treatments-xml/data/14/54/B3/1454B336FF90A25E6E9E4D26FCDA2A28.xml

<document id="3C89380D9CB121B0FD59617FA1C4C03A" ID="10.11646/zootaxa.4170.3.6" ID-DOI="10.11646/zootaxa.4170.3.6" ID-GBIF-Dataset="83ea9489-9668-4132-b965-42183147c6b4" ID-ISSN="1175-5326" ID-Zenodo-Dep="260633" ID-ZooBank="3FB4ADDC-E67B-42AD-98D1-956E42211694" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1474873756993" checkinUser="plazi" docAuthor="Sung, Yik-Hei, Hu, Ping, Wang, Jian, Liu, Hai-Jun &amp; Wang, Ying-Yong" docDate="2016" docId="1454B336FF90A25E6E9E4D26FCDA2A28" docLanguage="en" docName="zootaxa.4170.3.6.pdf" docOrigin="Zootaxa 4170 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amolops albispinus Sung, Wang and Wang, sp. nov." docType="treatment" docVersion="7" lastPageNumber="534" masterDocId="E86DCB4EFF95A2576E094B21FFF52B68" masterDocTitle="A new species of Amolops (Anura: Ranidae) from southern China" masterLastPageNumber="538" masterPageNumber="525" pageNumber="530" updateTime="1698387244303" updateUser="plazi">
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<mods:title id="E9A9182C1A0CF4FB1FB45D2D996C0C7F">A new species of Amolops (Anura: Ranidae) from southern China</mods:title>
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<mods:namePart id="E75A0651CD3E5E31F23A3244D60130B1">Sung, Yik-Hei</mods:namePart>
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<mods:namePart id="6F1748C648336F28E028C8808AAB0CD4">Hu, Ping</mods:namePart>
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<mods:namePart id="E27A3EEF182AB9F3A68537649056A6A8">Wang, Jian</mods:namePart>
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<mods:namePart id="B2BD8480EC02209A9C1C826FD5B1F268">Liu, Hai-Jun</mods:namePart>
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<mods:namePart id="66F0B42F6606A41EA15194E443984FE9">Wang, Ying-Yong</mods:namePart>
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<heading id="C70AB54CFF90A2526E9E4D26FCFA2D4A" bold="true" box="[151,783,1543,1570]" fontSize="11" level="1" pageId="5" pageNumber="530" reason="1">
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<taxonomicName id="5BFD79A3FF90A2526E9E4D26FD5C2D4A" ID-CoL="CW9H" authority="Sung, Wang and Wang" authorityName="Sung, Wang and Wang" box="[151,681,1543,1570]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="5" pageNumber="530" phylum="Chordata" rank="species" species="albispinus" status="sp. nov.">
<emphasis id="AE89DE32FF90A2526E9E4D26FE712D49" bold="true" box="[151,388,1543,1569]" italics="true" pageId="5" pageNumber="530">Amolops albispinus</emphasis>
Sung, Wang and Wang
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<taxonomicNameLabel id="B5BA6349FF90A2526CB84D29FCFA2D4A" box="[689,783,1544,1570]" pageId="5" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="04C61EA5FF90A2526E964D0BFF172D2B" box="[159,226,1578,1603]" captionStart="FIGURE 3" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[190,1388,201,1904]" captionTargetId="figure@6.[183,1399,195,1915]" captionTargetPageId="6" captionText="FIGURE 3. Dorsolateral view of adult male holotype SYS a 003454 of Amolops albispinus sp. nov. in life; B: ventral view of the holotype in life; C: hand of the holotype in life; D: foot of the holotype in life; E: Dorsal view of the holotype in preserve; F: ventral view of the holotype in preserve; G: close-up of the head of the holotype in life." httpUri="https://zenodo.org/record/260636/files/figure.png" pageId="5" pageNumber="530">Fig. 3</figureCitation>
and 
<figureCitation id="04C61EA5FF90A2526F114D0AFED22D2B" box="[280,295,1579,1603]" captionStart="FIGURE 4" captionStartId="9.[151,250,1641,1663]" captionTargetBox="[151,1436,375,1616]" captionTargetId="figure@9.[151,1436,375,1616]" captionTargetPageId="9" captionText="FIGURE 4. Comparisons of morphological characteristics between Amolops albispinus sp. nov. and species in the Amolops ricketti and Amolops daiyunensis species groups. A: Amolops albispinus sp. nov.; B: Amolops ricketti; C: Amolops wuyiensis; D: Amolops daiyunensi; E. Amolops hongkongnesis. 1: dorsolateral view; 2: close-up of the head; 3: ventral view of the hand; 4: ventral view of the leg." httpUri="https://zenodo.org/record/260637/files/figure.png" pageId="5" pageNumber="530">4</figureCitation>
)
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<paragraph id="9C420220FF90A2526E9E4D53FABC2DC7" blockId="5.[151,1436,1650,1999]" pageId="5" pageNumber="530">
<materialsCitation id="2C95087DFF90A2526E9E4D53FABC2DC7" ID-GBIF-Occurrence="1315044626" collectingDate="1926-01" collectionCode="SYS" collectorName="Jian Wang &amp; Zu-Yao Liu &amp; Zhi-Tong Lyu" country="China" elevation="260" latitude="22.58189" location="Mt. Wutong" longLatPrecision="1" longitude="114.200745" municipality="Shenzhen City" pageId="5" pageNumber="530" specimenCount="1" stateProvince="Guangdong Province" typeStatus="holotype">
<emphasis id="AE89DE32FF90A2526E9E4D53FEF82DE3" bold="true" box="[151,269,1650,1675]" pageId="5" pageNumber="530">
<typeStatus id="4346BC82FF90A2526E9E4D53FEFD2DE3" box="[151,264,1650,1675]" pageId="5" pageNumber="530" type="holotype">Holotype</typeStatus>
:
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<collectionCode id="FAEC9AE5FF90A2526F1C4D52FEBD2DE3" box="[277,328,1651,1675]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="5" pageNumber="530">SYS</collectionCode>
a003454, adult male, collected by 
<collectorName id="310867F6FF90A2526CDC4D53FCB82DE3" box="[725,845,1650,1675]" pageId="5" pageNumber="530">Jian Wang</collectorName>
, 
<collectorName id="310867F6FF90A2526D504D52FC292DE3" box="[857,988,1650,1675]" pageId="5" pageNumber="530">Zu-Yao Liu</collectorName>
and 
<collectorName id="310867F6FF90A2526A1D4D53FB442DE3" box="[1044,1201,1650,1675]" pageId="5" pageNumber="530">Zhi-Tong Lyu</collectorName>
on 
<collectingDate id="F807DD08FF90A2526AD54D53FAAD2DE3" box="[1244,1368,1650,1675]" pageId="5" pageNumber="530" value="1926-01">
<date id="E84324E0FF90A2526AD54D53FAAD2DE3" box="[1244,1368,1650,1675]" pageId="5" pageNumber="530" value="1926-01">26 January</date>
</collectingDate>
, 2015 from 
<location id="992254FBFF90A2526EDD4DB6FEA92DC7" LSID="urn:lsid:plazi:treatment:1454B336FF90A25E6E9E4D26FCDA2A28:992254FBFF90A2526EDD4DB6FEA92DC7" box="[212,348,1687,1711]" country="China" latitude="22.58189" longLatPrecision="1" longitude="114.200745" municipality="Shenzhen City" name="Mt. Wutong" pageId="5" pageNumber="530" stateProvince="Guangdong Province">Mt. Wutong</location>
(
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, 
<geoCoordinate id="F9C964E7FF90A2526C1C4DB7FD5B2DC7" box="[533,686,1686,1711]" direction="east" orientation="longitude" pageId="5" pageNumber="530" precision="1" value="114.200745">114°12′2.7″E</geoCoordinate>
; 
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<quantity id="5B05AFC5FF90A2526CB04DB7FCF52DC6" box="[697,768,1686,1711]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="530" unit="m" value="260.0">260 m</quantity>
a.s.l.
</elevation>
), 
<collectingMunicipality id="7C26985AFF90A2526D5A4DB7FC0D2DC7" box="[851,1016,1686,1711]" pageId="5" pageNumber="530">Shenzhen City</collectingMunicipality>
, 
<collectingRegion id="5E39CCC2FF90A2526A0A4DB7FB012DC7" box="[1027,1268,1686,1711]" country="China" name="Guangdong" pageId="5" pageNumber="530">Guangdong Province</collectingRegion>
, 
<collectingCountry id="E4EA42B0FF90A2526AF74DB7FAB12DC7" box="[1278,1348,1686,1711]" name="China" pageId="5" pageNumber="530">China</collectingCountry>
.
</materialsCitation>
</paragraph>
<paragraph id="9C420220FF90A2526ECE4D9BFA602CA7" blockId="5.[151,1436,1650,1999]" pageId="5" pageNumber="530">
<materialsCitation id="2C95087DFF90A2526ECE4D9BFC622D9F" ID-GBIF-Occurrence="1315044629" county="Mt. Wutong" location="Ying-Yong Wang" municipality="Zhi-Tong Lyu" pageId="5" pageNumber="530" specimenCount="13" specimenCount-adult="13" typeStatus="paratype">
<emphasis id="AE89DE32FF90A2526ECE4D9BFEC82DBB" bold="true" box="[199,317,1722,1747]" pageId="5" pageNumber="530">
<typeStatus id="4346BC82FF90A2526ECE4D9BFECD2DBB" box="[199,312,1722,1747]" pageId="5" pageNumber="530" type="paratype">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="8AFBC9A9FF90A2526F4E4D9AFE502DBB" box="[327,421,1722,1747]" pageId="5" pageNumber="530" type="adult">13 adult</specimenCount>
specimens, collected 
<location id="992254FBFF90A2526CAC4D9BFC9B2DBB" LSID="urn:lsid:plazi:treatment:1454B336FF90A25E6E9E4D26FCDA2A28:992254FBFF90A2526CAC4D9BFC9B2DBB" box="[677,878,1722,1747]" county="Mt. Wutong" municipality="Zhi-Tong Lyu" name="Ying-Yong Wang" pageId="5" pageNumber="530">Ying-Yong Wang</location>
, 
<location id="992254FBFF90A2526D744D9BFBC22DBB" LSID="urn:lsid:plazi:treatment:1454B336FF90A25E6E9E4D26FCDA2A28:992254FBFF90A2526D744D9BFBC22DBB" box="[893,1079,1722,1747]" county="Mt. Wutong" municipality="Zhi-Tong Lyu" name="Jian-Huan Yang" pageId="5" pageNumber="530">Jian-Huan Yang</location>
, 
<location id="992254FBFF90A2526A4C4D9BFB3C2DBB" LSID="urn:lsid:plazi:treatment:1454B336FF90A25E6E9E4D26FCDA2A28:992254FBFF90A2526A4C4D9BFB3C2DBB" box="[1093,1225,1722,1747]" county="Mt. Wutong" municipality="Zhi-Tong Lyu" name="Run-Lin Li" pageId="5" pageNumber="530">Run-Lin Li</location>
, 
<location id="992254FBFF90A2526AD14D9AFA952DBB" LSID="urn:lsid:plazi:treatment:1454B336FF90A25E6E9E4D26FCDA2A28:992254FBFF90A2526AD14D9AFA952DBB" box="[1240,1376,1722,1747]" county="Mt. Wutong" municipality="Zhi-Tong Lyu" name="Zu-Yao Liu" pageId="5" pageNumber="530">Zu-Yao Liu</location>
, 
<location id="992254FBFF90A2526B664D9BFF2E2D9F" LSID="urn:lsid:plazi:treatment:1454B336FF90A25E6E9E4D26FCDA2A28:992254FBFF90A2526B664D9BFF2E2D9F" county="Mt. Wutong" municipality="Zhi-Tong Lyu" name="Jian Wang" pageId="5" pageNumber="530">Jian Wang</location>
, and 
<collectingMunicipality id="7C26985AFF90A2526F1C4DFFFE442D9F" box="[277,433,1758,1783]" pageId="5" pageNumber="530">Zhi-Tong Lyu</collectingMunicipality>
from 
<collectingCounty id="75237AACFF90A2526FFC4DFEFD882D9F" box="[501,637,1759,1783]" pageId="5" pageNumber="530">Mt. Wutong</collectingCounty>
at elevations between 
<quantity id="5B05AFC5FF90A2526D744DFFFC622D9F" box="[893,919,1758,1783]" metricMagnitude="2" metricUnit="m" metricValue="2.925" metricValueMax="5.0" metricValueMin="0.85" pageId="5" pageNumber="530" unit="m" value="292.5" valueMax="500.0" valueMin="85.0">85</quantity>
</materialsCitation>
– 
<materialsCitation id="2C95087DFF90A2526DAF4DFFFE082C73" ID-GBIF-Occurrence="1315044627" collectingDate="2012-01-13" collectionCode="SYS" location="Seven" municipality="Yong Wang" pageId="5" pageNumber="530" specimenCount="500" typeStatus="paratype">
<specimenCount id="8AFBC9A9FF90A2526DAF4DFFFC002D9E" box="[934,1013,1758,1783]" pageId="5" pageNumber="530" type="generic">500 m.</specimenCount>
<location id="992254FBFF90A2526DF54DFEFBB52D9F" LSID="urn:lsid:plazi:treatment:1454B336FF90A25E6E9E4D26FCDA2A28:992254FBFF90A2526DF54DFEFBB52D9F" box="[1020,1088,1759,1783]" municipality="Yong Wang" name="Seven" pageId="5" pageNumber="530">Seven</location>
males: 
<collectionCode id="FAEC9AE5FF90A2526A9E4DFEFB3E2D9F" box="[1175,1227,1759,1783]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="5" pageNumber="530">SYS</collectionCode>
a003364 by Ying- 
<collectingMunicipality id="7C26985AFF90A2526E9E4C22FEEF2C73" box="[151,282,1795,1819]" pageId="5" pageNumber="530">Yong Wang</collectingMunicipality>
on 
<collectingDate id="F807DD08FF90A2526F4B4C22FE082C73" box="[322,509,1795,1819]" pageId="5" pageNumber="530" value="2012-01-13">
<date id="E84324E0FF90A2526F4B4C22FE082C73" box="[322,509,1795,1819]" pageId="5" pageNumber="530" value="2012-01-13">13 January 2012</date>
</collectingDate>
</materialsCitation>
, SYS a001509 by Jian-Huan Yang and Run-Lin Li on 
<date id="E84324E0FF90A2526A504C23FB0D2C73" box="[1113,1272,1794,1819]" pageId="5" pageNumber="530" value="2012-03-05">5 March 2012</date>
, SYS a003271 and 3272 by Zu-Tao Liu, Jian Wang and Zhi-Tong Lyu on 
<date id="E84324E0FF90A2526D354C07FBE82C57" box="[828,1053,1830,1855]" pageId="5" pageNumber="530" value="2014-09-16">16 September 2014</date>
, SYS a003452 by Jian Wang and Run-Lin Li on 
<date id="E84324E0FF90A2526F4D4C6BFDF62C0B" box="[324,515,1866,1891]" pageId="5" pageNumber="530" value="2015-01-26">26 January 2015</date>
, SYS a003473 by Zu-Yao Liu, Jian Wang and Zhi-Tong Lyu on 
<date id="E84324E0FF90A2526AEE4C6AFA6D2C0B" box="[1255,1432,1866,1891]" pageId="5" pageNumber="530" value="2015-03-13">13 March 2015</date>
, and SYS a004511 by Zhi-Tong Lyu and Jian Wang on 
<date id="E84324E0FF90A2526D034C4EFC392CEF" box="[778,972,1902,1927]" pageId="5" pageNumber="530" value="2015-10-22">22 October 2015</date>
; six females: SYS a001508, 1513, 1514, and 1526 by Ying-Yong Wang, Jian-Huan Yang, Run-Lin Li on 
<date id="E84324E0FF90A2526D634CB2FBFE2CC3" box="[874,1035,1938,1963]" pageId="5" pageNumber="530" value="2012-03-08">8 March 2012</date>
, SYS a003270 by Zu-Yao Liu, Jian Wang and Zhi-Tong Lyu on 
<date id="E84324E0FF90A2526FDD4C97FD442CA7" box="[468,689,1974,1999]" pageId="5" pageNumber="530" value="2014-09-16">16 September 2014</date>
, SYS a003453 by Jian Wang and Run-Lin Li on 
<date id="E84324E0FF90A2526ADD4C97FA642CA7" box="[1236,1425,1974,1999]" pageId="5" pageNumber="530" value="2015-01-26">26 January 2015</date>
.
</paragraph>
<caption id="C88252A8FF93A2516E9E4CB1FBD92C8B" httpUri="https://zenodo.org/record/260636/files/figure.png" pageId="6" pageNumber="531" targetBox="[190,1388,201,1904]" targetPageId="6">
<paragraph id="9C420220FF93A2516E9E4CB1FBD92C8B" blockId="6.[151,1436,1936,2019]" pageId="6" pageNumber="531">
<emphasis id="AE89DE32FF93A2516E9E4CB1FEE62CCE" bold="true" box="[151,275,1936,1958]" pageId="6" pageNumber="531">FIGURE 3.</emphasis>
Dorsolateral view of adult male holotype SYS a003454 of 
<taxonomicName id="5BFD79A3FF93A2516D784CB1FBCC2CCE" box="[881,1081,1936,1958]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="6" pageNumber="531" phylum="Chordata" rank="species" species="albispinus" status="sp. nov.">
<emphasis id="AE89DE32FF93A2516D784CB1FBCC2CCE" box="[881,1081,1936,1958]" italics="true" pageId="6" pageNumber="531">Amolops albispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5BA6349FF93A2516A364CB1FB652CCE" box="[1087,1168,1936,1958]" pageId="6" pageNumber="531" rank="species">
<emphasis id="AE89DE32FF93A2516A364CB1FB652CCE" bold="true" box="[1087,1168,1936,1958]" pageId="6" pageNumber="531">sp. nov.</emphasis>
</taxonomicNameLabel>
in life; 
<emphasis id="AE89DE32FF93A2516AEB4CB1FB0E2CCE" bold="true" box="[1250,1275,1936,1958]" pageId="6" pageNumber="531">B:</emphasis>
ventral view of the holotype in life; 
<emphasis id="AE89DE32FF93A2516F6C4C8FFE752CAC" bold="true" box="[357,384,1966,1988]" pageId="6" pageNumber="531">C:</emphasis>
hand of the holotype in life; 
<emphasis id="AE89DE32FF93A2516CA14C8FFD362CAC" bold="true" box="[680,707,1966,1988]" pageId="6" pageNumber="531">D:</emphasis>
foot of the holotype in life; 
<emphasis id="AE89DE32FF93A2516DEA4C8FFC0E2CAC" bold="true" box="[995,1019,1966,1988]" pageId="6" pageNumber="531">E:</emphasis>
Dorsal view of the holotype in preserve; 
<emphasis id="AE89DE32FF93A2516E9E4CECFF5A2C8B" bold="true" box="[151,175,1997,2019]" pageId="6" pageNumber="531">F:</emphasis>
ventral view of the holotype in preserve; 
<emphasis id="AE89DE32FF93A2516C5F4CECFD9F2C8B" bold="true" box="[598,618,1997,2019]" pageId="6" pageNumber="531">G</emphasis>
: close-up of the head of the holotype in life.
</paragraph>
</caption>
<paragraph id="9C420220FF92A2506ECE4BB6FAC72B90" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
Other specimens examined include five juveniles: SYS a001510 collected by Jian-Huan Yang and Run-Lin Li on 
<date id="E84324E0FF92A2506EB64B9DFE9D2BBC" box="[191,360,188,213]" pageId="7" pageNumber="532" value="2012-03-05">5 March 2012</date>
, 
<materialsCitation id="2C95087DFF92A2506F714B9DFADB2B90" ID-GBIF-Occurrence="1315044628" collectingDate="2012-03-10" collectingDateMax="2015-03-13" collectingDateMin="2012-03-10" collectionCode="SYS" collectorName="Jian-Huan Yang &amp; Run-Lin Li &amp; Jian Wang &amp; Zhi-Tong Lyu" location="Mt. Wutong" pageId="7" pageNumber="532" specimenCount="1">
<collectionCode id="FAEC9AE5FF92A2506F714B9DFE5E2BBC" box="[376,427,188,212]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="7" pageNumber="532">SYS</collectionCode>
a001532 collected by 
<collectorName id="310867F6FF92A2506CB44B9DFC832BBD" box="[701,886,188,213]" pageId="7" pageNumber="532">Jian-Huan Yang</collectorName>
and 
<collectorName id="310867F6FF92A2506DBF4B9DFBCC2BBD" box="[950,1081,188,213]" pageId="7" pageNumber="532">Run-Lin Li</collectorName>
on 
<collectingDate id="F807DD08FF92A2506A644B9DFAD02BBC" box="[1133,1317,188,213]" pageId="7" pageNumber="532" value="2012-03-10">
<date id="E84324E0FF92A2506A644B9DFAD02BBC" box="[1133,1317,188,213]" pageId="7" pageNumber="532" value="2012-03-10">10 March 2012</date>
</collectingDate>
, and 
<collectionCode id="FAEC9AE5FF92A2506B604B9DFA692BBC" box="[1385,1436,188,212]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14836" name="Zhongshan (Sun Yatsen) University" pageId="7" pageNumber="532">SYS</collectionCode>
a003474, 3475 and 3481 collected by 
<collectorName id="310867F6FF92A2506C4B4BFEFD422B90" box="[578,695,223,248]" pageId="7" pageNumber="532">Jian Wang</collectorName>
and 
<collectorName id="310867F6FF92A2506CE44BFEFC7C2B90" box="[749,905,223,248]" pageId="7" pageNumber="532">Zhi-Tong Lyu</collectorName>
on 
<date id="E84324E0FF92A2506DBA4BC1FBAB2B90" box="[947,1118,223,248]" pageId="7" pageNumber="532" value="2015-03-13">
<collectingDate id="F807DD08FF92A2506DBA4BC1FBAB2B90" box="[947,1118,223,248]" pageId="7" pageNumber="532" value="2015-03-13">13 March 2015</collectingDate>
</date>
from 
<location id="992254FBFF92A2506AAB4BC1FADB2B90" LSID="urn:lsid:plazi:treatment:1454B336FF90A25E6E9E4D26FCDA2A28:992254FBFF92A2506AAB4BC1FADB2B90" box="[1186,1326,224,248]" name="Mt. Wutong" pageId="7" pageNumber="532">Mt. Wutong</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="D4E751ABFF92A2506ECE4A25FCCA29ED" pageId="7" pageNumber="532" type="diagnosis">
<paragraph id="9C420220FF92A2506ECE4A25FDDA2AEF" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
<emphasis id="AE89DE32FF92A2506ECE4A25FEB42A75" bold="true" box="[199,321,260,285]" pageId="7" pageNumber="532">Diagnosis.</emphasis>
The presence of an abdominal sucker in the tadpole is the diagnostic character for the genus 
<taxonomicName id="5BFD79A3FF92A2506E9E4A06FDEE2A28" authority="Rao &amp; Wilkinson 2007" authorityName="Rao &amp; Wilkinson" authorityYear="2007" box="[151,539,295,320]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="7" pageNumber="532" phylum="Chordata" rank="genus">
<emphasis id="AE89DE32FF92A2506E9E4A06FF0C2A28" box="[151,249,295,320]" italics="true" pageId="7" pageNumber="532">Amolops</emphasis>
(
<bibRefCitation id="F86C7FD1FF92A2506F014A09FDE62A28" author="Rao" box="[264,531,295,320]" pageId="7" pageNumber="532" refString="Rao, D. Q. &amp; Wilkinson, J. A. (2007) A new species of Amolops (Anura: Ranidae) from southwest China. Copeia, 2007, 913 - 919. http: // dx. doi. org / 10.1643 / 0045 - 8511 (2007) 7 [913: ANSOAA] 2.0. CO; 2" type="journal article" year="2007">Rao &amp; Wilkinson 2007</bibRefCitation>
)
</taxonomicName>
. However, because the tadpole of the new species remains to be discovered, we assigned the new species to this genus based on the morphological and genetic similarity of the adult specimens to those of 
<taxonomicName id="5BFD79A3FF92A2506EFE4A51FE902AE0" box="[247,357,367,392]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="ricketti">
<emphasis id="AE89DE32FF92A2506EFE4A51FE902AE0" box="[247,357,367,392]" italics="true" pageId="7" pageNumber="532">A. ricketti</emphasis>
</taxonomicName>
and 
<taxonomicName id="5BFD79A3FF92A2506F954A51FDDD2AE0" box="[412,552,368,392]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="wuyiensis">
<emphasis id="AE89DE32FF92A2506F954A51FDDD2AE0" box="[412,552,368,392]" italics="true" pageId="7" pageNumber="532">A. wuyiensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9C420220FF92A2506ECE4AB5FCCA29ED" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
<taxonomicName id="5BFD79A3FF92A2506ECE4AB5FE592AC5" box="[199,428,404,429]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="7" pageNumber="532" phylum="Chordata" rank="species" species="albispinus" status="sp. nov.">
<emphasis id="AE89DE32FF92A2506ECE4AB5FE592AC5" box="[199,428,404,429]" italics="true" pageId="7" pageNumber="532">Amolops albispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5BA6349FF92A2506FB44AB5FDD52AC5" box="[445,544,404,429]" pageId="7" pageNumber="532" rank="species">
<emphasis id="AE89DE32FF92A2506FB44AB5FDD52AC5" bold="true" box="[445,544,404,429]" pageId="7" pageNumber="532">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from its congeners by a combination of the following morphological characteristics: (1) presence of white conical spines on the upper and lower lips, loreal and temporal regions, excluding the tympanum; (2) relatively small body size, SVL 
<quantity id="5B05AFC5FF92A2506DA74AFDFBA42A9C" box="[942,1105,476,501]" metricMagnitude="-2" metricUnit="m" metricValue="3.925" metricValueMax="4.24" metricValueMin="3.61" pageId="7" pageNumber="532" unit="mm" value="39.25" valueMax="42.4" valueMin="36.1">36.1–42.4 mm</quantity>
in adult males and 
<quantity id="5B05AFC5FF92A2506B254AFDFF36297F" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="5.09" metricValueMin="4.31" pageId="7" pageNumber="532" unit="mm" value="47.0" valueMax="50.9" valueMin="43.1">43.1–50.9 mm</quantity>
in adult females; (3) dorsal skin of body very rough with numerous raised large warts; (4) dorsal body olivebrown with dark brown blotches; (5) presence of strongly developed vomerine teeth; (6) absence of vocal sacs; (7) absence of tarsal glands; (8) absence of the dorsolateral folds; (9) presence of a circummarginal groove on the disk of the first finger; (10) absence of outer metatarsal tubercles.
</paragraph>
</subSubSection>
<subSubSection id="D4E751ABFF92A25F6ECE49AEFC6A2C14" lastPageId="8" lastPageNumber="533" pageId="7" pageNumber="532" type="description">
<paragraph id="9C420220FF92A2506ECE49AEFB822835" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
<emphasis id="AE89DE32FF92A2506ECE49AEFE0029C0" bold="true" box="[199,501,655,680]" pageId="7" pageNumber="532">
Description of 
<typeStatus id="4346BC82FF92A2506F7649AEFE0529C0" box="[383,496,655,680]" pageId="7" pageNumber="532" type="holotype">Holotype</typeStatus>
:
</emphasis>
Head width approximately equal to head length (HDW/HDL 1.0); snout short (SNT/HDL 0.4) and rounded in profile, projecting beyond lower jaw; nostril closer to tip of snout than eye; loreal region concave; top of head flat; eye large and convex (EYE/HDL 0.3); eye diameter shorter than snout length (EYE/SNT 0.9); canthus rostralis distinct; pineal body barely visible; tympanum small, edge faintly distinct; supratympanic fold broad, from back of eye to shoulder; choanae large; vomerine teeth on well-developed ridges, converging posteriorly; tongue cordiform, deeply notched posteriorly; vocal sacs absent.
</paragraph>
<paragraph id="9C420220FF92A2506ECE4846FF192F78" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">Forelimbs moderately robust; hands moderately long (ML/SVL 0.3); relative finger lengths I&lt;II&lt;IV&lt;III; finger tips on I–IV dilated to wide oval disks with circummarginal grooves, relative width of finger disks I&lt;II&lt;III=IV; nuptial pad on first finger prominent with strongly developed white conical spines; subarticular tubercles prominent, rounded; inner and outer metacarpal tubercle slightly elongated; no finger webbing or lateral fringes.</paragraph>
<paragraph id="9C420220FF92A2506ECE4F3DFCC62FAD" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
Hindlimbs long and robust (TIB/SVL 0.5; PL/SVL 0.5); relative toe lengths I&lt;II&lt;V&lt;III&lt;IV; tips of all toes expanded to well-developed oval discs with circummarginal grooves; subarticular tubercles oval and distinct; inner metatarsal tubercles laterally compressed and pronounced; outer metatarsal tubercles absent; toes fully webbed, webbing formula 
<date id="E84324E0FF92A2506F694FA6FE392FC8" box="[352,460,1159,1184]" pageId="7" pageNumber="532">I1–1II1–2</date>
¾ 
<date id="E84324E0FF92A2506FEB4FA6FD872FC8" box="[482,626,1159,1184]" pageId="7" pageNumber="532">III1–3IV3–1</date>
following 
<bibRefCitation id="F86C7FD1FF92A2506CE64FA9FC602FC8" author="Savage" box="[751,917,1160,1184]" pageId="7" pageNumber="532" refString="Savage, J. M. (1997) Digital webbing formulae for anurans: a refinement. Herpetological Review, 28, 131." type="journal article" year="1997">Savage (1997)</bibRefCitation>
; lateral fringe present; tibio-tarsal articulation reaching snout, when hindlimb stretched alongside of body.
</paragraph>
<paragraph id="9C420220FF92A2506ECE4FEEFAEC2E3D" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">Skin on dorsal surface of head, trunk, and limbs very rough with numerous tubercles and large raised warts; prominent conical spines on the upper and lower lips, loreal and temporal regions, excluding the tympanum; numerous small tubercles and ridges on the throat and ventral surfaces of trunk and limbs; dorsolateral fold absent; posterior part of upper lip swollen; rictal gland prominent and ellipsoidal, posterior to corner of mouth.</paragraph>
<paragraph id="9C420220FF92A2506ECE4E7EFC7E2EF4" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
<emphasis id="AE89DE32FF92A2506ECE4E7EFD962E10" bold="true" box="[199,611,1375,1400]" pageId="7" pageNumber="532">Measurement of holotype (in mm).</emphasis>
SVL 36.8; HDL 13.2, HDW 13.4; SNT 5.2; IOD 3.3; EYE 4.6; TMP 1.5; TEY 1.3; TIB 17.3; ML 11.4; PL 19.2; F2D 1.8; F3D 2.2; T4D 1.4.
</paragraph>
<paragraph id="9C420220FF92A2506ECE4E86FBCE2DD5" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
<emphasis id="AE89DE32FF92A2506ECE4E86FE942EA8" bold="true" box="[199,353,1447,1472]" pageId="7" pageNumber="532">Color in life.</emphasis>
Dorsal surface olive-brown with raised dark brown blotches on the dorsal surface of head and trunk and small dark brown spots on the dorsal surface of fingers, upper arms, tarsi, tibias, thighs and toes; faint dark transverse bars on dorsal surface of fingers, lower arms, tarsi, tibias, thighs and toes; posterior edge of upper lip and rictal gland copper; white conical spines on upper and lower lips, loreal and temporal regions, excluding tympanum; nuptial pad and spines white; dorsal surfaces of discs on fingers copper; copper flecks on the lateral sides of body; ventral surface of the throat, chest, and belly opaque creamy white, some grey flecks on the throat and chest; ventral surface of the hands and feet dark grey; ventral surfaces of the upper arms, lower arms, tarsi, tibias, and thighs pink with copper flecks; rear of thighs copper with dark mottling.
</paragraph>
<paragraph id="9C420220FF92A2506ECE4DE6FE602C40" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
<emphasis id="AE89DE32FF92A2506ECE4DE6FE3A2D88" bold="true" box="[199,463,1735,1760]" pageId="7" pageNumber="532">Color in preservative.</emphasis>
On dorsum, color fades to dark olive with dark brown blotches, transverse bars, and spots; ventral surface yellow with grey mottling on throat and chest; edge of the ventral surface of the belly with a hint of orange (
<figureCitation id="04C61EA5FF92A2506F4A4C2EFE722C40" box="[323,391,1807,1832]" captionStart="FIGURE 3" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[190,1388,201,1904]" captionTargetId="figure@6.[183,1399,195,1915]" captionTargetPageId="6" captionText="FIGURE 3. Dorsolateral view of adult male holotype SYS a 003454 of Amolops albispinus sp. nov. in life; B: ventral view of the holotype in life; C: hand of the holotype in life; D: foot of the holotype in life; E: Dorsal view of the holotype in preserve; F: ventral view of the holotype in preserve; G: close-up of the head of the holotype in life." httpUri="https://zenodo.org/record/260636/files/figure.png" pageId="7" pageNumber="532">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="9C420220FF92A2506ECE4C15FDCA2CD1" blockId="7.[151,1437,151,1977]" pageId="7" pageNumber="532">
<emphasis id="AE89DE32FF92A2506ECE4C15FECE2C25" bold="true" box="[199,315,1844,1869]" pageId="7" pageNumber="532">Variation.</emphasis>
Measurements of 
<typeStatus id="4346BC82FF92A2506C194C14FDB52C25" box="[528,576,1845,1869]" pageId="7" pageNumber="532">type</typeStatus>
series are given in 
<tableCitation id="D17F379BFF92A2506D174C15FC862C24" box="[798,883,1844,1869]" captionStart="TABLE 3" captionStartId="8.[151,239,152,175]" captionTargetBox="[159,1297,234,959]" captionTargetPageId="8" captionText="TABLE 3. Measurements [in mm; mean ± SD (range)] of the type series of Amolops albispinus sp. nov. See Material and methods for abbreviations." httpUri="http://table.plazi.org/id/C88252A8FF9DA25F6E9E4BB9FE402BA7" pageId="7" pageNumber="532" tableUuid="C88252A8FF9DA25F6E9E4BB9FE402BA7">Table 3</tableCitation>
. All specimens were very similar in morphology and color pattern. However, the main diagnostic character of this species (i.e., white conical spines on upper and lower lips, temporal and loreal regions), are obvious on adult males but subtle on adult females, which indicates that it is a secondary sexual character.
</paragraph>
<caption id="C88252A8FF9DA25F6E9E4BB9FE402BA7" ID-Table-UUID="C88252A8FF9DA25F6E9E4BB9FE402BA7" httpUri="http://table.plazi.org/id/C88252A8FF9DA25F6E9E4BB9FE402BA7" pageId="8" pageNumber="533" targetBox="[159,1297,234,959]" targetIsTable="true" targetPageId="8">
<paragraph id="9C420220FF9DA25F6E9E4BB9FE402BA7" blockId="8.[151,1435,151,207]" pageId="8" pageNumber="533">
<emphasis id="AE89DE32FF9DA25F6E9E4BB9FEF62BC7" bold="true" box="[151,259,152,175]" pageId="8" pageNumber="533">TABLE 3</emphasis>
. Measurements [in mm; mean±SD (range)] of the type series of 
<taxonomicName id="5BFD79A3FF9DA25F6DBD4BB6FB702BC7" box="[948,1157,151,175]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="albispinus" status="sp. nov.">
<emphasis id="AE89DE32FF9DA25F6DBD4BB6FB702BC7" box="[948,1157,151,175]" italics="true" pageId="8" pageNumber="533">Amolops albispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5BA6349FF9DA25F6A854BB9FB142BC7" box="[1164,1249,152,175]" pageId="8" pageNumber="533" rank="species">
<emphasis id="AE89DE32FF9DA25F6A854BB9FB142BC7" bold="true" box="[1164,1249,152,175]" pageId="8" pageNumber="533">sp. nov.</emphasis>
</taxonomicNameLabel>
See Material and methods for abbreviations.
</paragraph>
</caption>
<paragraph id="9C420220FF9DA25F6F624BCAFB1E28D7" pageId="8" pageNumber="533">
<table id="EEFDF080FF9D5DA86E964BCBFAE428D7" box="[159,1297,234,959]" gridcols="3" gridrows="18" pageId="8" pageNumber="533">
<tr id="22CD0062FF9D5DA86E964BCBFAE42A68" box="[159,1297,234,256]" gridrow="0" pageId="8" pageNumber="533" rowspan-1="1" rowspan-2="1">
<th id="611C691EFF9D5DA86E964BCBFDB32A68" box="[159,582,234,256]" gridcol="0" gridrow="0" pageId="8" pageNumber="533">
<taxonomicName id="5BFD79A3FF9DA25F6F624BCAFE1B2A68" box="[363,494,234,256]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="albispinus" status="sp. nov.">
<emphasis id="AE89DE32FF9DA25F6F624BCAFE1B2A68" box="[363,494,234,256]" italics="true" pageId="8" pageNumber="533">A. albispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5BA6349FF9DA25F6FFC4BCBFDB32A68" box="[501,582,234,256]" pageId="8" pageNumber="533" rank="species">
<emphasis id="AE89DE32FF9DA25F6FFC4BCBFDB32A68" bold="true" box="[501,582,234,256]" pageId="8" pageNumber="533">sp. nov.</emphasis>
</taxonomicNameLabel>
</th>
</tr>
<tr id="22CD0062FF9D5DA86E964A35FAE42A42" box="[159,1297,276,298]" gridrow="1" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964A35FDB32A42" box="[159,582,276,298]" gridcol="0" gridrow="1" pageId="8" pageNumber="533">Males (n=8)</th>
<td id="611C691EFF9D5DA86CDA4A35FC5C2A42" box="[723,937,276,298]" gridcol="1" gridrow="1" pageId="8" pageNumber="533">Females (n=6)</td>
<td id="611C691EFF9D5DA86A324A35FAE42A42" box="[1083,1297,276,298]" gridcol="2" gridrow="1" pageId="8" pageNumber="533">Juveniles (n=5)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964A1CFAE42A3B" box="[159,1297,317,339]" gridrow="2" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964A1CFDB32A3B" box="[159,582,317,339]" gridcol="0" gridrow="2" pageId="8" pageNumber="533">SVL 40.0±2.2 (36.7–42.4)</th>
<td id="611C691EFF9D5DA86CDA4A1CFC5C2A3B" box="[723,937,317,339]" gridcol="1" gridrow="2" pageId="8" pageNumber="533">48.3±2.8 (43.1–51.9)</td>
<td id="611C691EFF9D5DA86A324A1CFAE42A3B" box="[1083,1297,317,339]" gridcol="2" gridrow="2" pageId="8" pageNumber="533">25.7±5.2 (21.5–33.5)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964A47FAE42A14" box="[159,1297,358,380]" gridrow="3" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964A47FDB32A14" box="[159,582,358,380]" gridcol="0" gridrow="3" pageId="8" pageNumber="533">HDL 14.3±0.7 (13.2–15.1)</th>
<td id="611C691EFF9D5DA86CDA4A47FC5C2A14" box="[723,937,358,380]" gridcol="1" gridrow="3" pageId="8" pageNumber="533">17.4±0.8 (16.2–18.4)</td>
<td id="611C691EFF9D5DA86A324A47FAE42A14" box="[1083,1297,358,380]" gridcol="2" gridrow="3" pageId="8" pageNumber="533">8.2±1.3 (6.9–9.9)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964AB1FAE42ACE" box="[159,1297,400,422]" gridrow="4" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964AB1FDB32ACE" box="[159,582,400,422]" gridcol="0" gridrow="4" pageId="8" pageNumber="533">HDW 15.2±1.1 (13.4–16.9)</th>
<td id="611C691EFF9D5DA86CDA4AB1FC5C2ACE" box="[723,937,400,422]" gridcol="1" gridrow="4" pageId="8" pageNumber="533">17.8±1.2 (16.0–19.5)</td>
<td id="611C691EFF9D5DA86A324AB1FAE42ACE" box="[1083,1297,400,422]" gridcol="2" gridrow="4" pageId="8" pageNumber="533">9.5±2.0 (7.3–12.0)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964A98FAE42AA7" box="[159,1297,441,463]" gridrow="5" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964A98FDB32AA7" box="[159,582,441,463]" gridcol="0" gridrow="5" pageId="8" pageNumber="533">SNT 6.0±0.7 (5.0–7.0)</th>
<td id="611C691EFF9D5DA86CDA4A98FC5C2AA7" box="[723,937,441,463]" gridcol="1" gridrow="5" pageId="8" pageNumber="533">6.5±0.9 (5.8–8.1)</td>
<td id="611C691EFF9D5DA86A324A98FAE42AA7" box="[1083,1297,441,463]" gridcol="2" gridrow="5" pageId="8" pageNumber="533">4.0±0.6 (3.4–4.6)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964AC3FAE42A90" box="[159,1297,482,504]" gridrow="6" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964AC3FDB32A90" box="[159,582,482,504]" gridcol="0" gridrow="6" pageId="8" pageNumber="533">IOD 3.5±0.5 (2.8–4.0)</th>
<td id="611C691EFF9D5DA86CDA4AC3FC5C2A90" box="[723,937,482,504]" gridcol="1" gridrow="6" pageId="8" pageNumber="533">4.0±0.5 (3.2–4.5)</td>
<td id="611C691EFF9D5DA86A324AC3FAE42A90" box="[1083,1297,482,504]" gridcol="2" gridrow="6" pageId="8" pageNumber="533">2.4±0.5 (1.6–3.0)</td>
</tr>
<tr id="22CD0062FF9D5DA86E96492DFAE4294A" box="[159,1297,524,546]" gridrow="7" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E96492DFDB3294A" box="[159,582,524,546]" gridcol="0" gridrow="7" pageId="8" pageNumber="533">EYE 5.0±0.6 (3.9–5.8)</th>
<td id="611C691EFF9D5DA86CDA492DFC5C294A" box="[723,937,524,546]" gridcol="1" gridrow="7" pageId="8" pageNumber="533">5.6±0.4 (5.1–6.3)</td>
<td id="611C691EFF9D5DA86A32492DFAE4294A" box="[1083,1297,524,546]" gridcol="2" gridrow="7" pageId="8" pageNumber="533">3.7±0.9 (2.9–5.1)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964914FAE42923" box="[159,1297,565,587]" gridrow="8" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964914FDB32923" box="[159,582,565,587]" gridcol="0" gridrow="8" pageId="8" pageNumber="533">TMP 2.1±0.3 (1.5–2.5)</th>
<td id="611C691EFF9D5DA86CDA4914FC5C2923" box="[723,937,565,587]" gridcol="1" gridrow="8" pageId="8" pageNumber="533">2.8±0.5 (2.2–3.3)</td>
<td id="611C691EFF9D5DA86A324914FAE42923" box="[1083,1297,565,587]" gridcol="2" gridrow="8" pageId="8" pageNumber="533">1.2±0.4 (0.7–1.8)</td>
</tr>
<tr id="22CD0062FF9D5DA86E96497FFAE4291C" box="[159,1297,606,628]" gridrow="9" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E96497FFDB3291C" box="[159,582,606,628]" gridcol="0" gridrow="9" pageId="8" pageNumber="533">TEY 1.4±0.2 (1.3–1.7)</th>
<td id="611C691EFF9D5DA86CDA497FFC5C291C" box="[723,937,606,628]" gridcol="1" gridrow="9" pageId="8" pageNumber="533">1.8±0.5 (0.8–2.2)</td>
<td id="611C691EFF9D5DA86A32497FFAE4291C" box="[1083,1297,606,628]" gridcol="2" gridrow="9" pageId="8" pageNumber="533">0.7±0.2 (0.5–0.9)</td>
</tr>
<tr id="22CD0062FF9D5DA86E9649A9FAE429F6" box="[159,1297,648,670]" gridrow="10" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E9649A9FDB329F6" box="[159,582,648,670]" gridcol="0" gridrow="10" pageId="8" pageNumber="533">TIB 19.4±1.9 (17.2–22.4)</th>
<td id="611C691EFF9D5DA86CDA49A9FC5C29F6" box="[723,937,648,670]" gridcol="1" gridrow="10" pageId="8" pageNumber="533">22.9±2.1 (21.0–26.4)</td>
<td id="611C691EFF9D5DA86A3249A9FAE429F6" box="[1083,1297,648,670]" gridcol="2" gridrow="10" pageId="8" pageNumber="533">12.9±2.7 (10.0–17.1)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964990FAE429AF" box="[159,1297,689,711]" gridrow="11" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964990FDB329AF" box="[159,582,689,711]" gridcol="0" gridrow="11" pageId="8" pageNumber="533">ML 11.2±0.9 (9.7–12.3)</th>
<td id="611C691EFF9D5DA86CDA4990FC5C29AF" box="[723,937,689,711]" gridcol="1" gridrow="11" pageId="8" pageNumber="533">13.1±0.6 (12.1–13.8)</td>
<td id="611C691EFF9D5DA86A324990FAE429AF" box="[1083,1297,689,711]" gridcol="2" gridrow="11" pageId="8" pageNumber="533">7.9±1.5 (6.5–9.8)</td>
</tr>
<tr id="22CD0062FF9D5DA86E9649FBFAE42998" box="[159,1297,730,752]" gridrow="12" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E9649FBFDB32998" box="[159,582,730,752]" gridcol="0" gridrow="12" pageId="8" pageNumber="533">PL 20.1±1.4 (18.2–22.4)</th>
<td id="611C691EFF9D5DA86CDA49FBFC5C2998" box="[723,937,730,752]" gridcol="1" gridrow="12" pageId="8" pageNumber="533">22.7±1.5 (20.2–24.1)</td>
<td id="611C691EFF9D5DA86A3249FBFAE42998" box="[1083,1297,730,752]" gridcol="2" gridrow="12" pageId="8" pageNumber="533">12.5±2.3 (10.7–16.1)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964825FAE42872" box="[159,1297,772,794]" gridrow="13" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964825FDB32872" box="[159,582,772,794]" gridcol="0" gridrow="13" pageId="8" pageNumber="533">F2D 2.1±0.2 (1.8–2.4)</th>
<td id="611C691EFF9D5DA86CDA4825FC5C2872" box="[723,937,772,794]" gridcol="1" gridrow="13" pageId="8" pageNumber="533">2.4±0.4 (1.8–2.9)</td>
<td id="611C691EFF9D5DA86A324825FAE42872" box="[1083,1297,772,794]" gridcol="2" gridrow="13" pageId="8" pageNumber="533">1.3±0.4 (1.0–1.8)</td>
</tr>
<tr id="22CD0062FF9D5DA86E96480CFAE4282B" box="[159,1297,813,835]" gridrow="14" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E96480CFDB3282B" box="[159,582,813,835]" gridcol="0" gridrow="14" pageId="8" pageNumber="533">F3D 2.5±0.2 (2.2–2.7)</th>
<td id="611C691EFF9D5DA86CDA480CFC5C282B" box="[723,937,813,835]" gridcol="1" gridrow="14" pageId="8" pageNumber="533">2.8±0.3 (2.2–3.2)</td>
<td id="611C691EFF9D5DA86A32480CFAE4282B" box="[1083,1297,813,835]" gridcol="2" gridrow="14" pageId="8" pageNumber="533">1.5±0.4 (1.1–2.2)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964877FAE42804" box="[159,1297,854,876]" gridrow="15" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964877FDB32804" box="[159,582,854,876]" gridcol="0" gridrow="15" pageId="8" pageNumber="533">T4D 1.4±0.2 (1.2–1.7)</th>
<td id="611C691EFF9D5DA86CDA4877FC5C2804" box="[723,937,854,876]" gridcol="1" gridrow="15" pageId="8" pageNumber="533">1.6±0.2 (1.4–1.9)</td>
<td id="611C691EFF9D5DA86A324877FAE42804" box="[1083,1297,854,876]" gridcol="2" gridrow="15" pageId="8" pageNumber="533">0.8±0.2 (0.6–1.1)</td>
</tr>
<tr id="22CD0062FF9D5DA86E9648A1FAE428FE" box="[159,1297,896,918]" gridrow="16" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E9648A1FDB328FE" box="[159,582,896,918]" gridcol="0" gridrow="16" pageId="8" pageNumber="533">HDW/HDL 1.1±0.1 (1.0–1.2)</th>
<td id="611C691EFF9D5DA86CDA48A1FC5C28FE" box="[723,937,896,918]" gridcol="1" gridrow="16" pageId="8" pageNumber="533">1.0±0.0 (1.0–1.1)</td>
<td id="611C691EFF9D5DA86A3248A1FAE428FE" box="[1083,1297,896,918]" gridcol="2" gridrow="16" pageId="8" pageNumber="533">1.2±0.1 (1.0–1.3)</td>
</tr>
<tr id="22CD0062FF9D5DA86E964888FAE428D7" box="[159,1297,937,959]" gridrow="17" pageId="8" pageNumber="533">
<th id="611C691EFF9D5DA86E964888FDB328D7" box="[159,582,937,959]" gridcol="0" gridrow="17" pageId="8" pageNumber="533">HDL/SVL 0.4±0.0 (0.3–0.4)</th>
<td id="611C691EFF9D5DA86CDA4888FC5C28D7" box="[723,937,937,959]" gridcol="1" gridrow="17" pageId="8" pageNumber="533">0.4±0.0 (0.3–0.4)</td>
<td id="611C691EFF9D5DA86A324888FAE428D7" box="[1083,1297,937,959]" gridcol="2" gridrow="17" pageId="8" pageNumber="533">0.3±0.0 (0.3–0.4)</td>
</tr>
</table>
</paragraph>
<paragraph id="9C420220FF9DA25F6ECE4F25FB282EC0" blockId="8.[151,1436,1028,2025]" pageId="8" pageNumber="533">
<emphasis id="AE89DE32FF9DA25F6ECE4F25FE982F75" bold="true" box="[199,365,1028,1053]" pageId="8" pageNumber="533">Comparisons.</emphasis>
Morphologically, 
<taxonomicName id="5BFD79A3FF9DA25F6C454F25FCC52F75" box="[588,816,1028,1053]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="albispinus" status="sp. nov.">
<emphasis id="AE89DE32FF9DA25F6C454F25FCC52F75" box="[588,816,1028,1053]" italics="true" pageId="8" pageNumber="533">Amolops albispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5BA6349FF9DA25F6D364F25FC542F75" box="[831,929,1028,1053]" pageId="8" pageNumber="533" rank="species">
<emphasis id="AE89DE32FF9DA25F6D364F25FC542F75" bold="true" box="[831,929,1028,1053]" pageId="8" pageNumber="533">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="5BFD79A3FF9DA25F6A444F25FB362F75" box="[1101,1219,1028,1053]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="ricketti">
<emphasis id="AE89DE32FF9DA25F6A444F25FB362F75" box="[1101,1219,1028,1053]" italics="true" pageId="8" pageNumber="533">A. ricketti</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6AD34F25FA9B2F74" box="[1242,1390,1028,1052]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="wuyiensis">
<emphasis id="AE89DE32FF9DA25F6AD34F25FA9B2F74" box="[1242,1390,1028,1052]" italics="true" pageId="8" pageNumber="533">A. wuyiensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6B8D4F25FEEE2F28" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="daiyunensis">
<emphasis id="AE89DE32FF9DA25F6B8D4F25FEEE2F28" italics="true" pageId="8" pageNumber="533">A. daiyunensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="5BFD79A3FF9DA25F6F534F09FDD52F28" box="[346,544,1063,1088]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="hongkongensis">
<emphasis id="AE89DE32FF9DA25F6F534F09FDD52F28" box="[346,544,1063,1088]" italics="true" pageId="8" pageNumber="533">A. hongkongensis</emphasis>
</taxonomicName>
(in parenthesis), to which it is most closely related (
<figureCitation id="04C61EA5FF9DA25F6A7B4F06FB422F28" box="[1138,1207,1063,1088]" captionStart="FIGURE 2" captionStartId="5.[151,250,1410,1432]" captionTargetBox="[151,1420,466,1368]" captionTargetId="figure@5.[151,1436,463,1372]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian inference tree derived from partial DNA sequences of the mitochondrial 16 S rRNA and COI genes. Numbers above branches are bootstrap support values for ML / BI inferences respectively." httpUri="https://zenodo.org/record/260635/files/figure.png" pageId="8" pageNumber="533">Fig. 2</figureCitation>
), by the presence of white strongly developed conical spines on upper and lower lips and loreal and temporal regions, excluding tympanum (vs. absence; 
<figureCitation id="04C61EA5FF9DA25F6FA74F4EFE042FE0" box="[430,497,1135,1160]" captionStart="FIGURE 3" captionStartId="6.[151,250,1936,1958]" captionTargetBox="[190,1388,201,1904]" captionTargetId="figure@6.[183,1399,195,1915]" captionTargetPageId="6" captionText="FIGURE 3. Dorsolateral view of adult male holotype SYS a 003454 of Amolops albispinus sp. nov. in life; B: ventral view of the holotype in life; C: hand of the holotype in life; D: foot of the holotype in life; E: Dorsal view of the holotype in preserve; F: ventral view of the holotype in preserve; G: close-up of the head of the holotype in life." httpUri="https://zenodo.org/record/260636/files/figure.png" pageId="8" pageNumber="533">Fig. 3</figureCitation>
), and by the presence of numerous raised large warts on the dorsal skin of the body (vs. dorsal skin without such warts). Further, it differs from 
<taxonomicName id="5BFD79A3FF9DA25F6D304FB5FC5D2FC5" box="[825,936,1172,1197]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="ricketti">
<emphasis id="AE89DE32FF9DA25F6D304FB5FC5D2FC5" box="[825,936,1172,1197]" italics="true" pageId="8" pageNumber="533">A. ricketti</emphasis>
</taxonomicName>
by having a relative small body size at SVL 
<quantity id="5B05AFC5FF9DA25F6E9E4F96FEC82FA7" box="[151,317,1207,1232]" metricMagnitude="-2" metricUnit="m" metricValue="3.9549999999999996" metricValueMax="4.24" metricValueMin="3.67" pageId="8" pageNumber="533" unit="mm" value="39.55" valueMax="42.4" valueMin="36.7">36.7–42.4 mm</quantity>
in adult males (vs. 42.0–60.5), and 
<quantity id="5B05AFC5FF9DA25F6CE84F96FC722FA7" box="[737,903,1207,1232]" metricMagnitude="-2" metricUnit="m" metricValue="4.75" metricValueMax="5.1899999999999995" metricValueMin="4.31" pageId="8" pageNumber="533" unit="mm" value="47.5" valueMax="51.9" valueMin="43.1">43.1–51.9 mm</quantity>
in adult females (vs. 53.5–67.0), and a dorsal color of olive-brown with dark brown blotches (vs. dorsal color olive-brown or brown with light-colored wormlike marks); from 
<taxonomicName id="5BFD79A3FF9DA25F6F7D4E21FDF32E70" box="[372,518,1280,1304]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="wuyiensis">
<emphasis id="AE89DE32FF9DA25F6F7D4E21FDF32E70" box="[372,518,1280,1304]" italics="true" pageId="8" pageNumber="533">A. wuyiensis</emphasis>
</taxonomicName>
by the presence of vomerine teeth (vs. absence), absence of vocal sacs (vs. presence), and white nuptial spines (vs. black); from 
<taxonomicName id="5BFD79A3FF9DA25F6D114E05FC352E55" box="[792,960,1316,1341]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="daiyunensis">
<emphasis id="AE89DE32FF9DA25F6D114E05FCDC2E53" box="[792,809,1316,1339]" italics="true" pageId="8" pageNumber="533">A</emphasis>
. 
<emphasis id="AE89DE32FF9DA25F6D354E05FC352E55" box="[828,960,1316,1341]" italics="true" pageId="8" pageNumber="533">daiyunensis</emphasis>
</taxonomicName>
by the presence of vomerine teeth (vs. absence), absence of vocal sacs (vs. presence), prominent conical nuptial spines (vs. fine, particle-like), and absence of tarsal gland (vs. presence); and from 
<taxonomicName id="5BFD79A3FF9DA25F6CBD4E4DFC8F2EED" box="[692,890,1388,1413]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="hongkongensis">
<emphasis id="AE89DE32FF9DA25F6CBD4E4DFC8F2EED" box="[692,890,1388,1413]" italics="true" pageId="8" pageNumber="533">A. hongkongensis</emphasis>
</taxonomicName>
by the presence of vomerine teeth (vs. absence), absence of vocal sacs (vs. presence), and prominent conical nuptial spines (vs. fine, particle-like).
</paragraph>
<paragraph id="9C420220FF9DA25F6ECE4E94FC6A2C14" blockId="8.[151,1436,1028,2025]" pageId="8" pageNumber="533">
In addition to the presence of white, strongly developed, conical spines on the upper and lower lips and loreal and temporal region, 
<taxonomicName id="5BFD79A3FF9DA25F6F844EF6FD992E98" box="[397,620,1495,1520]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="albispinus" status="sp. nov.">
<emphasis id="AE89DE32FF9DA25F6F844EF6FD992E98" box="[397,620,1495,1520]" italics="true" pageId="8" pageNumber="533">Amolops albispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5BA6349FF9DA25F6C7C4EF6FD272E98" box="[629,722,1495,1520]" pageId="8" pageNumber="533" rank="species">
<emphasis id="AE89DE32FF9DA25F6C7C4EF6FD272E98" bold="true" box="[629,722,1495,1520]" pageId="8" pageNumber="533">sp. nov.</emphasis>
</taxonomicNameLabel>
differs from the remaining 46 species of 
<taxonomicName id="5BFD79A3FF9DA25F6ABD4EF6FAE32E98" box="[1204,1302,1495,1520]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="genus">
<emphasis id="AE89DE32FF9DA25F6ABD4EF6FAE32E98" box="[1204,1302,1495,1520]" italics="true" pageId="8" pageNumber="533">Amolops</emphasis>
</taxonomicName>
as follows: from 
<taxonomicName id="5BFD79A3FF9DA25F6EDD4EDDFE932D7D" box="[212,358,1532,1557]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="akhaorum">
<emphasis id="AE89DE32FF9DA25F6EDD4EDDFE932D7D" box="[212,358,1532,1557]" italics="true" pageId="8" pageNumber="533">A. akhaorum</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6F7A4EDDFDD52D7C" box="[371,544,1532,1556]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="aniqiaoensis">
<emphasis id="AE89DE32FF9DA25F6F7A4EDDFDD52D7C" box="[371,544,1532,1556]" italics="true" pageId="8" pageNumber="533">A. aniqiaoensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6C274EDDFD2D2D7D" box="[558,728,1532,1557]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="archotaphus">
<emphasis id="AE89DE32FF9DA25F6C274EDDFD2D2D7D" box="[558,728,1532,1557]" italics="true" pageId="8" pageNumber="533">A. archotaphus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6CEF4EDDFCAB2D7D" box="[742,862,1532,1557]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="bellulus">
<emphasis id="AE89DE32FF9DA25F6CEF4EDDFCAB2D7D" box="[742,862,1532,1557]" italics="true" pageId="8" pageNumber="533">A. bellulus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6D654EDDFBDD2D7D" box="[876,1064,1532,1557]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="chakrataensis">
<emphasis id="AE89DE32FF9DA25F6D654EDDFBDD2D7D" box="[876,1064,1532,1557]" italics="true" pageId="8" pageNumber="533">A. chakrataensis</emphasis>
</taxonomicName>
, 
<emphasis id="AE89DE32FF9DA25F6A3C4EDDFA612D7D" box="[1077,1428,1532,1557]" italics="true" pageId="8" pageNumber="533">
<taxonomicName id="5BFD79A3FF9DA25F6A3C4EDDFB272D7D" box="[1077,1234,1532,1557]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="chayuensis">A. chayuensis</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6AD44EDDFA612D7D" box="[1245,1428,1532,1557]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="chunganensis">A. chunganensis</taxonomicName>
</emphasis>
, 
<taxonomicName id="5BFD79A3FF9DA25F6E9E4D01FEDC2D50" box="[151,297,1568,1592]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="compotrix">
<emphasis id="AE89DE32FF9DA25F6E9E4D01FEDC2D50" box="[151,297,1568,1592]" italics="true" pageId="8" pageNumber="533">A. compotrix</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6F334D01FE192D50" box="[314,492,1567,1592]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="cremnobatus">
<emphasis id="AE89DE32FF9DA25F6F334D01FE192D50" box="[314,492,1567,1592]" italics="true" pageId="8" pageNumber="533">A. cremnobatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6FF54D01FDAB2D50" box="[508,606,1568,1592]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="cucae">
<emphasis id="AE89DE32FF9DA25F6FF54D01FDAB2D50" box="[508,606,1568,1592]" italics="true" pageId="8" pageNumber="533">A. cucae</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6C664D01FD012D50" box="[623,756,1567,1592]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="gerbillus">
<emphasis id="AE89DE32FF9DA25F6C664D01FD012D50" box="[623,756,1567,1592]" italics="true" pageId="8" pageNumber="533">A. gerbillus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6D0D4D01FC812D50" box="[772,884,1567,1592]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="iriodes">
<emphasis id="AE89DE32FF9DA25F6D0D4D01FC812D50" box="[772,884,1567,1592]" italics="true" pageId="8" pageNumber="533">A. iriodes</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6D8D4D01FBF72D50" box="[900,1026,1568,1592]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="jaunsari">
<emphasis id="AE89DE32FF9DA25F6D8D4D01FBF72D50" box="[900,1026,1568,1592]" italics="true" pageId="8" pageNumber="533">A. jaunsari</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6A1A4D01FB4B2D50" box="[1043,1214,1567,1592]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="kohimaensis">
<emphasis id="AE89DE32FF9DA25F6A1A4D01FB4B2D50" box="[1043,1214,1567,1592]" italics="true" pageId="8" pageNumber="533">A. kohimaensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6AC64D01FA812D50" box="[1231,1396,1567,1592]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="longimanus">
<emphasis id="AE89DE32FF9DA25F6AC64D01FA812D50" box="[1231,1396,1567,1592]" italics="true" pageId="8" pageNumber="533">A. longimanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6B8D4D01FEB72D34" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="mengyangensis">
<emphasis id="AE89DE32FF9DA25F6B8D4D01FEB72D34" italics="true" pageId="8" pageNumber="533">A. mengyangensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6F5D4D65FE3A2D34" box="[340,463,1604,1628]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="minutus">
<emphasis id="AE89DE32FF9DA25F6F5D4D65FE3A2D34" box="[340,463,1604,1628]" italics="true" pageId="8" pageNumber="533">A. minutus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6FE84D65FD862D35" box="[481,627,1604,1629]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="monticola">
<emphasis id="AE89DE32FF9DA25F6FE84D65FD862D35" box="[481,627,1604,1629]" italics="true" pageId="8" pageNumber="533">A. monticola</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6C8C4D65FCB52D35" box="[645,832,1604,1629]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="nyingchiensis">
<emphasis id="AE89DE32FF9DA25F6C8C4D65FCB52D35" box="[645,832,1604,1629]" italics="true" pageId="8" pageNumber="533">A. nyingchiensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="5BFD79A3FF9DA25F6D8C4D65FC1D2D34" box="[901,1000,1604,1628]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="vitrea">
<emphasis id="AE89DE32FF9DA25F6D8C4D65FC1D2D34" box="[901,1000,1604,1628]" italics="true" pageId="8" pageNumber="533">A. vitrea</emphasis>
</taxonomicName>
by the lack of dorsolateral folds (vs. presence); from 
<taxonomicName id="5BFD79A3FF9DA25F6F464D49FE202DE8" box="[335,469,1640,1664]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="formosus">
<emphasis id="AE89DE32FF9DA25F6F464D49FE202DE8" box="[335,469,1640,1664]" italics="true" pageId="8" pageNumber="533">A. formosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6FED4D49FD752DE8" box="[484,640,1639,1664]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="granulosus">
<emphasis id="AE89DE32FF9DA25F6FED4D49FD752DE8" box="[484,640,1639,1664]" italics="true" pageId="8" pageNumber="533">A. granulosus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6C864D49FCCB2DE8" box="[655,830,1640,1664]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="jinjiangensis">
<emphasis id="AE89DE32FF9DA25F6C864D49FCCB2DE8" box="[655,830,1640,1664]" italics="true" pageId="8" pageNumber="533">A. jinjiangensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6D444D49FBF32DE8" box="[845,1030,1639,1664]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="kangtingensis">
<emphasis id="AE89DE32FF9DA25F6D444D49FBF32DE8" box="[845,1030,1639,1664]" italics="true" pageId="8" pageNumber="533">A. kangtingensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6A1C4D49FB2E2DE8" box="[1045,1243,1639,1664]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="liangshanensis">
<emphasis id="AE89DE32FF9DA25F6A1C4D49FB2E2DE8" box="[1045,1243,1639,1664]" italics="true" pageId="8" pageNumber="533">A. liangshanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6AE34D49FA802DE8" box="[1258,1397,1639,1664]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="lifanensis">
<emphasis id="AE89DE32FF9DA25F6AE34D49FA802DE8" box="[1258,1397,1639,1664]" italics="true" pageId="8" pageNumber="533">A. lifanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6B8D4D49FF0E2DCD" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="loloensis">
<emphasis id="AE89DE32FF9DA25F6B8D4D49FF0E2DCD" italics="true" pageId="8" pageNumber="533">A. loloensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6F184DADFE432DCC" box="[273,438,1676,1700]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="mantzorum">
<emphasis id="AE89DE32FF9DA25F6F184DADFE432DCC" box="[273,438,1676,1700]" italics="true" pageId="8" pageNumber="533">A. mantzorum</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6FC54DADFD862DCD" box="[460,627,1676,1701]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="nidorbellus">
<emphasis id="AE89DE32FF9DA25F6FC54DADFD862DCD" box="[460,627,1676,1701]" italics="true" pageId="8" pageNumber="533">A. nidorbellus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6C834DADFC932DCD" box="[650,870,1676,1701]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="tuberodepressus">
<emphasis id="AE89DE32FF9DA25F6C834DADFC932DCD" box="[650,870,1676,1701]" italics="true" pageId="8" pageNumber="533">A. tuberodepressus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="5BFD79A3FF9DA25F6DBD4DADFB792DCD" box="[948,1164,1676,1701]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="viridimaculatus">
<emphasis id="AE89DE32FF9DA25F6DBD4DADFB792DCD" box="[948,1164,1676,1701]" italics="true" pageId="8" pageNumber="533">A. viridimaculatus</emphasis>
</taxonomicName>
by the presence of a circummarginal groove on disk of first finger (vs. absence); from 
<taxonomicName id="5BFD79A3FF9DA25F6D8B4D91FBFD2DA0" box="[898,1032,1711,1736]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="afghanus">
<emphasis id="AE89DE32FF9DA25F6D8B4D91FBFD2DA0" box="[898,1032,1711,1736]" italics="true" pageId="8" pageNumber="533">A. afghanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6A114D91FB432DA0" box="[1048,1206,1712,1736]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="assamensis">
<emphasis id="AE89DE32FF9DA25F6A114D91FB432DA0" box="[1048,1206,1712,1736]" italics="true" pageId="8" pageNumber="533">A. assamensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6ACC4D91FA812DA0" box="[1221,1396,1711,1736]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="himalayanus">
<emphasis id="AE89DE32FF9DA25F6ACC4D91FA812DA0" box="[1221,1396,1711,1736]" italics="true" pageId="8" pageNumber="533">A. himalayanus</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6B8D4D91FEAD2D85" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="indoburmanensis">
<emphasis id="AE89DE32FF9DA25F6B8D4D91FEAD2D85" italics="true" pageId="8" pageNumber="533">A. indoburmanensis</emphasis>
</taxonomicName>
, 
<emphasis id="AE89DE32FF9DA25F6F634DF5FDE92D84" box="[362,540,1748,1772]" italics="true" pageId="8" pageNumber="533">
<taxonomicName id="5BFD79A3FF9DA25F6F634DF5FDED2D84" box="[362,536,1748,1772]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="marmoratus">A. marmoratus</taxonomicName>
,
</emphasis>
and 
<taxonomicName id="5BFD79A3FF9DA25F6C524DF5FCDF2D85" box="[603,810,1748,1773]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="spinapectoralis">
<emphasis id="AE89DE32FF9DA25F6C524DF5FCDF2D85" box="[603,810,1748,1773]" italics="true" pageId="8" pageNumber="533">A. spinapectoralis</emphasis>
</taxonomicName>
by the absence of vocal sacs (vs. presence); from 
<taxonomicName id="5BFD79A3FF9DA25F6B8D4DF5FF1B2C78" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="daorum">
<emphasis id="AE89DE32FF9DA25F6B8D4DF5FF1B2C78" italics="true" pageId="8" pageNumber="533">A. daorum</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6F094DD9FE5C2C78" box="[256,425,1783,1808]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="hainanensis">
<emphasis id="AE89DE32FF9DA25F6F094DD9FE5C2C78" box="[256,425,1783,1808]" italics="true" pageId="8" pageNumber="533">A. hainanensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6FB34DD9FDD12C78" box="[442,548,1783,1808]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="panhai">
<emphasis id="AE89DE32FF9DA25F6FB34DD9FDD12C78" box="[442,548,1783,1808]" italics="true" pageId="8" pageNumber="533">A panhai</emphasis>
</taxonomicName>
, and 
<taxonomicName id="5BFD79A3FF9DA25F6C604DD9FD1E2C78" box="[617,747,1784,1808]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="torrentis">
<emphasis id="AE89DE32FF9DA25F6C604DD9FD1E2C78" box="[617,747,1784,1808]" italics="true" pageId="8" pageNumber="533">A. torrentis</emphasis>
</taxonomicName>
by the presence of strongly developed vomerine teeth (vs. vomerine teeth weak or absent); from 
<taxonomicName id="5BFD79A3FF9DA25F6C444C3DFCF72C5D" box="[589,770,1820,1845]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="caelumnoctis">
<emphasis id="AE89DE32FF9DA25F6C444C3DFCF72C5D" box="[589,770,1820,1845]" italics="true" pageId="8" pageNumber="533">A. caelumnoctis</emphasis>
</taxonomicName>
, 
<taxonomicName id="5BFD79A3FF9DA25F6D184C3DFC402C5D" box="[785,949,1820,1845]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="medogensis">
<emphasis id="AE89DE32FF9DA25F6D184C3DFC402C5D" box="[785,949,1820,1845]" italics="true" pageId="8" pageNumber="533">A. medogensis</emphasis>
</taxonomicName>
, and 
<taxonomicName id="5BFD79A3FF9DA25F6DFF4C3DFB462C5D" box="[1014,1203,1820,1845]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="splendissimus">
<emphasis id="AE89DE32FF9DA25F6DFF4C3DFB462C5D" box="[1014,1203,1820,1845]" italics="true" pageId="8" pageNumber="533">A. splendissimus</emphasis>
</taxonomicName>
by a dorsal color of olive-brown with dark brown blotches (vs. dorsal color of green or bright yellow); and from 
<taxonomicName id="5BFD79A3FF9DA25F6AB54C61FABE2C30" box="[1212,1355,1855,1880]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="larutensis">
<emphasis id="AE89DE32FF9DA25F6AB54C61FABE2C30" box="[1212,1355,1855,1880]" italics="true" pageId="8" pageNumber="533">A. larutensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="5BFD79A3FF9DA25F6B8D4C61FEF12C15" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="kaulbacki">
<emphasis id="AE89DE32FF9DA25F6B8D4C61FEF12C15" italics="true" pageId="8" pageNumber="533">A. kaulbacki</emphasis>
</taxonomicName>
by the absence of outer metatarsal tubercles (vs. presence).
</paragraph>
</subSubSection>
<subSubSection id="D4E751ABFF9DA25F6ECE4CA6FDBC2C80" pageId="8" pageNumber="533" type="etymology">
<paragraph id="9C420220FF9DA25F6ECE4CA6FDBC2C80" blockId="8.[151,1436,1028,2025]" pageId="8" pageNumber="533">
<emphasis id="AE89DE32FF9DA25F6ECE4CA6FEBB2CC8" bold="true" box="[199,334,1927,1952]" pageId="8" pageNumber="533">Etymology.</emphasis>
The specific name, 
<taxonomicName id="5BFD79A3FF9DA25F6C384CA6FD512CC8" box="[561,676,1927,1952]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="8" pageNumber="533" phylum="Chordata" rank="species" species="albispinus">
<emphasis id="AE89DE32FF9DA25F6C384CA6FD512CC8" box="[561,676,1927,1952]" italics="true" pageId="8" pageNumber="533">albispinus</emphasis>
</taxonomicName>
, refers to the “white spines” on the upper and lower lips, and loreal and temporal regions, which are the diagnostic features of this new species. As an English common name we suggest “White-spined Cascade Frog”.
</paragraph>
</subSubSection>
<subSubSection id="D4E751ABFF9CA25E6ECE4BB6FCDA2A28" pageId="9" pageNumber="534" type="distribution">
<paragraph id="9C420220FF9CA25E6ECE4BB6FCDA2A28" blockId="9.[151,1436,151,320]" pageId="9" pageNumber="534">
<emphasis id="AE89DE32FF9CA25E6ECE4BB6FE032BD8" bold="true" box="[199,502,151,176]" pageId="9" pageNumber="534">Distribution and ecology.</emphasis>
Currently, 
<taxonomicName id="5BFD79A3FF9CA25E6C714BB9FCF92BD8" box="[632,780,151,176]" class="Amphibia" family="Ranidae" genus="Amolops" kingdom="Animalia" order="Anura" pageId="9" pageNumber="534" phylum="Chordata" rank="species" species="albispinus" status="sp. nov.">
<emphasis id="AE89DE32FF9CA25E6C714BB9FCF92BD8" box="[632,780,151,176]" italics="true" pageId="9" pageNumber="534">A. albispinus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B5BA6349FF9CA25E6D1C4BB6FC842BD8" box="[789,881,151,176]" pageId="9" pageNumber="534" rank="species">
<emphasis id="AE89DE32FF9CA25E6D1C4BB6FC842BD8" bold="true" box="[789,881,151,176]" pageId="9" pageNumber="534">sp. nov.</emphasis>
</taxonomicNameLabel>
is known from the 
<typeStatus id="4346BC82FF9CA25E6A5C4BB9FB702BD8" box="[1109,1157,152,176]" pageId="9" pageNumber="534">type</typeStatus>
locality of Mt. Wutong, and from Mt. Paiya, which is 
<quantity id="5B05AFC5FF9CA25E6FE94B9DFDD32BBD" box="[480,550,188,213]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="534" unit="km" value="30.0">30 km</quantity>
from Mt. Wutong, in Shenzhen City, 
<collectingRegion id="5E39CCC2FF9CA25E6DC04B9DFB4C2BBD" box="[969,1209,188,213]" country="China" name="Guangdong" pageId="9" pageNumber="534">Guangdong Province</collectingRegion>
, 
<collectingCountry id="E4EA42B0FF9CA25E6ACA4B9DFAFF2BBD" box="[1219,1290,188,213]" name="China" pageId="9" pageNumber="534">China</collectingCountry>
. This species is common in Mt. Wutong throughout the year, whereas, it was observed to be rare in Mt. Paiya (only one specimen (SYS a002436) found). It inhabits low to mid-elevation (
<quantity id="5B05AFC5FF9CA25E6D214A25FC6B2A74" box="[808,926,260,285]" metricMagnitude="2" metricUnit="m" metricValue="2.8" metricValueMax="5.0" metricValueMin="0.6" pageId="9" pageNumber="534" unit="m" value="280.0" valueMax="500.0" valueMin="60.0">60–500 m</quantity>
) rocky, fast-flowing streams surrounding by moist subtropical secondary evergreen broadleaved forests.
</paragraph>
</subSubSection>
</treatment>
</document>