,1591]"pageId="22"pageNumber="38"refString="Boulenger, G. A. (1881) Description of a new species of Enyalius in the Brussels Museum. Proceedings of the Zoological Society of London, 1881, 246 - 247, 1 plate."type="book chapter"year="1881">Boulenger 1881</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraphid="8B9436EBFFAD605169838D35D3BEF9E2"blockId="22.[151,761,1627,1680]"box="[151,714,1627,1650]"pageId="22"pageNumber="23">Proposed standard English name: red-eyed woodlizards</paragraph>
<paragraphid="8B9436EBFFAD605169838D14D38DF903"blockId="22.[151,761,1627,1680]"box="[151,761,1658,1680]"pageId="22"pageNumber="23">Proposed standard Spanish name: lagartijas de palo ojirrojas</paragraph>
<bibRefCitationid="EFBA4B1AFFAD605168848DDAD307F959"author="Boulenger"box="[400,627,1716,1738]"pageId="22"pageNumber="38"refString="Boulenger, G. A. (1881) Description of a new species of Enyalius in the Brussels Museum. Proceedings of the Zoological Society of London, 1881, 246 - 247, 1 plate."type="book chapter"year="1881">Boulenger (1881:246)</bibRefCitation>
<bibRefCitationid="EFBA4B1AFFAD605168BB8DBCD3E5F97B"author="Boulenger"box="[431,657,1746,1768]"pageId="22"pageNumber="38"refString="Boulenger, G. A. (1885) Catalogue of the Lizards in the British Museum (Natural History) II. Taylor and Francis, London, 497 pp."type="book"year="1885">Boulenger (1885:115)</bibRefCitation>
<bibRefCitationid="EFBA4B1AFFAD60516B8F8DBAD22CF97B"author="Burt"box="[667,856,1746,1769]"pageId="22"pageNumber="38"refString="Burt, C. E. & Burt, M. D. (1931) South American lizards in the collection of the American Museum of Natural History. Bulletin of the American Museum of Natural History, 61, 227 - 395."type="journal article"year="1931">Burt & Burt (1931</bibRefCitation>
:267
</treatmentCitation>
; 1933:24);
<bibRefCitationid="EFBA4B1AFFAD60516AEE8DBAD418F97B"author="Peters"box="[1018,1388,1746,1769]"pageId="22"pageNumber="40"refString="Peters, J. A. & Donoso-Barros, R. (1970) Catalogue of the neotropical Squamata: Part II. Lizards and amphisbaenians. United States National Museum Bulletin, 297, 1 - 293."type="journal article"year="1970">Peters & Donoso-Barros (1970:115)</bibRefCitation>
with a bright red iris in both sexes (but see Remarks) and a dark gular patch restricted to the inner aspect of the gular fold in males. Other species of
(1) dorsal head scales conical or multicarinate, strongly projecting dorsally; (2) posterior superciliaries not enlarged relative to adjacent scales; (3) scales on lateral edge of skull roof just posterior to superciliaries strongly projecting; the projection is more pronounced in adults; (4) one or two enlarged pretympanic scales present; (5) gular scales conical or multicarinate, strongly projecting ventrally; (6) dorsal neck scales homogeneous in size, mostly conical or cycloid and slightly imbricate; lateral neck scales similar in size to dorsal neck scales except for some enlarged conical scales, which sometimes form an oblique row extending from tympanum to scapular region; (7) vertebrals larger than adjacent dorsals, forming distinct raised middorsal crest that extends onto tail as a pair of crests; (8) nuchal region with continuous and usually (92.3%) single middorsal crest; (9) dorsals flat, usually (76.9%) smooth, and homogeneous in size; (10) longitudinal row of raised, enlarged scales between dorsals and flank scales usually (84.6%) present; when present this row is continuous; (11) scales on flanks heterogeneous in size, with most scales similar in size and shape to dorsals and a few enlarged, circular, and keeled scales; (12) ventrals keeled; (13) fore limb scales keeled dorsally and ventrally; (14) hind limb scales keeled dorsally and ventrally; scattered enlarged scales absent; dorsal scales of pes homogeneous in size; (15) caudals heterogeneous, increasing in size posteriorly on each segment (6–8 scales in lateral view), not modified as conspicuous spines (
<figureCitationid="13102A6EFFA3605F6C7B8BB3D1D2FE84"captionStart="FIGURE 3"captionStartId="12.[151,250,1517,1540]"captionTargetBox="[280,1305,321,1491]"captionTargetId="figure@12.[274,1314,306,1505]"captionTargetPageId="12"captionText="FIGURE 3. Caudal segments from the anterior third of the tail of ten species of Hoplocercinae in lateral view. (A) Enyalioides cofanorum, (B) E. heterolepis, (C) E. laticeps, (D) E. microlepis, (E) E. oshaughnessyi, (F) E. palpebralis, (G) E. praestabilis, (H) E. rubrigularis, (I) E. touzeti, and (J) Morunasaurus annularis. Scale bars = 5 mm. Illustration by D. Paucar."httpUri="https://zenodo.org/record/207076/files/figure.png"pageId="24"pageNumber="25">Fig. 3</figureCitation>
); (16) tail compressed laterally. Meristic and morphometric characters are presented in
<tableCitationid="C6A90350FFA3605F6D6F8B91D5BAFE84"box="[1147,1230,255,279]"captionStart="TABLE 1"captionStartId="6.[151,235,1304,1325]"captionTargetBox="[151,1436,1377,1892]"captionTargetPageId="6"captionText="TABLE 1. Summary of counts and measurements (mm) for species of Hoplocercinae. Range (first line) and mean ± standard"httpUri="http://table.plazi.org/id/DF546663FFBD604169838E76D4E8FABD"pageId="24"pageNumber="25"tableUuid="DF546663FFBD604169838E76D4E8FABD">Table 1</tableCitation>
<emphasisid="B95FEAF9FFA3605F69D18A4CD0E0FEA8"bold="true"box="[197,404,290,315]"pageId="24"pageNumber="25">Coloration in life</emphasis>
(
<figureCitationid="13102A6EFFA3605F68B28A4CD099FEA9"box="[422,493,290,314]"captionStart="FIGURE 4"captionStartId="14.[151,250,1897,1920]"captionTargetBox="[204,1383,190,1874]"captionTargetId="figure@14.[204,1384,184,1876]"captionTargetPageId="14"captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae."httpUri="https://zenodo.org/record/207077/files/figure.png"pageId="24"pageNumber="25">Fig. 4</figureCitation>
). Adult male KU 109630: dorsal background olive green with light green and turquoise blue spots on flanks, tail, and dorsal surfaces of legs; labials and chin greenish yellow; gular region light blue with black patch under gular fold; chest greenish tan; venter and posterior surfaces of thighs tan; ventral surface of tail olive tan; iris rusty orange; throat and tongue flesh colored (W.E. Duellman,
<dateid="FF95102BFFA3605F6DB18AE4D42DFE31"box="[1189,1369,394,418]"pageId="24"pageNumber="25"value="1963-03-30">30 March 1963</date>
, field notes). All adult males seen by the authors alive or in photographs have a red iris (
<figureCitationid="13102A6EFFA3605F6D258AC3D502FE56"box="[1073,1142,429,453]"captionStart="FIGURE 4"captionStartId="14.[151,250,1897,1920]"captionTargetBox="[204,1383,190,1874]"captionTargetId="figure@14.[204,1384,184,1876]"captionTargetPageId="14"captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae."httpUri="https://zenodo.org/record/207077/files/figure.png"pageId="24"pageNumber="25">Fig. 4</figureCitation>
Adult female (based on photograph by P.S. Hamilton,
<figureCitationid="13102A6EFFA3605F6A3F8AA1D204FE74"box="[811,880,463,487]"captionStart="FIGURE 4"captionStartId="14.[151,250,1897,1920]"captionTargetBox="[204,1383,190,1874]"captionTargetId="figure@14.[204,1384,184,1876]"captionTargetPageId="14"captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae."httpUri="https://zenodo.org/record/207077/files/figure.png"pageId="24"pageNumber="25">Fig. 4</figureCitation>
): dorsal background light brown with light green and light blue spots on flanks, tail, and hind limbs; labials light green, greenish brown, or reddish brown; distal margin of superciliaries, canthals, some suboculars, and some pretympanics bright green, light blue post-tympanic blotch; light green scales on chin and gular region; iris red.
<figureCitationid="13102A6EFFA3605F6D5A89ACD5E6FD49"box="[1102,1170,706,730]"captionStart="FIGURE 7"captionStartId="23.[151,250,1564,1587]"captionTargetBox="[197,1383,204,1538]"captionTargetId="figure@23.[189,1403,169,1564]"captionTargetPageId="23"captionText="FIGURE 7. Distribution of Enyalioides microlepis (dots) and E. oshaughnessyi (triangles). A, Alto Tambo; B, Bilsa Ecological Reserve (see Remarks under E. oshaughnessyi)."httpUri="https://zenodo.org/record/207084/files/figure.png"pageId="24"pageNumber="25">Fig. 7</figureCitation>
). This species is known to occur in sympatry with
<bibRefCitationid="EFBA4B1AFFA3605F6DD18844D4E3FCD1"author="Burt"box="[1221,1431,810,834]"pageId="24"pageNumber="38"refString="Burt, C. E. & Burt, M. D. (1931) South American lizards in the collection of the American Museum of Natural History. Bulletin of the American Museum of Natural History, 61, 227 - 395."type="journal article"year="1931">Burt & Burt 1931</bibRefCitation>
;
<bibRefCitationid="EFBA4B1AFFA3605F69838820D098FCF6"author="Peters"box="[151,492,845,869]"pageId="24"pageNumber="40"refString="Peters, J. A. & Donoso-Barros, R. (1970) Catalogue of the neotropical Squamata: Part II. Lizards and amphisbaenians. United States National Museum Bulletin, 297, 1 - 293."type="journal article"year="1970">Peters & Donoso-Barros 1970</bibRefCitation>
;
<bibRefCitationid="EFBA4B1AFFA3605F68EC8823D3CAFCF6"author="Almendariz"box="[504,702,845,869]"pageId="24"pageNumber="38"refString="Almendariz, A. (1992) Anfibios y reptiles. Revista Politecnica, Escuela Politecnica Nacional, 16, 89 - 162."type="journal article"year="1992">Almendáriz 1992</bibRefCitation>
;
<bibRefCitationid="EFBA4B1AFFA3605F6BDD8823D2C8FCF6"author="Torres-Carvajal"box="[713,956,845,869]"pageId="24"pageNumber="41"refString="Torres-Carvajal, O. (2001) Lizards of Ecuador: checklist, distribution, and systematic references. Smithsonian Herpetological Information Service, 131, 1 - 35."type="journal article"year="2001">Torres-Carvajal 2001</bibRefCitation>
16))). According to these authors, this pattern can be explained by fixed morphological differences between the species evolving prior to the evolution of reciprocal monophyly of their mtDNA alleles (see also remarks under
<figureCitationid="13102A6EFFA3605F69838FA4D1AFFB71"box="[151,219,1226,1250]"captionStart="FIGURE 7"captionStartId="23.[151,250,1564,1587]"captionTargetBox="[197,1383,204,1538]"captionTargetId="figure@23.[189,1403,169,1564]"captionTargetPageId="23"captionText="FIGURE 7. Distribution of Enyalioides microlepis (dots) and E. oshaughnessyi (triangles). A, Alto Tambo; B, Bilsa Ecological Reserve (see Remarks under E. oshaughnessyi)."httpUri="https://zenodo.org/record/207084/files/figure.png"pageId="24"pageNumber="25">Fig. 7</figureCitation>
), both have reddish-brown irises (
<figureCitationid="13102A6EFFA3605F6B4F8FA4D3D5FB71"box="[603,673,1226,1250]"captionStart="FIGURE 4"captionStartId="14.[151,250,1897,1920]"captionTargetBox="[204,1383,190,1874]"captionTargetId="figure@14.[204,1384,184,1876]"captionTargetPageId="14"captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae."httpUri="https://zenodo.org/record/207077/files/figure.png"pageId="24"pageNumber="25">Fig. 4</figureCitation>
” seen by the authors (live specimens and photographs) have red irises (
<figureCitationid="13102A6EFFA3605F6A338F83D218FA96"box="[807,876,1261,1285]"captionStart="FIGURE 4"captionStartId="14.[151,250,1897,1920]"captionTargetBox="[204,1383,190,1874]"captionTargetId="figure@14.[204,1384,184,1876]"captionTargetPageId="14"captionText="FIGURE 4. Photographs of live lizards representing twelve species of Hoplocercinae."httpUri="https://zenodo.org/record/207077/files/figure.png"pageId="24"pageNumber="25">Fig. 4</figureCitation>
), including both male and female lizards recently photographed near the locality where
<figureCitationid="13102A6EFFA3605F6BD48E5CD277FAD9"box="[704,771,1330,1354]"captionStart="FIGURE 7"captionStartId="23.[151,250,1564,1587]"captionTargetBox="[197,1383,204,1538]"captionTargetId="figure@23.[189,1403,169,1564]"captionTargetPageId="23"captionText="FIGURE 7. Distribution of Enyalioides microlepis (dots) and E. oshaughnessyi (triangles). A, Alto Tambo; B, Bilsa Ecological Reserve (see Remarks under E. oshaughnessyi)."httpUri="https://zenodo.org/record/207084/files/figure.png"pageId="24"pageNumber="25">Fig. 7</figureCitation>
