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<document id="E0DF412CF43875EEF53FB8C0B4565082" ID-CLB-Dataset="24604" ID-DOI="10.11646/zootaxa.4711.2.6" ID-GBIF-Dataset="57cfb289-89a3-48b0-901c-c76fb029d2c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3577059" ID-ZooBank="0A02D6A2-9DF4-415D-9A1C-51B313F59998" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576479411414" checkinUser="plazi" docAuthor="Li, Xu, Che, Xing-Jin &amp; Zhou, Wei" docDate="2019" docId="03D247023911FFE3FF69FDA431FFFD2E" docLanguage="en" docName="zootaxa.4711.2.6.pdf" docOrigin="Zootaxa 4711 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Homatula cryptoclathrata Li &amp; Che &amp; Zhou 2019, sp. nov." docType="treatment" docVersion="7" lastPageNumber="341" masterDocId="FFEB3F7A391BFFE8FFFEFFA4311DFF85" masterDocTitle="Loaches of Homatula (Teleostei: Nemacheilidae) from the upper Salween River in Yunnan, China with description of three new species" masterLastPageNumber="348" masterPageNumber="330" pageNumber="340" updateTime="1698774435811" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="A81B5CFCCD22B8665225A85428BC42B5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D616E728C44B29B3AA51ED0DB9FF49E3">
<mods:title id="CCF6567360258B1D9FF88CC62A85E07B">Loaches of Homatula (Teleostei: Nemacheilidae) from the upper Salween River in Yunnan, China with description of three new species</mods:title>
</mods:titleInfo>
<mods:name id="F521687AC6F68AA7CECFA5E458B1123C" type="personal">
<mods:role id="5B7D2421BFD73915E80167B9A36D39D5">
<mods:roleTerm id="5F82D4CD6E23271EB650834E81880DEB">Author</mods:roleTerm>
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<mods:namePart id="A6F2F2A4AAB6C2B8243B7E36EE5E759E">Li, Xu</mods:namePart>
</mods:name>
<mods:name id="113D90DB53AC70D445E3BF96A50AC505" type="personal">
<mods:role id="CDF83B7196824A2D259743CE42A5D064">
<mods:roleTerm id="0AFF45D80BD1D40A375F3F7B2F729B2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E8166C6D76FF881BE4DF9EC99DE045E8">Che, Xing-Jin</mods:namePart>
</mods:name>
<mods:name id="7F2C52C81F9880CE986A3A3705D344C3" type="personal">
<mods:role id="D990D93D053529C40B29CE54419CD63E">
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<mods:namePart id="0A90EF658C1B398EBEE17BB619AB9793">Zhou, Wei</mods:namePart>
</mods:name>
<mods:typeOfResource id="CDFAF27305EFA070C4A3487B466AFC9A">text</mods:typeOfResource>
<mods:relatedItem id="7EA4E6287FBDF2EE0262BF84A02F059A" type="host">
<mods:titleInfo id="9CFB09F51728DAC1489DB09CF23A35FF">
<mods:title id="0995632263D2A2AD312623E214503F98">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="780A3A7A3FC666659B63537A1FF31177">
<mods:date id="9E89CCBAC6CA74A3688BECBE18B22614">2019</mods:date>
<mods:detail id="15D542537AB7074DCAFA249A6E4650D8" type="pubDate">
<mods:number id="F67E434456484FC5E92D2F150E978932">2019-12-16</mods:number>
</mods:detail>
<mods:detail id="416D84309AFEB7CD6A18F622B4E7F7CD" type="volume">
<mods:number id="0B1E79F3521A21A0BDCD803DC2B5D8A8">4711</mods:number>
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<mods:detail id="3226F950CBBBE4219AB6538336209AFE" type="issue">
<mods:number id="22DBE88BE2A13A3934D3462C40CA065F">2</mods:number>
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<mods:start id="6B6D0ED917A69948F962F1CBE2D18CB5">330</mods:start>
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<mods:classification id="5F94C2EB94B14DE2D902AC48F100C962">journal article</mods:classification>
<mods:identifier id="DBF9F490C98F9A85B2D43300E9C6D9FD" type="CLB-Dataset">24604</mods:identifier>
<mods:identifier id="7541C30F2FC6498382D6406E676D1E03" type="DOI">10.11646/zootaxa.4711.2.6</mods:identifier>
<mods:identifier id="646275ECE90088B9C96F823BD667EF79" type="GBIF-Dataset">57cfb289-89a3-48b0-901c-c76fb029d2c7</mods:identifier>
<mods:identifier id="D0FF049807A14775CD734901B6688747" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DCDCD06864DFA7087665A145EB1B0E44" type="Zenodo-Dep">3577059</mods:identifier>
<mods:identifier id="FBC27862D65FA9A7503EED1D928F9BC1" type="ZooBank">0A02D6A2-9DF4-415D-9A1C-51B313F59998</mods:identifier>
</mods:mods>
<treatment id="03D247023911FFE3FF69FDA431FFFD2E" ID-GBIF-Taxon="160747845" LSID="urn:lsid:plazi:treatment:03D247023911FFE3FF69FDA431FFFD2E" httpUri="http://treatment.plazi.org/id/03D247023911FFE3FF69FDA431FFFD2E" lastPageId="11" lastPageNumber="341" pageId="10" pageNumber="340">
<subSubSection id="C361A59F3911FFE2FF69FDA4332BFD9E" box="[151,566,512,539]" pageId="10" pageNumber="340" type="nomenclature">
<paragraph id="8BC4F6143911FFE2FF69FDA4332BFD9E" blockId="10.[151,566,512,575]" box="[151,566,512,539]" pageId="10" pageNumber="340">
<heading id="D08C41783911FFE2FF69FDA4332BFD9E" bold="true" box="[151,566,512,539]" fontSize="11" level="1" pageId="10" pageNumber="340" reason="1">
<emphasis id="B90F2A063911FFE2FF69FDA4332BFD9E" bold="true" box="[151,566,512,539]" pageId="10" pageNumber="340">
<taxonomicName id="4C7B8D973911FFE2FF69FDA430CCFD9E" authority="Li &amp; Che &amp; Zhou, 2019" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[151,465,512,539]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="cryptoclathrata" status="sp. nov.">
<emphasis id="B90F2A063911FFE2FF69FDA430CCFD9E" bold="true" box="[151,465,512,539]" italics="true" pageId="10" pageNumber="340">Homatula cryptoclathrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C977D3911FFE2FE26FDA5332BFD9E" box="[472,566,513,539]" pageId="10" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3911FFE2FF69FD803050FDBA" box="[151,333,548,575]" pageId="10" pageNumber="340" type="description">
<paragraph id="8BC4F6143911FFE2FF69FD803050FDBA" blockId="10.[151,566,512,575]" box="[151,333,548,575]" pageId="10" pageNumber="340">
(
<figureCitation id="1340EA913911FFE2FF5EFD8031EAFDBA" box="[160,247,548,575]" captionStart="FIGURE 2" captionStartId="5.[151,250,932,957]" captionTargetBox="[155,1432,320,847]" captionTargetId="figure@5.[151,1436,271,892]" captionTargetPageId="5" captionText="FIGURE 2. Relationship between anterior end of adipose crest and anal-fin base of Homatula. (A) H. anteridorsalis, SWFC 1610019, paratype, 108.1 mm SL; (B) H. cryptoclathrata, SWFC 1002001, holotype, 138.1 mm SL; (C) H. nigra, SWFC 1901005, holotype, 99.8 mm SL; (D) H. pycnolepis, SWFC 0612085, 99.6 mm SL; (E) H. laxiclathra, SWFC 1810010, 97.0 mm SL); (F) H. nanpanjiangensis, SWFC 1903001, 97.1 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3577063" httpUri="https://zenodo.org/record/3577063/files/figure.png" pageId="10" pageNumber="340">Fig. 2B</figureCitation>
,
<figureCitation id="1340EA913911FFE2FEFDFD803058FDBA" box="[259,325,548,575]" captionStart="FIGURE 9" captionStartId="11.[151,250,1507,1532]" captionTargetBox="[151,1435,697,1487]" captionTargetId="figure@11.[151,1435,697,1487]" captionTargetPageId="11" captionText="FIGURE 9. Homatula cryptoclathrata, SWFC 1002001, holotype, 138.1 mm SL; China: Yunnan: Baoshan City: Changning Co.: Gengga Town: Mangping Village." figureDoi="http://doi.org/10.5281/zenodo.3577079" httpUri="https://zenodo.org/record/3577079/files/figure.png" pageId="10" pageNumber="340">Fig. 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3911FFE2FF69FDC93439FD4B" pageId="10" pageNumber="340" type="materials_examined">
<paragraph id="8BC4F6143911FFE2FF69FDC93542FD2E" blockId="10.[151,1437,620,2015]" pageId="10" pageNumber="340">
<materialsCitation id="3B13FC493911FFE2FF69FDC93542FD2E" ID-GBIF-Occurrence="2512877916" collectingDate="2019-02-20" country="China" county="Changning County" latitude="24.328514" location="Gengga Town" longLatPrecision="1" longitude="99.537605" municipality="Baoshan City" pageId="10" pageNumber="340" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B90F2A063911FFE2FF69FDC93016FD02" bold="true" box="[151,267,621,647]" pageId="10" pageNumber="340">
<typeStatus id="54C048B63911FFE2FF69FDC9301BFD02" box="[151,262,621,647]" pageId="10" pageNumber="340" type="holotype">Holotype</typeStatus>
.
</emphasis>
SWFC
<quantity id="4C835BF13911FFE2FE90FDC93347FD02" box="[366,602,621,647]" metricMagnitude="6" metricUnit="m" metricValue="1.0020011381" pageId="10" pageNumber="340" unit="mm" value="1.0020011381E9">1002001, 138.1 mm</quantity>
SL;
<collectingCountry id="F36CB6843911FFE2FD67FDC833FDFD03" box="[665,736,620,646]" name="China" pageId="10" pageNumber="340">China</collectingCountry>
:
<collectingRegion id="49BF38F63911FFE2FD10FDC93255FD02" box="[750,840,621,647]" country="China" name="Yunnan" pageId="10" pageNumber="340">Yunnan</collectingRegion>
:
<collectingMunicipality id="6BA06C6E3911FFE2FCA9FDC832E5FD03" box="[855,1016,620,646]" pageId="10" pageNumber="340">Baoshan City</collectingMunicipality>
:
<collectingCounty id="62A58E983911FFE2FBF9FDC835F9FD02" box="[1031,1252,620,647]" pageId="10" pageNumber="340">Changning County</collectingCounty>
:
<location id="8EA4A0CF3911FFE2FB0DFDC93484FD02" LSID="urn:lsid:plazi:treatment:03D247023911FFE3FF69FDA431FFFD2E:8EA4A0CF3911FFE2FB0DFDC93484FD02" box="[1267,1433,621,647]" country="China" county="Changning County" latitude="24.328514" longLatPrecision="1" longitude="99.537605" municipality="Baoshan City" name="Gengga Town" pageId="10" pageNumber="340" stateProvince="Yunnan">Gengga Town</location>
:
<location id="8EA4A0CF3911FFE2FF69FD343049FD2F" LSID="urn:lsid:plazi:treatment:03D247023911FFE3FF69FDA431FFFD2E:8EA4A0CF3911FFE2FF69FD343049FD2F" box="[151,340,656,682]" country="China" county="Changning County" latitude="24.328514" longLatPrecision="1" longitude="99.537605" municipality="Baoshan City" name="Manging Village" pageId="10" pageNumber="340" stateProvince="Yunnan">Manging Village</location>
(
<geoCoordinate id="EE4F90D33911FFE2FE9DFD343316FD2E" box="[355,523,656,683]" degrees="24" direction="north" minutes="19" orientation="latitude" pageId="10" pageNumber="340" precision="1" seconds="42.65" value="24.328514">24°19'42.65&quot;N</geoCoordinate>
<geoCoordinate id="EE4F90D33911FFE2FDECFD3433A7FD2E" box="[530,698,656,683]" degrees="99" direction="east" minutes="32" orientation="longitude" pageId="10" pageNumber="340" precision="1" seconds="15.39" value="99.537605">99°32'15.39&quot;E</geoCoordinate>
); collected by XJ.
<location id="8EA4A0CF3911FFE2FC74FD3432A7FD2F" LSID="urn:lsid:plazi:treatment:03D247023911FFE3FF69FDA431FFFD2E:8EA4A0CF3911FFE2FC74FD3432A7FD2F" box="[906,954,656,682]" country="China" county="Changning County" latitude="24.328514" longLatPrecision="1" longitude="99.537605" municipality="Baoshan City" name="Che" pageId="10" pageNumber="340" stateProvince="Yunnan">Che</location>
,
<date id="FFC5D0D43911FFE2FC3BFD353541FD2E" box="[965,1116,656,683]" pageId="10" pageNumber="340" value="2019-02-20">
<collectingDate id="EF81293C3911FFE2FC3BFD353541FD2E" box="[965,1116,656,683]" pageId="10" pageNumber="340" value="2019-02-20">20 Feb. 2019</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC4F6143911FFE2FF39FD113439FD4B" blockId="10.[151,1437,620,2015]" box="[199,1316,692,719]" pageId="10" pageNumber="340">
<materialsCitation id="3B13FC493911FFE2FF39FD11343DFD4B" ID-GBIF-Occurrence="2512877917" box="[199,1312,692,719]" collectingDate="2019-02-20" country="China" county="Changning County" latitude="24.328514" location="Gengga Town" longLatPrecision="1" longitude="99.537605" municipality="Baoshan City" pageId="10" pageNumber="340" specimenCount="10" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="B90F2A063911FFE2FF39FD11305BFD4A" bold="true" box="[199,326,693,719]" pageId="10" pageNumber="340">
<typeStatus id="54C048B63911FFE2FF39FD11305CFD4A" box="[199,321,693,719]" pageId="10" pageNumber="340" type="paratype">Paratypes</typeStatus>
.
</emphasis>
SWFC 1902002-19020011,
<specimenCount id="9D7D3D9D3911FFE2FD76FD1133D2FD4A" box="[648,719,693,719]" pageId="10" pageNumber="340" type="generic">10 ex.</specimenCount>
,
<quantity id="4C835BF13911FFE2FD25FD10329BFD4A" box="[731,902,692,719]" metricMagnitude="-2" metricUnit="m" metricValue="8.165" metricValueMax="11.01" metricValueMin="5.32" pageId="10" pageNumber="340" unit="mm" value="81.65" valueMax="110.1" valueMin="53.2">53.2-110.1 mm</quantity>
SL; other data same as the holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3911FFE2FF39FD7C33A9FBFA" pageId="10" pageNumber="340" type="diagnosis">
<paragraph id="8BC4F6143911FFE2FF39FD7C33A9FBFA" blockId="10.[151,1437,620,2015]" pageId="10" pageNumber="340">
<emphasis id="B90F2A063911FFE2FF39FD7C305FFD76" bold="true" box="[199,322,728,755]" pageId="10" pageNumber="340">Diagnosis.</emphasis>
<taxonomicName id="4C7B8D973911FFE2FEB4FD7C336CFD77" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[330,625,728,754]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="cryptoclathrata">
<emphasis id="B90F2A063911FFE2FEB4FD7C336CFD77" box="[330,625,728,754]" italics="true" pageId="10" pageNumber="340">Homatula cryptoclathrata</emphasis>
</taxonomicName>
differs from other species of densely-scaled group in
<taxonomicName id="4C7B8D973911FFE2FB23FD7C3450FD77" authorityName="Nichols" authorityYear="1925" box="[1245,1357,728,754]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="genus">
<emphasis id="B90F2A063911FFE2FB23FD7C3450FD77" box="[1245,1357,728,754]" italics="true" pageId="10" pageNumber="340">Homatula</emphasis>
</taxonomicName>
by the following combination of characters: bars on flank across lateral line equal to or greater than 25 (vs. equal to or less than
<quantity id="4C835BF13911FFE2FF31FC853015FCBF" box="[207,264,800,827]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="10" pageNumber="340" unit="in" value="20.0">20 in</quantity>
<taxonomicName id="4C7B8D973911FFE2FEF1FC8530DEFCBF" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[271,451,800,826]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A063911FFE2FEF1FC8530DEFCBF" box="[271,451,800,826]" italics="true" pageId="10" pageNumber="340">H. acuticephala</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B8D973911FFE2FE07FC853392FCBF" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[505,655,800,826]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A063911FFE2FE07FC853392FCBF" box="[505,655,800,826]" italics="true" pageId="10" pageNumber="340">H. erhaiensis</emphasis>
</taxonomicName>
); bars on flank vertical along lateral line (vs. no marks on flank in
<taxonomicName id="4C7B8D973911FFE2FA7FFC8531C9FCDB" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A063911FFE2FA7FFC8531C9FCDB" italics="true" pageId="10" pageNumber="340">H. nigra</emphasis>
</taxonomicName>
); no free protrusions between the pelvic fins (vs. a pair of free protrusions between the pelvic fins in
<taxonomicName id="4C7B8D973911FFE2FAB7FCE131F5FC07" authorityName="Hu &amp; Zhang" authorityYear="2010" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A063911FFE2FAB7FCE131F5FC07" italics="true" pageId="10" pageNumber="340">H. pycnolepis</emphasis>
</taxonomicName>
); gill opening smaller, its upper angle level with lower edge of eye (vs. gill opening larger, its upper angle aligned with middle point of eye in
<taxonomicName id="4C7B8D973911FFE2FDDBFC2933C4FC23" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[549,729,908,934]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A063911FFE2FDDBFC2933C4FC23" box="[549,729,908,934]" italics="true" pageId="10" pageNumber="340">H. anguillioides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B8D973911FFE2FD18FC293284FC23" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[742,921,908,934]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A063911FFE2FD18FC293284FC23" box="[742,921,908,934]" italics="true" pageId="10" pageNumber="340">H. acuticephala</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B8D973911FFE2FC59FC293552FC23" authorityName="Endruweit, Min &amp; Yang" authorityYear="2018" box="[935,1103,908,934]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="coccinocola">
<emphasis id="B90F2A063911FFE2FC59FC293552FC23" box="[935,1103,908,934]" italics="true" pageId="10" pageNumber="340">H. coccinocola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B8D973911FFE2FBA3FC2935EFFC23" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[1117,1266,908,934]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A063911FFE2FBA3FC2935EFFC23" box="[1117,1266,908,934]" italics="true" pageId="10" pageNumber="340">H. erhaiensis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7B8D973911FFE2FAD0FC2931D1FC4F" authorityName="Hu &amp; Zhang" authorityYear="2010" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A063911FFE2FAD0FC2931D1FC4F" italics="true" pageId="10" pageNumber="340">H. pycnolepis</emphasis>
</taxonomicName>
) (
<figureCitation id="1340EA913911FFE2FF18FC143036FC4E" box="[230,299,944,971]" captionStart="FIGURE 5" captionStartId="7.[151,250,559,584]" captionTargetBox="[166,1421,174,540]" captionTargetId="figure@7.[166,1421,174,540]" captionTargetPageId="7" captionText="FIGURE 5. Gill openings of Homatula. (A) Larger gill opening; upper angle aligned with middle point of eye (H. pycnolepis, SWFC 0612085, 99.6 mm SL); (B) smaller gill opening; upper angle level with lower edge of eye (H. cryptoclathrata, SWFC 1610006, holotype, 121.6 mm SL)." figureDoi="http://doi.org/10.5281/zenodo.3577071" httpUri="https://zenodo.org/record/3577071/files/figure.png" pageId="10" pageNumber="340">Fig. 5</figureCitation>
); with pelvic axillary lobe (vs. without pelvic axillary lobe in
<taxonomicName id="4C7B8D973911FFE2FC14FC153583FC4F" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[1002,1182,944,970]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A063911FFE2FC14FC153583FC4F" box="[1002,1182,944,970]" italics="true" pageId="10" pageNumber="340">H. acuticephala</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B8D973911FFE2FB28FC153496FC4F" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[1238,1419,944,970]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A063911FFE2FB28FC153496FC4F" box="[1238,1419,944,970]" italics="true" pageId="10" pageNumber="340">H. anguillioides</emphasis>
</taxonomicName>
); marked notch on lower jaw (vs. lower jaw spoon-shaped and lacking notch in
<taxonomicName id="4C7B8D973911FFE2FBF7FC713582FC6B" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" box="[1033,1183,980,1006]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="erhaiensis">
<emphasis id="B90F2A063911FFE2FBF7FC713582FC6B" box="[1033,1183,980,1006]" italics="true" pageId="10" pageNumber="340">H. erhaiensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B8D973911FFE2FB28FC713496FC6B" authorityName="Min, Yang &amp; Chen" authorityYear="2012" box="[1238,1419,980,1006]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="wuliangensis">
<emphasis id="B90F2A063911FFE2FB28FC713496FC6B" box="[1238,1419,980,1006]" italics="true" pageId="10" pageNumber="340">H. wuliangensis</emphasis>
</taxonomicName>
); dorsal-fin base longer than length of longest branched dorsal-fin ray (vs. shorter than longest branched dorsal-fin ray in
<taxonomicName id="4C7B8D973911FFE2FF1DFBB930B9FBB3" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[227,420,1052,1078]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="anteridorsalis">
<emphasis id="B90F2A063911FFE2FF1DFBB930B9FBB3" box="[227,420,1052,1078]" italics="true" pageId="10" pageNumber="340">H. anteridorsalis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B8D973911FFE2FE4AFBB9330BFBB3" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[436,534,1053,1078]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="nigra">
<emphasis id="B90F2A063911FFE2FE4AFBB9330BFBB3" box="[436,534,1053,1078]" italics="true" pageId="10" pageNumber="340">H. nigra</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7B8D973911FFE2FDA9FBB933EEFBB3" authorityName="Hu &amp; Zhang" authorityYear="2010" box="[599,755,1052,1078]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A063911FFE2FDA9FBB933EEFBB3" box="[599,755,1052,1078]" italics="true" pageId="10" pageNumber="340">H. pycnolepis</emphasis>
</taxonomicName>
); bars on anterior flank crowded together, faintly visible in fresh specimens (vs. obvious in
<taxonomicName id="4C7B8D973911FFE2FE02FBE533B2FBDF" baseAuthorityName="Zhou &amp; He" baseAuthorityYear="1993" box="[508,687,1088,1114]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="acuticephala">
<emphasis id="B90F2A063911FFE2FE02FBE533B2FBDF" box="[508,687,1088,1114]" italics="true" pageId="10" pageNumber="340">H. acuticephala</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B8D973911FFE2FD42FBE5326DFBDF" baseAuthorityName="Zhu &amp; Wang" baseAuthorityYear="1985" box="[700,880,1088,1114]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="anguillioides">
<emphasis id="B90F2A063911FFE2FD42FBE5326DFBDF" box="[700,880,1088,1114]" italics="true" pageId="10" pageNumber="340">H. anguillioides</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7B8D973911FFE2FC80FBE53520FBDF" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[894,1085,1088,1114]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="anteridorsalis">
<emphasis id="B90F2A063911FFE2FC80FBE53520FBDF" box="[894,1085,1088,1114]" italics="true" pageId="10" pageNumber="340">H. anteridorsalis</emphasis>
</taxonomicName>
,
<emphasis id="B90F2A063911FFE2FBB4FBE53017FBFB" italics="true" pageId="10" pageNumber="340">
<taxonomicName id="4C7B8D973911FFE2FBB4FBE535DDFBDF" authorityName="Endruweit" authorityYear="2015" box="[1098,1216,1088,1114]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="change">H. change</taxonomicName>
,
<taxonomicName id="4C7B8D973911FFE2FB34FBE5346BFBDF" authorityName="Endruweit, Min &amp; Yang" authorityYear="2018" box="[1226,1398,1088,1114]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="coccinocola">H. coccinocola</taxonomicName>
,
<taxonomicName id="4C7B8D973911FFE2FA7FFBE53017FBFB" baseAuthorityName="Zhu &amp; Cao" baseAuthorityYear="1988" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="erhaiensis">H. erhaiensis</taxonomicName>
</emphasis>
,
<taxonomicName id="4C7B8D973911FFE2FEE6FBC130AFFBFB" authorityName="Hu &amp; Zhang" authorityYear="2010" box="[280,434,1124,1150]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="pycnolepis">
<emphasis id="B90F2A063911FFE2FEE6FBC130AFFBFB" box="[280,434,1124,1150]" italics="true" pageId="10" pageNumber="340">H. pycnolepis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C7B8D973911FFE2FE11FBC133BEFBFB" authorityName="Min, Yang &amp; Chen" authorityYear="2012" box="[495,675,1124,1150]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="340" phylum="Chordata" rank="species" species="wuliangensis">
<emphasis id="B90F2A063911FFE2FE11FBC133BEFBFB" box="[495,675,1124,1150]" italics="true" pageId="10" pageNumber="340">H. wuliangensis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3911FFE3FF39FB2C33A6FE0E" lastPageId="11" lastPageNumber="341" pageId="10" pageNumber="340" type="description">
<paragraph id="8BC4F6143911FFE2FF39FB2C3519FB6E" blockId="10.[151,1437,620,2015]" pageId="10" pageNumber="340">
<emphasis id="B90F2A063911FFE2FF39FB2C3044FB26" bold="true" box="[199,345,1160,1187]" pageId="10" pageNumber="340">Description.</emphasis>
Based on
<typeStatus id="54C048B63911FFE2FE33FB2C3332FB27" box="[461,559,1160,1186]" pageId="10" pageNumber="340" type="holotype">holotype</typeStatus>
and
<specimenCount id="9D7D3D9D3911FFE2FD98FB2D33EBFB26" box="[614,758,1161,1187]" pageId="10" pageNumber="340" type="generic" typeStatus="paratypes">10 paratypes</specimenCount>
, maximum standard length is
<quantity id="4C835BF13911FFE2FBB8FB2D35A6FB26" box="[1094,1211,1161,1187]" metricMagnitude="-1" metricUnit="m" metricValue="1.408" pageId="10" pageNumber="340" unit="mm" value="140.8">140.8 mm</quantity>
. Morphometric data are shown in
<tableCitation id="C6F9C3AF3911FFE2FECDFB083090FB42" box="[307,397,1196,1223]" captionStart="TABLE 2" captionStartId="14.[151,239,470,496]" captionTargetBox="[165,1975,507,1700]" captionText="TABLE 2. Morphometric data of three new Homatula species and H. pycnolepis" httpUri="http://table.plazi.org/id/DF04A69C3915FFE6FF69FE7232E8FE75" pageId="10" pageNumber="340" tableUuid="DF04A69C3915FFE6FF69FE7232E8FE75">Table 2</tableCitation>
. Dorsal-fin rays iii, 8½; anal-fin rays iii, 5½; pectoral-fin rays i, 9–10; pelvic-fin rays i, 7; caudal-fin branched rays 9+8; vertebrae 4+42 (
<specimenCount id="9D7D3D9D3911FFE2FD57FB753235FB6E" box="[681,808,1232,1259]" pageId="10" pageNumber="340" type="generic">1 specimen</specimenCount>
)–43 (
<specimenCount id="9D7D3D9D3911FFE2FC96FB7532E5FB6E" box="[872,1016,1232,1259]" pageId="10" pageNumber="340" type="generic">2 specimens</specimenCount>
).
</paragraph>
<paragraph id="8BC4F6143911FFE2FF39FB503284FAFF" blockId="10.[151,1437,620,2015]" pageId="10" pageNumber="340">Body elongate and cylindrical, anteriorly slightly depressed, posteriorly compressed laterally. Maximum body depth immediately in front of dorsal fin. Dorsal and ventral profiles almost straight. Except for scaleless head and thorax, body entirely covered by small scales, with scattered scales on abdomen. Caudal peduncle compressed laterally. Lateral line complete, extending directly along midline of body.</paragraph>
<paragraph id="8BC4F6143911FFE2FF39FA203223F98F" blockId="10.[151,1437,620,2015]" pageId="10" pageNumber="340">Head short and depressed in frontal view, wider than high, roughly triangular in dorsal view. Snout blunt, slightly shorter than postorbital length of head. Eye small, close to dorsal profile of head, invisible from ventral view. Interorbital space wide and flat. Nostrils closely set, nearer to anterior margin of eye than to snout tip; anterior nare pierced in front side of a flap-like tube with pointed tip.</paragraph>
<paragraph id="8BC4F6143911FFE2FF39F9B03516F93A" blockId="10.[151,1437,620,2015]" pageId="10" pageNumber="340">
Mouth inferior and arched. Lips thick, slightly furrowed, but not papillated; upper lip with small median incision, lower lip with marked median incision. Jaws covered by lips; upper jaw with well developed processus dentiformis corresponding with marked notch on lower jaw. Three pairs of barbels; two rostral pairs, inner barbel not reaching corner of mouth and outer barbel reaching vertical line of anterior nostril; one maxillary barbel pair long, reaching to vertical line between middle and posterior margin of eye (
<figureCitation id="1340EA913911FFE2FC5BF90032E3F93A" box="[933,1022,1700,1727]" captionStart="FIGURE 7" captionStartId="8.[151,250,1933,1958]" captionTargetBox="[217,1391,853,1887]" captionTargetId="figure@8.[151,1436,838,1899]" captionTargetPageId="8" captionText="FIGURE 7. Mouths of Homatula. (A) H. anteridorsalis, SWFC 1610006, holotype, 121.6 mm SL; (B) H. cryptoclathrata, SWFC 1002001, holotype, 138.1 mm SL; (C) H. nigra, SWFC 1901005, holotype, 99.8 mm SL; (D) H. pycnolepis, SWFC 0612085, 99.6 mm SL." figureDoi="http://doi.org/10.5281/zenodo.3577075" httpUri="https://zenodo.org/record/3577075/files/figure.png" pageId="10" pageNumber="340">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="8BC4F6143911FFE3FF39F96C3289FF52" blockId="10.[151,1437,620,2015]" lastBlockId="11.[151,1437,153,683]" lastPageId="11" lastPageNumber="341" pageId="10" pageNumber="340">Dorsal fin with 3 simple and 8–9 branched rays; distal margin convex; origin almost midway between snout tip and caudal-fin base. Pectoral fin with 1 simple and 9–10 branched rays, inserted vertically through posteriormost point of operculum, tip of longest fin ray not extending beyond halfway to insertion of pelvic fin. Pelvic fin with 1 simple and 6–7 branched rays, inserted below first or second branched rays of dorsal fin, tip of longest fin ray not extending beyond half distance between pelvic-fin insertion and anal-fin origin. Pelvic axillary lobe with pointed tip. Anal fin with 3 simple and 5 branched rays, with convex distal edge; origin closer to pelvic-fin insertion than to caudal-fin base. Posterior margin of caudal fin oblique or micro-concave. Caudal peduncle uniformly deep; with adipose crests along dorsal and ventral midlines. Adipose crest along dorsal midline of caudal peduncle not extend- ing through posterior end of anal-fin base. Anus located posteriorly, approximately 1/3 the distance between the pelvic-fin insertion and anal-fin origin, closer to the caudal-fin base.</paragraph>
<paragraph id="8BC4F6143910FFE3FF39FF4433A6FE0E" blockId="11.[151,1437,153,683]" pageId="11" pageNumber="341">
<emphasis id="B90F2A063910FFE3FF39FF44304FFF7E" bold="true" box="[199,338,224,251]" pageId="11" pageNumber="341">Coloration:</emphasis>
Flank of fresh specimens dark yellow, abdomen pale yellow.After fixation in 95% alcohol, flank is brown, abdomen grayish yellow. Flank with 25 to 27 brown vertical bars; width of each bar approximately equal to or narrower than spaces between bars. Bars in front of anal-fin origin narrow and dense, gradually becoming wider towards caudal-fin base. Dark brown vertical bar on caudal-fin base. Dorsal surface of pectoral, pelvic, and anal fins reddish orange, dorsal and caudal fins yellowish.
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3910FFE3FF39FE303271FDBA" pageId="11" pageNumber="341" type="distribution">
<paragraph id="8BC4F6143910FFE3FF39FE3033E3FE56" blockId="11.[151,1437,153,683]" pageId="11" pageNumber="341">
<emphasis id="B90F2A063910FFE3FF39FE30307EFE2A" bold="true" box="[199,355,404,431]" pageId="11" pageNumber="341">Distribution:</emphasis>
Known only from the Damengtong-he, a branch of the Mengboluo-he, the first tributary of the Nu-jiang drainagec (the upper Salween River) (
<figureCitation id="1340EA913910FFE3FD52FE1D33EDFE56" box="[684,752,441,467]" captionStart="FIGURE 8" captionStartId="9.[151,250,1995,2020]" captionTargetBox="[159,1425,702,1967]" captionTargetId="figure@9.[151,1436,695,1973]" captionTargetPageId="9" captionText="FIGURE 8. Distribution map of the densely-scaled group of Homatula. A symbol may represent several adjacent localities." figureDoi="http://doi.org/10.5281/zenodo.3577077" httpUri="https://zenodo.org/record/3577077/files/figure.png" pageId="11" pageNumber="341">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8BC4F6143910FFE3FF39FE783271FDBA" blockId="11.[151,1437,153,683]" pageId="11" pageNumber="341">
<emphasis id="B90F2A063910FFE3FF39FE7830D6FE72" bold="true" box="[199,459,476,503]" pageId="11" pageNumber="341">Habitat and Ecology:</emphasis>
<taxonomicName id="4C7B8D973910FFE3FE2AFE7933E6FE72" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[468,763,477,503]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="341" phylum="Chordata" rank="species" species="cryptoclathrata">
<emphasis id="B90F2A063910FFE3FE2AFE7933E6FE72" box="[468,763,477,503]" italics="true" pageId="11" pageNumber="341">Homatula cryptoclathrata</emphasis>
</taxonomicName>
is omnivorous, mainly feeding on algae attached to rocks, organic residues, and small aquatic insects. It inhabits flowing and clear stream environments and dwells on the stream bottom, shuttling back and forth among rocks and gravel.
</paragraph>
</subSubSection>
<subSubSection id="C361A59F3910FFE3FF39FDED31FFFD2E" pageId="11" pageNumber="341" type="etymology">
<paragraph id="8BC4F6143910FFE3FF39FDED31FFFD2E" blockId="11.[151,1437,153,683]" pageId="11" pageNumber="341">
<emphasis id="B90F2A063910FFE3FF39FDED304FFDE6" bold="true" box="[199,338,585,611]" pageId="11" pageNumber="341">Etymology:</emphasis>
The specific epithet,
<emphasis id="B90F2A063910FFE3FDBEFDED33E8FDE6" box="[576,757,585,611]" italics="true" pageId="11" pageNumber="341">
<taxonomicName id="4C7B8D973910FFE3FDBEFDED33ECFDE6" authorityName="Li &amp; Che &amp; Zhou" authorityYear="2019" box="[576,753,585,611]" class="Actinopterygii" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="341" phylum="Chordata" rank="species" species="cryptoclathrata">cryptoclathrata</taxonomicName>
,
</emphasis>
an adjective, is the combination of the Latin
<emphasis id="B90F2A063910FFE3FB17FDEE342CFDE6" box="[1257,1329,586,611]" italics="true" pageId="11" pageNumber="341">crypto</emphasis>
– (cryptic or hidden) and –
<emphasis id="B90F2A063910FFE3FEACFDC930D9FD02" box="[338,452,621,647]" italics="true" pageId="11" pageNumber="341">clathratus</emphasis>
(barred), alluding to the indistinct pattern of bars on the flank in front of the dorsal-fin origin.
</paragraph>
</subSubSection>
</treatment>
</document>
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