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<document id="D2D20E1EC6F5E6F60F4E30B6184B0A55" ID-DOI="10.37520/aemnp.2021.007" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037282" ID-ZooBank="FE96D8BE-6087-436C-8728-7BE849CA13B7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624895298315" checkinUser="carolina" docAuthor="Gapon, Dmitry A." docDate="2021" docId="03EC7368FFC3FF86FC34FB6DFD1EFE0E" docLanguage="en" docName="ActaEntMusNatPra.61.1.133-161.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.007" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Exosehirus validus" docType="treatment" docVersion="3" lastPageNumber="151" masterDocId="FFD50B10FFC8FF94FF97FFB3FFACFFFB" masterDocTitle="Revision of the genus Exosehirus (Heteroptera: Cydnidae), with the description of two new species" masterLastPageNumber="161" masterPageNumber="133" pageNumber="144" updateTime="1698977273770" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="7932F7FA2BDD741AF47EAB11F3CFDCDB">Revision of the genus Exosehirus (Heteroptera: Cydnidae), with the description of two new species</mods:title>
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<mods:date id="16775CEC8EE0314D113E649A77D14AC9">2021</mods:date>
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<subSubSection id="C35F91F5FFC3FF9FFC34FB6DFA9FFB0E" box="[931,1331,1246,1269]" pageId="11" pageNumber="144" type="nomenclature">
<paragraph id="8BFAC27EFFC3FF9FFC34FB6DFA9FFB0E" blockId="11.[931,1331,1246,1269]" box="[931,1331,1246,1269]" pageId="11" pageNumber="144">
<heading id="D0B27512FFC3FF9FFC34FB6DFA9FFB0E" box="[931,1331,1246,1269]" centered="true" fontSize="10" level="2" pageId="11" pageNumber="144" reason="3">
<taxonomicName id="4C45B9FDFFC3FF9FFC34FB6DFA9FFB0E" authority="(Jakovlev, 1877)" baseAuthorityName="Jakovlev" baseAuthorityYear="1877" box="[931,1331,1246,1269]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="144" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="B9311E6CFFC3FF9FFC34FB6DFA9FFB0E" bold="true" box="[931,1331,1246,1269]" pageId="11" pageNumber="144">
<emphasis id="B9311E6CFFC3FF9FFC34FB6DFBD9FB0E" bold="true" box="[931,1141,1246,1269]" italics="true" pageId="11" pageNumber="144">Exosehirus validus</emphasis>
(Jakovlev, 1877)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C35F91F5FFC3FF9FFCEFFB4FFAF3FAF4" box="[888,1375,1276,1295]" pageId="11" pageNumber="144" type="description">
<paragraph id="8BFAC27EFFC3FF9FFCEFFB4FFAF3FAF4" blockId="11.[827,1444,1276,2065]" box="[888,1375,1276,1295]" pageId="11" pageNumber="144">
(
<figureCitation id="137EDEFBFFC3FF9FFCE9FB4FFC69FAF4" box="[894,965,1276,1295]" captionStart="Fig" captionStartId="4.[151,182,1834,1853]" captionTargetBox="[164,1391,218,1805]" captionTargetId="figure-13@4.[151,1436,189,1820]" captionTargetPageId="4" captionText="Fig. 2. Exosehirus species, general appearance from dorsal side and type labels. A – E. validus (Jakovlev, 1877), holotype; B – E. elamensis sp. nov., holotype; C – E. essedonius sp. nov., holotype; D – E. marginatus (Signoret, 1881), without abdomen, from Turkey." figureDoi="http://doi.org/10.5281/zenodo.5037288" httpUri="https://zenodo.org/record/5037288/files/figure.png" pageId="11" pageNumber="144">Figs 2A</figureCitation>
, 
<figureCitation id="137EDEFBFFC3FF9FFC5AFB4FFBB7FAF4" box="[973,1051,1276,1295]" captionStart="Fig" captionStartId="5.[150,181,2008,2027]" captionTargetBox="[150,1435,204,1928]" captionTargetId="figure-11@5.[150,1435,189,1928]" captionTargetPageId="5" captionText="Fig. 3. Head (A–F), pronotum (G–K) and external scent efferent system (L) of Exosehirus species.A, G, L – E. validus (Jakovlev, 1877); B, H – E. sargon Linnavuori, 1984; C, I – E. elamensis sp. nov.; D, E, J – E. essedonius sp. nov.; F, K – E. marginatus (Signoret, 1881). Scale bars: 0.25 mm (A, C–F, L) and 0.5 mm (G, I–K). B and H after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037290" httpUri="https://zenodo.org/record/5037290/files/figure.png" pageId="11" pageNumber="144">3A, G, L</figureCitation>
, 
<figureCitation id="137EDEFBFFC3FF9FFBB3FB4FFB25FAF4" box="[1060,1161,1276,1295]" captionStart="Fig" captionStartId="7.[151,182,1951,1970]" captionTargetBox="[153,1433,194,1836]" captionTargetId="figure-11@7.[150,1435,189,1852]" captionTargetPageId="7" captionText="Fig. 4. Pygophore, paramere and aedeagus of Exosehirus species.A, F, G, H – E. validus (Jakovlev, 1877); B, K – E. sargon Linnavuori, 1984; C – E. elamensis sp. nov.; D, I – E. essedonius sp. nov.; E, J – E. marginatus (Signoret, 1881). Pygophore in ventrocaudal (A–E) and dorsal (F) view; paramere in lateral (G) and dorsal (H–J) view; aedeagus in lateral view (K). Abbreviations: d.inf – dorsal infolding of pygophore; l.inf – lateral infolding of pygophore; l.r – lateral ridge of subgenital plate; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding of pygophore. Scale bars: 0.25 mm (A, C, D, E, F) and 0.14 mm (G–J). B and K after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037292" httpUri="https://zenodo.org/record/5037292/files/figure.png" pageId="11" pageNumber="144">4A, F, G, H</figureCitation>
, 
<figureCitation id="137EDEFBFFC3FF9FFB05FB4FFB6AFAF4" box="[1170,1222,1276,1295]" captionStart="Fig" captionStartId="8.[151,182,2030,2049]" captionTargetBox="[203,1427,417,1998]" captionTargetId="figure-13@8.[151,1436,411,2018]" captionTargetPageId="8" captionText="Fig. 5. Uninflated aedeagi of Exosehirus species.A, B – E. validus (Jakovlev, 1877); C, D – E. essedonius sp. nov. Lateral (A, C) and ventral (B, D) view. Abbreviations: vl.l – ventrolateral lobe of conjunctiva; dl.l – dorsolateral lobe; da.l – dorsoapical lobe. Scale bar: 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.5037294" httpUri="https://zenodo.org/record/5037294/files/figure.png" pageId="11" pageNumber="144">5A, B</figureCitation>
, 
<figureCitation id="137EDEFBFFC3FF9FFB58FB4FFAA8FAF4" box="[1231,1284,1276,1295]" captionStart="Fig" captionStartId="9.[151,183,1535,1554]" captionTargetBox="[154,1434,208,1514]" captionTargetId="figure-270@9.[151,1436,189,1518]" captionTargetPageId="9" captionText="Fig. 6. Wet preparations of conjunctival lobes of Exosehirus species. A–C – E. validus (Jakovlev, 1877); D–F – E. sargon Linnavuori, 1984; G–I – E. elamensis sp. nov.; J–L – E. essedonius sp. nov. Ventrolateral lobe (A, D, G, J); dorsolateral lobe (B, E, H, K); dorsoapical lobe (C, F, I, L). Scale bar: 0.14 mm. D–F after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037296" httpUri="https://zenodo.org/record/5037296/files/figure.png" pageId="11" pageNumber="144">6A–C</figureCitation>
, 
<figureCitation id="137EDEFBFFC3FF9FFA9AFB4FFAB5FAF4" box="[1293,1305,1276,1295]" captionStart="Fig" captionStartId="12.[943,974,1953,1972]" captionTargetBox="[156,1438,197,2079]" captionTargetId="figure-13@12.[151,1436,189,2079]" captionTargetPageId="12" captionText="Fig. 8. Completely inflated aedeagus of Exosehirus vali- dus (Jakovlev, 1877). A – ventral view; B – lateral view; C – dorsal view.Abbreviations: vl.l – ventrolateral lobe of conjunctiva;dl.l – dorsolateral lobe; da.l – dorsoapical lobe. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5037300" httpUri="https://zenodo.org/record/5037300/files/figure.png" pageId="11" pageNumber="144">8</figureCitation>
, 
<figureCitation id="137EDEFBFFC3FF9FFAB5FB4FFA95FAF4" box="[1314,1337,1276,1295]" captionStart="Fig" captionStartId="19.[151,182,1978,1997]" captionTargetBox="[162,1434,196,1904]" captionTargetId="figure-10@19.[150,1435,189,1915]" captionTargetPageId="19" captionText="Fig. 12. Female terminalia of Exosehirus validus (Jakovlev, 1877).A – dorsal view; B – ventral view.Abbreviations: a.s – arcuate sclerites; a.l.g – anterior limb of gonangulum; f.r – first rami; g.c – gynatrial cone; gc.I – gonocoxites I; gc.II – gonocoxites II; gp.I – gonapophyses I; gp.II – gonapophyses II; p.i.m – posterior intervalvular membrane; p.l.g – posterior limb of gonangulum; p.p – pouches of posterior part of gynatrial sac; pt.IX – patatergites IX; r.s – ring sclerites; s.r – second rami; v.f – ventral fold of posterior intervalvular membrane. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5037312" httpUri="https://zenodo.org/record/5037312/files/figure.png" pageId="11" pageNumber="144">12</figureCitation>
, 
<figureCitation id="137EDEFBFFC3FF9FFAD5FB4FFAF4FAF4" box="[1346,1368,1276,1295]" captionStart="Fig" captionStartId="20.[151,182,2032,2051]" captionTargetBox="[206,1365,190,1934]" captionTargetId="figure-13@20.[186,1402,165,1953]" captionTargetPageId="20" captionText="Fig. 13. Posterior part of gynatrial sac of Exosehirus validus (Jakovlev, 1877). A – dorsoposterior view; B – dorsolateral view. Abbreviation: g.c – gynatrial cone. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5037314" httpUri="https://zenodo.org/record/5037314/files/figure.png" pageId="11" pageNumber="144">13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F91F5FFC3FF84FCACFAADFE6EFBEA" lastPageId="16" lastPageNumber="149" pageId="11" pageNumber="144" type="materials_examined">
<paragraph id="8BFAC27EFFC3FF9FFCACFAADFA30F7EA" blockId="11.[827,1444,1276,2065]" pageId="11" pageNumber="144">
<taxonomicName id="4C45B9FDFFC3FF9FFCACFAADFB0EFACA" authority="Jakovlev, 1877: 76" authorityName="Jakovlev" authorityPageNumber="76" authorityYear="1877" box="[827,1186,1310,1329]" class="Insecta" family="Cydnidae" genus="Gnathoconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="144" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="B9311E6CFFC3FF9FFCACFAADFC5FFACA" box="[827,1011,1310,1329]" italics="true" pageId="11" pageNumber="144">Gnathoconus validus</emphasis>
Jakovlev, 1877: 76
</taxonomicName>
; Pඎඍඈඇ (1886: 9); Oඌ*üൺඇංඇ (1906: 19, 1910: 5). 
<taxonomicName id="4C45B9FDFFC3FF9FFCACFAE7FC60FA9C" baseAuthorityName="Jakovlev" baseAuthorityYear="1877" box="[827,972,1364,1383]" class="Insecta" family="Cydnidae" genus="Legnotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="144" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="B9311E6CFFC3FF9FFCACFAE7FC60FA9C" box="[827,972,1364,1383]" italics="true" pageId="11" pageNumber="144">Legnotus validus</emphasis>
</taxonomicName>
: Hඈൻൾ*©අൺඇൽඍ (1961: 199); Pඎඍඌ*üκඈඏ (1965: 55). 
<taxonomicName id="4C45B9FDFFC3FF9FFCACFADDFC71FA7A" baseAuthorityName="Jakovlev" baseAuthorityYear="1877" box="[827,989,1390,1409]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="144" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="B9311E6CFFC3FF9FFCACFADDFC71FA7A" box="[827,989,1390,1409]" italics="true" pageId="11" pageNumber="144">Exosehirus validus</emphasis>
</taxonomicName>
: WൺǤඇൾ*© (1963: 107); Lංඌ (1999: 223; 2006: 142). 
<emphasis id="B9311E6CFFC3FF9FFCACFA2BFC6EFA50" bold="true" box="[827,962,1432,1451]" pageId="11" pageNumber="144">Type material.</emphasis>
Hඈඅඈඍඒඉൾ: 
<specimenCount id="9D4309F7FFC3FF9FFBA7FA21FBECFA50" box="[1072,1088,1426,1451]" pageId="11" pageNumber="144" type="female">♀</specimenCount>
[mounted on short rectangle], “[golden square] // 354 [pink circle with numbers] // [golden circle] // “Shaku [
<emphasis id="B9311E6CFFC3FF9FFCD5FA7DFCD4FA1A" bold="true" box="[834,888,1486,1505]" pageId="11" pageNumber="144">
<collectingCountry id="F35282EEFFC3FF9FFCD5FA7DFCD4FA1A" box="[834,888,1486,1505]" name="Iran" pageId="11" pageNumber="144">IRAN</collectingCountry>
</emphasis>
, 
<emphasis id="B9311E6CFFC3FF9FFC15FA7DFBBBFA1A" bold="true" box="[898,1047,1486,1505]" pageId="11" pageNumber="144">Gඈඅൾඌඍൺඇ Pඋඈඏ.</emphasis>
, Shahkuh Range near Shahrud, 
<geoCoordinate id="EE71A4B9FFC3FF9FFAB4FA7DFA30FA1A" box="[1315,1436,1486,1505]" degrees="36" direction="north" minutes="30" orientation="latitude" pageId="11" pageNumber="144" precision="1" seconds="00.9" value="36.50025">36°30’00.9”N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFC3FF9FFCACFA5BFC1FFA00" box="[827,947,1512,1531]" degrees="54" direction="east" minutes="24" orientation="longitude" pageId="11" pageNumber="144" precision="1" seconds="21.3" value="54.40592">54°24’21.3”E</geoCoordinate>
]” // “soli… [illegible handwriting; probably “solitarius”]” // “k. 
<collectionCode id="ED545ABBFFC3FF9FFCFEF9B0FCD4F9ED" box="[873,888,1539,1558]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="144" type="Museum">V</collectionCode>
. Yakovleva [coll. of B. Jakovlev]” // “ 
<taxonomicName id="4C45B9FDFFC3FF9FFB20F9B0FA84F9ED" authorityName="Fieber" authorityYear="1861" box="[1207,1320,1539,1558]" class="Insecta" family="Cydnidae" genus="Gnathoconus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="159" phylum="Arthropoda" rank="genus">Gnathoconus</taxonomicName>
/ 
<taxonomicName id="4C45B9FDFFC3FF9FFAA0F9B0FA31F9ED" authority="Jak." authorityName="Jak." box="[1335,1437,1539,1558]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="144" phylum="Arthropoda" rank="species" species="validus">validus Jak.</taxonomicName>
/ B. Jakowlew det. // 
<typeStatus id="54FE7CDCFFC3FF9FFC67F9ADFBE5F9CA" box="[1008,1097,1566,1585]" pageId="11" pageNumber="144" type="holotype">Holotypus</typeStatus>
[red rectangle]” (
<collectionCode id="ED545ABBFFC3FF9FFB77F9ADFABCF9CA" box="[1248,1296,1566,1585]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="144">ZISP</collectionCode>
). 
<emphasis id="B9311E6CFFC3FF9FFCACF98BFBFDF9B0" bold="true" box="[827,1105,1592,1611]" pageId="11" pageNumber="144">Additional material examined</emphasis>
(all in 
<collectionCode id="ED545ABBFFC3FF9FFB01F98BFB6AF9B0" box="[1174,1222,1592,1611]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="11" pageNumber="144">ZISP</collectionCode>
). Without exact locality, “A. Gekk”, date unknown, coll. B. Jakovlev, 1* J. 
<emphasis id="B9311E6CFFC3FF9FFB69F9E0FC43F97A" bold="true" pageId="11" pageNumber="144">
<collectingCountry id="F35282EEFFC3FF9FFB69F9E0FA34F99D" box="[1278,1432,1619,1638]" name="Kazakhstan" pageId="11" pageNumber="144">KAZAKHSTAN</collectingCountry>
: 
<collectionCode id="ED545ABBFFC3FF9FFCACF9DDFCEFF97A" box="[827,835,1646,1665]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="144">T</collectionCode>
ඎඋΚൾඌඍൺඇ Pඋඈඏ.:
</emphasis>
“st.[ation] Arys’” [Arys Town, 
<geoCoordinate id="EE71A4B9FFC3FF9FFA88F9DDFA37F97A" box="[1311,1435,1646,1665]" degrees="42" direction="north" minutes="25" orientation="latitude" pageId="11" pageNumber="144" precision="1" seconds="60.0" value="42.433334">42°25′60.0″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFC3FF9FFCACF93BFC1AF960" box="[827,950,1672,1691]" degrees="68" direction="east" minutes="48" orientation="longitude" pageId="11" pageNumber="144" precision="1" seconds="00.0" value="68.8">68°48′00.0″E</geoCoordinate>
], 
<date id="FFFBE4BEFFC3FF9FFC50F93BFB89F960" box="[967,1061,1672,1691]" pageId="11" pageNumber="144" value="1938-04-20">20.iv.1938</date>
, F. Lukjanovitsh, 1* J. 
<emphasis id="B9311E6CFFC3FF9FFA90F93BFC1FF94D" bold="true" pageId="11" pageNumber="144">
<collectingCountry id="F35282EEFFC3FF9FFA90F93BFA3BF960" box="[1287,1431,1672,1691]" name="Uzbekistan" pageId="11" pageNumber="144">UZBEKISTAN</collectingCountry>
: Nൺඏඈං Pඋඈඏ.:
</emphasis>
Khatyrchi Distr.: Yargak near “Khatyrchi [Yangirabad]”, 19,20. 
<date id="FFFBE4BEFFC3FF9FFCE4F90DFC1BF92A" box="[883,951,1726,1745]" pageId="11" pageNumber="144" value="1929-06">vi.1929</date>
, L. Zimin leg., 2 JJ, 1 J; Changir near “Khatyrchi [Yangirabad]”, 
<geoCoordinate id="EE71A4B9FFC3FF9FFC52F96BFB92F910" box="[965,1086,1752,1771]" degrees="40" direction="north" minutes="06" orientation="latitude" pageId="11" pageNumber="144" precision="1" seconds="00.0" value="40.1">40°06′00.0″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFC3FF9FFBD1F96BFB13F910" box="[1094,1215,1752,1771]" degrees="65" direction="east" minutes="56" orientation="longitude" pageId="11" pageNumber="144" precision="1" seconds="00.0" value="65.933334">65°56′00.0″E</geoCoordinate>
, 
<date id="FFFBE4BEFFC3FF9FFB5CF96BFA86F910" box="[1227,1322,1752,1771]" pageId="11" pageNumber="144" value="1930-06-15">15.vi.1930</date>
, 4*+39 JJ, 3*+ 
<specimenCount id="9D4309F7FFC3FF9FFCCBF940FC31F8FE" box="[860,925,1772,1798]" pageId="11" pageNumber="144" type="female">37 ♀♀</specimenCount>
, 
<date id="FFFBE4BEFFC3FF9FFC3BF940FBA2F8FD" box="[940,1038,1779,1798]" pageId="11" pageNumber="144" value="1930-04-28">28.iv.1930</date>
, 1 J, L. Zimin leg. 
<emphasis id="B9311E6CFFC3FF9FFB42F940FCD4F8DA" bold="true" pageId="11" pageNumber="144">KൺඌΗΚൺൽൺඋ’ංඇඌΚൺඒൺ Pඋඈඏ.:</emphasis>
Guzar Distr.: Guzar, 
<geoCoordinate id="EE71A4B9FFC3FF9FFBABF8BDFB1FF8DA" box="[1084,1203,1806,1825]" degrees="38" direction="north" minutes="35" orientation="latitude" pageId="11" pageNumber="144" precision="1" seconds="13.4" value="38.587055">38°35′13.4″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFC3FF9FFB2CF8BDFA9EF8DA" box="[1211,1330,1806,1825]" degrees="66" direction="east" minutes="17" orientation="longitude" pageId="11" pageNumber="144" precision="1" seconds="01.8" value="66.28384">66°17′01.8″E</geoCoordinate>
, 
<date id="FFFBE4BEFFC3FF9FFAA9F8BDFA3BF8DA" box="[1342,1431,1806,1825]" pageId="11" pageNumber="144" value="1929-07-07">7.vii.1929</date>
, Elizarova, 
<specimenCount id="9D4309F7FFC3FF9FFC0BF89BFC12F8C0" box="[924,958,1826,1851]" pageId="11" pageNumber="144" type="female">1 ♀</specimenCount>
; Kammashi, 
<geoCoordinate id="EE71A4B9FFC3FF9FFBAFF89BFB03F8C0" box="[1080,1199,1832,1851]" degrees="38" direction="north" minutes="48" orientation="latitude" pageId="11" pageNumber="144" precision="1" seconds="34.6" value="38.80961">38°48′34.6″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFC3FF9FFB20F89BFA82F8C0" box="[1207,1326,1832,1851]" degrees="66" direction="east" minutes="28" orientation="longitude" pageId="11" pageNumber="144" precision="1" seconds="52.9" value="66.48136">66°28′52.9″E</geoCoordinate>
, 
<date id="FFFBE4BEFFC3FF9FFAAEF89BFA34F8C0" box="[1337,1432,1832,1851]" pageId="11" pageNumber="144" value="1930-06-27">27.vi.1930</date>
, 
<date id="FFFBE4BEFFC3FF9FFCACF8F0FC38F8AD" box="[827,916,1859,1878]" pageId="11" pageNumber="144" value="1931-07-05">5.vii.1931</date>
, 
<collectionCode id="ED545ABBFFC3FF9FFC37F8F0FC1CF8AD" box="[928,944,1859,1878]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="144" type="Museum">V</collectionCode>
. Gussakovskij leg., 1*+1 JJ, 
<specimenCount id="9D4309F7FFC3FF9FFB52F8F0FB4BF8AE" box="[1221,1255,1852,1878]" pageId="11" pageNumber="144" type="female">1 ♀</specimenCount>
; Kamchashi, NE of Guzar, c. 
<geoCoordinate id="EE71A4B9FFC3FF9FFC01F8EDFBA3F88A" box="[918,1039,1886,1905]" degrees="38" direction="north" minutes="47" orientation="latitude" pageId="11" pageNumber="144" precision="1" seconds="29.5" value="38.791527">38°47′29.5″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFC3FF9FFB8FF8EDFB3DF88A" box="[1048,1169,1886,1905]" degrees="66" direction="east" minutes="29" orientation="longitude" pageId="11" pageNumber="144" precision="1" seconds="15.1" value="66.487526">66°29′15.1″E</geoCoordinate>
, 
<date id="FFFBE4BEFFC3FF9FFB0AF8EDFB52F88A" box="[1181,1278,1886,1905]" pageId="11" pageNumber="144" value="1932-06-30">30.vi.1932</date>
, 
<collectionCode id="ED545ABBFFC3FF9FFA9DF8EDFAB6F88A" box="[1290,1306,1886,1905]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="11" pageNumber="144" type="Museum">V</collectionCode>
. Gussakovskij leg., 2*+1 JJ. 
<emphasis id="B9311E6CFFC3FF9FFC58F8CBFB25F870" bold="true" box="[975,1161,1912,1931]" pageId="11" pageNumber="144">SൺආൺඋΚൺඇൽ Pඋඈඏ.:</emphasis>
environs of Samarkand, date unknown, L. Barshchevskiy leg., 
<specimenCount id="9D4309F7FFC3FF9FFBF3F820FB2BF85E" box="[1124,1159,1932,1958]" pageId="11" pageNumber="144" type="female">1 ♀</specimenCount>
; Samarkand, 
<date id="FFFBE4BEFFC3FF9FFA95F820FAFFF85D" box="[1282,1363,1939,1958]" pageId="11" pageNumber="144" value="1921-04-05">5.iv.1921</date>
, Z. Ivanova leg., 1* J. 
<emphasis id="B9311E6CFFC3FF9FFC5AF81DFB7AF83A" bold="true" box="[973,1238,1966,1985]" pageId="11" pageNumber="144">SඎඋΚΗൺඇൽൺඋ’ංඇඌΚൺඒൺ Pඋඈඏ.:</emphasis>
Termez, 
<geoCoordinate id="EE71A4B9FFC3FF9FFAB1F81DFA37F83A" box="[1318,1435,1966,1985]" degrees="37" direction="north" minutes="15" orientation="latitude" pageId="11" pageNumber="144" precision="1" seconds="43.1" value="37.26197">37°15′43.1″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFC3FF9FFCACF87BFC1CF820" box="[827,944,1992,2011]" degrees="67" direction="east" minutes="19" orientation="longitude" pageId="11" pageNumber="144" precision="1" seconds="08.7" value="67.319084">67°19′08.7″E</geoCoordinate>
, 
<date id="FFFBE4BEFFC3FF9FFC2EF87BFBA3F820" box="[953,1039,1992,2011]" pageId="11" pageNumber="144" value="1912-05-18">18.v.1912</date>
, A-r Kiritshenko, 
<specimenCount id="9D4309F7FFC3FF9FFB3BF87BFB60F820" box="[1196,1228,1986,2011]" pageId="11" pageNumber="144" type="female">1 ♀</specimenCount>
, 1912, M. Siyazov leg., 1* 
<specimenCount id="9D4309F7FFC3FF9FFCC3F86FFCC8F80E" box="[852,868,2012,2037]" pageId="11" pageNumber="144" type="female">♀</specimenCount>
. 
<emphasis id="B9311E6CFFC3FF9FFCFAF850FBABF80D" bold="true" box="[877,1031,2019,2038]" pageId="11" pageNumber="144">
<collectionCode id="ED545ABBFFC3FF9FFCFAF850FCDAF80D" box="[877,886,2019,2038]" name="Tavera, Department of Geology and Geophysics" pageId="11" pageNumber="144">T</collectionCode>
ൺඌΗΚൾඇඍ Pඋඈඏ.:
</emphasis>
Tashkent, 
<date id="FFFBE4BEFFC3FF9FFBC8F850FB0BF80D" box="[1119,1191,2019,2038]" pageId="11" pageNumber="144" value="1912-07">vii.1912</date>
, Seslavin leg., 
<specimenCount id="9D4309F7FFC3FF9FFAB5F850FAECF80E" box="[1314,1344,2012,2038]" pageId="11" pageNumber="144" type="female">1 ♀</specimenCount>
; Yangiyul’ Distr., Yangiyul’ near Tashkent, 
<geoCoordinate id="EE71A4B9FFC3FF9FFBD9F84DFB6FF7EA" box="[1102,1219,2046,2065]" degrees="41" direction="north" minutes="05" orientation="latitude" pageId="11" pageNumber="144" precision="1" seconds="44.5" value="41.09569">41°05′44.5″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFC3FF9FFB5EF84DFA92F7EA" box="[1225,1342,2046,2065]" degrees="69" direction="east" minutes="04" orientation="longitude" pageId="11" pageNumber="144" precision="1" seconds="14.5" value="69.070694">69°04′14.5″E</geoCoordinate>
, 
<date id="FFFBE4BEFFC3FF9FFAD0F84DFA34F7EA" box="[1351,1432,2046,2065]" pageId="11" pageNumber="144" value="1940-04-02">2.iv.1940</date>
,
</paragraph>
<caption id="DF3A92F6FFC4FF98FC38F812FBF0F7E4" ID-DOI="http://doi.org/10.5281/zenodo.5037300" ID-Zenodo-Dep="5037300" httpUri="https://zenodo.org/record/5037300/files/figure.png" pageId="12" pageNumber="145" startId="12.[943,974,1953,1972]" targetBox="[156,1438,197,2079]" targetPageId="12">
<paragraph id="8BFAC27EFFC4FF98FC38F812FBF0F7E4" blockId="12.[943,1438,1953,2079]" pageId="12" pageNumber="145">
Fig. 8. Completely inflated aedeagus of 
<emphasis id="B9311E6CFFC4FF98FA98F812FC61F834" italics="true" pageId="12" pageNumber="145">
<taxonomicName id="4C45B9FDFFC4FF98FA98F812FAC2F84F" authorityName="Wagner" authorityYear="1963" box="[1295,1390,1953,1972]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="145" phylum="Arthropoda" rank="genus">Exosehirus</taxonomicName>
vali- dus
</emphasis>
(Jakovlev, 1877). A – ventral view; B – lateral view; C – dorsal view.Abbreviations: 
<emphasis id="B9311E6CFFC4FF98FB57F865FB77F812" box="[1216,1243,2006,2025]" italics="true" pageId="12" pageNumber="145">vl.l</emphasis>
– ventrolateral lobe of conjunctiva; 
<emphasis id="B9311E6CFFC4FF98FB8DF842FB9AF7FF" box="[1050,1078,2033,2052]" italics="true" pageId="12" pageNumber="145">dl.l</emphasis>
– dorsolateral lobe; 
<emphasis id="B9311E6CFFC4FF98FB4BF842FB50F7FF" box="[1244,1276,2033,2052]" italics="true" pageId="12" pageNumber="145">da.l</emphasis>
– dorsoapical lobe. Scale bar: 0.25 mm.
</paragraph>
</caption>
<caption id="DF3A92F6FFC5FF99FC57F87FFB50F7EE" ID-DOI="http://doi.org/10.5281/zenodo.5037304" ID-Zenodo-Dep="5037304" httpUri="https://zenodo.org/record/5037304/files/figure.png" pageId="13" pageNumber="146" startId="13.[960,992,1996,2015]" targetBox="[169,1437,202,2069]" targetPageId="13">
<paragraph id="8BFAC27EFFC5FF99FC57F87FFB50F7EE" blockId="13.[960,1437,1996,2069]" pageId="13" pageNumber="146">
Fig. 9. Completely inflated aedeagus of 
<taxonomicName id="4C45B9FDFFC5FF99FAAEF87FFBB8F801" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="146" phylum="Arthropoda" rank="species" species="elamensis" status="sp. nov.">
<emphasis id="B9311E6CFFC5FF99FAAEF87FFBB8F801" italics="true" pageId="13" pageNumber="146">Exosehirus elamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFC5FF99FB8BF854FBCDF801" box="[1052,1121,2023,2042]" pageId="13" pageNumber="146" rank="species">sp. nov.</taxonomicNameLabel>
A – ventral view; B – lateral view; C – dorsal view. Scale bar: 0.25 mm.
</paragraph>
</caption>
<caption id="DF3A92F6FFC6FF9AFC75F87DFAB2F7EC" ID-DOI="http://doi.org/10.5281/zenodo.5037308" ID-Zenodo-Dep="5037308" httpUri="https://zenodo.org/record/5037308/files/figure.png" pageId="14" pageNumber="147" startId="14.[994,1025,1998,2017]" targetBox="[163,1437,208,2071]" targetPageId="14">
<paragraph id="8BFAC27EFFC6FF9AFC75F87DFAB2F7EC" blockId="14.[994,1437,1998,2071]" pageId="14" pageNumber="147">
Fig. 10. Completely inflated aedeagus of 
<taxonomicName id="4C45B9FDFFC6FF9AFAAAF87DFB91F807" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="14" pageNumber="147" phylum="Arthropoda" rank="species" species="essedonius" status="sp. nov.">
<emphasis id="B9311E6CFFC6FF9AFAAAF87DFB91F807" italics="true" pageId="14" pageNumber="147">Exosehirus essedonius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFC6FF9AFBD6F85AFB2EF807" box="[1089,1154,2025,2044]" pageId="14" pageNumber="147" rank="species">sp. nov.</taxonomicNameLabel>
A – ventral view; B – lateral view; C – dorsal view. Scale bar: 0.25 mm.
</paragraph>
</caption>
<caption id="DF3A92F6FFC7FF9BFC3AF82BFA3DF7EE" ID-DOI="http://doi.org/10.5281/zenodo.5037310" ID-Zenodo-Dep="5037310" httpUri="https://zenodo.org/record/5037310/files/figure.png" pageId="15" pageNumber="148" startId="15.[941,972,1944,1963]" targetBox="[155,1437,204,2069]" targetPageId="15">
<paragraph id="8BFAC27EFFC7FF9BFC3AF82BFA3DF7EE" blockId="15.[941,1437,1944,2069]" pageId="15" pageNumber="148">
Fig.11. Completely inflated aedeagus of 
<emphasis id="B9311E6CFFC7FF9BFA96F82BFC77F83E" italics="true" pageId="15" pageNumber="148">
<taxonomicName id="4C45B9FDFFC7FF9BFA96F82BFAF3F850" authorityName="Wagner" authorityYear="1963" box="[1281,1375,1944,1963]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="15" pageNumber="148" phylum="Arthropoda" rank="genus">Exosehirus</taxonomicName>
margi- natus
</emphasis>
(
<bibRefCitation id="EFD4BF8FFFC7FF9BFC71F801FBC5F83E" author="SIGNORET V." box="[998,1129,1970,1989]" pageId="15" pageNumber="148" pagination="621 - 657" refId="ref19757" refString="SIGNORET V. 1881: Revue des Cydnides contenus dans la collection du Musee Civique d'Histoire Naturelle de Genes. Annali del Museo Civico di Storia Naturale di Genova 16: 621 - 657." type="journal article" year="1881">Signoret, 1881</bibRefCitation>
). A – ventral view; B – lateral view; C – dorsal view. Abbreviations: 
<emphasis id="B9311E6CFFC7FF9BFB58F87EFB4BF81B" box="[1231,1255,1997,2016]" italics="true" pageId="15" pageNumber="148">a.s</emphasis>
– spicules of apical lobes; 
<emphasis id="B9311E6CFFC7FF9BFC72F85BFBACF800" box="[997,1024,2024,2043]" italics="true" pageId="15" pageNumber="148">vl.l</emphasis>
– ventrolateral lobe of conjunctiva; 
<emphasis id="B9311E6CFFC7FF9BFAA2F85BFAFDF800" box="[1333,1361,2024,2043]" italics="true" pageId="15" pageNumber="148">dl.l</emphasis>
– dorso- lateral lobe; 
<emphasis id="B9311E6CFFC7FF9BFB8FF7B1FB94F7EE" box="[1048,1080,2050,2069]" italics="true" pageId="15" pageNumber="148">da.l</emphasis>
– dorsoapical lobe. Scale bar: 0.25 mm.
</paragraph>
</caption>
<paragraph id="8BFAC27EFFD8FF84FF00FF77FE68FCDA" blockId="16.[151,761,189,802]" pageId="16" pageNumber="149">
Chirkun leg., 
<specimenCount id="9D4309F7FFD8FF84FE85FF77FEE8FF2D" box="[274,324,189,215]" pageId="16" pageNumber="149" type="female">2 ♀♀</specimenCount>
. 
<emphasis id="B9311E6CFFD8FF84FEC5FF77FE59FF2C" bold="true" box="[338,501,196,215]" pageId="16" pageNumber="149">AඇൽංඓΗൺඇ Pඋඈඏ.:</emphasis>
Pakhtaabad Distr., “Fergana, Ispaskan [Izboskan]”, 
<geoCoordinate id="EE71A4B9FFD8FF84FEF3FF6DFE73FF0A" box="[356,479,222,241]" degrees="41" direction="north" minutes="01" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="56.0" value="41.032223">41°01′56.0″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FE70FF6DFDCDFF0A" box="[487,609,222,241]" degrees="72" direction="east" minutes="20" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="27.9" value="72.34109">72°20′27.9″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FDFAFF6DFD62FF0A" box="[621,718,222,241]" pageId="16" pageNumber="149" value="1913-06-12">12.vi.1913</date>
, A-r Kiritshenko, 1* 
<specimenCount id="9D4309F7FFD8FF84FEB5FF41FE9EFEF0" box="[290,306,242,267]" pageId="16" pageNumber="149" type="female">♀</specimenCount>
. 
<emphasis id="B9311E6CFFD8FF84FEAAFF4AFD2EFEF7" bold="true" box="[317,642,249,268]" pageId="16" pageNumber="149">
<collectingCountry id="F35282EEFFD8FF84FEAAFF4AFE43FEF7" box="[317,495,249,268]" name="Turkmenistan" pageId="16" pageNumber="149">TURKMENISTAN</collectingCountry>
: BൺඅΚൺඇ Pඋඈඏ.:
</emphasis>
“Kizyl-Arvat [Serdar]” Town, 
<geoCoordinate id="EE71A4B9FFD8FF84FEBAFEA7FE0FFEDC" box="[301,419,276,295]" degrees="38" direction="north" minutes="58" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="00.0" value="38.966667">38°58′00.0″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FE3CFEA7FD8DFEDC" box="[427,545,276,295]" degrees="56" direction="east" minutes="16" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="00.0" value="56.266666">56°16′00.0″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FDBBFEA7FD25FEDC" box="[556,649,276,295]" pageId="16" pageNumber="149" value="1953-06-11">11.vi.1953</date>
, O. Kryzhanovskij leg., 1 J, 1*+ 
<specimenCount id="9D4309F7FFD8FF84FEF7FE9DFE3CFEBA" box="[352,400,296,321]" pageId="16" pageNumber="149" type="female">1 ♀♀</specimenCount>
, 
<date id="FFFBE4BEFFD8FF84FE09FE9DFE56FEBA" box="[414,506,302,321]" pageId="16" pageNumber="149" value="1953-04-13">13.iv.1953</date>
, E. Arens, 1 J; Kopet Dag, 
<quantity id="4CBD6F9BFFD8FF84FF00FEFAFF63FEA7" box="[151,207,329,348]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="16" pageNumber="149" unit="km" value="15.0">15 km</quantity>
S Iskander station, 
<geoCoordinate id="EE71A4B9FFD8FF84FEE8FEFAFE59FEA7" box="[383,501,329,348]" degrees="38" direction="north" minutes="56" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="29.4" value="38.9415">38°56′29.4″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FE6BFEFAFDDDFEA7" box="[508,625,329,348]" degrees="55" direction="east" minutes="55" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="42.9" value="55.928585">55°55′42.9″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FDECFEFAFD7AFEA7" box="[635,726,329,348]" pageId="16" pageNumber="149" value="1953-06-13">13.vi.1953</date>
, O. Kryzhanovskij leg., 
<specimenCount id="9D4309F7FFD8FF84FED6FED7FECCFE8D" box="[321,352,349,375]" pageId="16" pageNumber="149" type="female">1 ♀</specimenCount>
; foothills of N slope of Syunt-Khasardag Range, 
<geoCoordinate id="EE71A4B9FFD8FF84FF00FECDFEA1FE6A" box="[151,269,382,401]" degrees="38" direction="north" minutes="34" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="58.7" value="38.58297">38°34′58.7″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FE85FECDFE2BFE6A" box="[274,391,382,401]" degrees="56" direction="east" minutes="26" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="12.3" value="56.436752">56°26′12.3″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FE07FECDFE49FE6A" box="[400,485,382,401]" pageId="16" pageNumber="149" value="1956-05-17">17.v.1956</date>
, G. Medvedev leg., 
<specimenCount id="9D4309F7FFD8FF84FD05FECDFD1EFE6A" box="[658,690,376,401]" pageId="16" pageNumber="149" type="female">1 ♀</specimenCount>
; Parhay gorge, 
<geoCoordinate id="EE71A4B9FFD8FF84FF47FE2AFEE9FE57" box="[208,325,409,428]" degrees="38" direction="north" minutes="28" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="50.4" value="38.480667">38°28′50.4″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FEDFFE2AFE11FE57" box="[328,445,409,428]" degrees="56" direction="east" minutes="14" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="57.8" value="56.24939">56°14′57.8″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FE54FE2AFDB4FE57" box="[451,536,409,428]" pageId="16" pageNumber="149" value="1981-05-25">25.v.1981</date>
,A. Gorochov, 1* J, 
<specimenCount id="9D4309F7FFD8FF84FD52FE2AFD5EFE50" box="[709,754,402,428]" pageId="16" pageNumber="149" type="female">2 ♀♀</specimenCount>
. 
<emphasis id="B9311E6CFFD8FF84FF00FE07FEA6FE3C" bold="true" box="[151,266,436,455]" pageId="16" pageNumber="149">AඌΗΚΗൺൻൺൽ:</emphasis>
<date id="FFFBE4BEFFD8FF84FE9AFE07FECBFE3C" box="[269,359,436,455]" pageId="16" pageNumber="149" value="1928-06-11">11.vi.1928</date>
, 
<collectionCode id="ED545ABBFFD8FF84FEF9FE07FED1FE3C" box="[366,381,436,455]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="149" type="Museum">V</collectionCode>
.Gussakovskij leg., 
<specimenCount id="9D4309F7FFD8FF84FD88FE07FDE0FE3D" box="[543,588,429,455]" pageId="16" pageNumber="149" type="female">2 ♀♀</specimenCount>
, 
<date id="FFFBE4BEFFD8FF84FDC2FE07FD02FE3C" box="[597,686,436,455]" pageId="16" pageNumber="149" value="1930-04-11">11.iv.1930</date>
, Shul’… [illegible handwriting], 
<specimenCount id="9D4309F7FFD8FF84FECBFE7DFED6FE1A" box="[348,378,456,481]" pageId="16" pageNumber="149" type="female">1 ♀</specimenCount>
, 
<date id="FFFBE4BEFFD8FF84FE14FE7DFE7BFE1A" box="[387,471,462,481]" pageId="16" pageNumber="149" value="1934-05-29">29.v.1934</date>
, 
<collectionCode id="ED545ABBFFD8FF84FE48FE7DFE42FE1A" box="[479,494,462,481]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="149" type="Museum">V</collectionCode>
.Popov leg., 
<specimenCount id="9D4309F7FFD8FF84FDC1FE7DFDD8FE1A" box="[598,628,456,481]" pageId="16" pageNumber="149" type="female">1 ♀</specimenCount>
;Archabil Distr. of Ashkhabad, Firyuza Settlm., 
<geoCoordinate id="EE71A4B9FFD8FF84FE31FE5AFDB7FE07" box="[422,539,489,508]" degrees="37" direction="north" minutes="54" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="49.8" value="37.913834">37°54′49.8″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FDB6FE5AFD3AFE07" box="[545,662,489,508]" degrees="58" direction="east" minutes="05" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="26.3" value="58.090637">58°05′26.3″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FD09FE5AFD59FE07" box="[670,757,489,508]" pageId="16" pageNumber="149" value="1972-05-21">21.v.1972</date>
, 
<collectionCode id="ED545ABBFFD8FF84FF00FDB7FF0AFDEC" box="[151,166,516,535]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="149" type="Museum">V</collectionCode>
. Golub leg., 1* 
<specimenCount id="9D4309F7FFD8FF84FEB2FE4EFE99FDED" box="[293,309,509,534]" pageId="16" pageNumber="149" type="female">♀</specimenCount>
. 
<emphasis id="B9311E6CFFD8FF84FEA9FDB7FE1AFDEC" bold="true" box="[318,438,516,535]" pageId="16" pageNumber="149">Lൾൻൺඉ Pඋඈඏ.:</emphasis>
Repetek railway station, 
<geoCoordinate id="EE71A4B9FFD8FF84FD13FDB7FD54FDEC" box="[644,760,516,535]" degrees="38" direction="north" minutes="33" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="45.0" value="38.5625">38°33′45.0″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FF00FDADFEA0FDCA" box="[151,268,542,561]" degrees="63" direction="east" minutes="10" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="37.9" value="63.177197">63°10′37.9″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FE81FDADFED8FDCA" box="[278,372,542,561]" pageId="16" pageNumber="149" value="1958-03-18">18.iii.1958</date>
, G. Medvedev leg., 1* 
<specimenCount id="9D4309F7FFD8FF84FDA8FDABFDE3FDCA" box="[575,591,536,561]" pageId="16" pageNumber="149" type="female">♀</specimenCount>
, 
<date id="FFFBE4BEFFD8FF84FDCDFDADFD00FDCA" box="[602,684,542,561]" pageId="16" pageNumber="149" value="1981-06-08">8.vi.1981</date>
, at light, S. Podlipaev, 
<specimenCount id="9D4309F7FFD8FF84FE9DFD8AFE85FDB0" box="[266,297,562,588]" pageId="16" pageNumber="149" type="female">1 ♀</specimenCount>
, 
<specimenCount id="9D4309F7FFD8FF84FEA5FD8AFE3EFDB7" box="[306,402,569,588]" pageId="16" pageNumber="149" type="generic">1 specimen</specimenCount>
of unknown sex. 
<emphasis id="B9311E6CFFD8FF84FDB2FD8AFD30FDB7" bold="true" box="[549,668,569,588]" pageId="16" pageNumber="149">Mൺඋඒ Pඋඈඏ.:</emphasis>
Bayramali Town, 
<geoCoordinate id="EE71A4B9FFD8FF84FF46FDE7FEEBFD9C" box="[209,327,596,615]" degrees="37" direction="north" minutes="37" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="00.0" value="37.616665">37°37′00.0″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FEDBFDE7FE6CFD9C" box="[332,448,596,615]" degrees="62" direction="east" minutes="09" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="00.0" value="62.15">62°09′00.0″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FE5EFDE7FDFDFD9C" box="[457,593,596,615]" pageId="16" pageNumber="149" value="1930-08-10" valueMax="1930-08-11" valueMin="1930-08-10">10–11.viii.1930</date>
, Bogush leg., 1* J; Badkhyz, 
<quantity id="4CBD6F9BFFD8FF84FF79FDDDFE88FD7A" box="[238,292,622,641]" metricMagnitude="4" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="149" unit="km" value="12.0">12 km</quantity>
W of Kala-i-Mor station, 
<geoCoordinate id="EE71A4B9FFD8FF84FD93FDDDFDD5FD7A" box="[516,633,622,641]" degrees="35" direction="north" minutes="39" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="19.9" value="35.65553">35°39′19.9″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FDE8FDDDFD59FD7A" box="[639,757,622,641]" degrees="62" direction="east" minutes="25" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="22.9" value="62.42303">62°25′22.9″E</geoCoordinate>
, 9,17. 
<date id="FFFBE4BEFFD8FF84FF55FD3AFEAEFD67" box="[194,258,649,668]" pageId="16" pageNumber="149" value="1957-04">iv.1957</date>
, G. Medvedev, 1* J, 
<specimenCount id="9D4309F7FFD8FF84FE51FD3AFE44FD60" box="[454,488,642,668]" pageId="16" pageNumber="149" type="female">1 ♀</specimenCount>
. 
<emphasis id="B9311E6CFFD8FF84FE64FD3AFF7FFD4C" bold="true" pageId="16" pageNumber="149">
<collectingCountry id="F35282EEFFD8FF84FE64FD3AFDDAFD67" box="[499,630,649,668]" name="Tajikistan" pageId="16" pageNumber="149">TAJIKISTAN</collectingCountry>
: SඈǤൽංඒඌΚൺඒൺ Pඋඈඏ.:
</emphasis>
Kanibadam Town, 
<geoCoordinate id="EE71A4B9FFD8FF84FEE2FD17FE46FD4C" box="[373,490,676,695]" degrees="40" direction="north" minutes="16" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="60.0" value="40.283333">40°16′60.0″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FE7AFD17FDCEFD4C" box="[493,610,676,695]" degrees="70" direction="east" minutes="25" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="00.0" value="70.416664">70°25′00.0″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FDFEFD17FD6AFD4C" box="[617,710,676,695]" pageId="16" pageNumber="149" value="1913-06-15">15.vi.1913</date>
, Minkevits leg., 1* J. 
<emphasis id="B9311E6CFFD8FF84FEB2FD0DFEF7FD2A" bold="true" box="[293,347,702,721]" pageId="16" pageNumber="149">
<collectingCountry id="F35282EEFFD8FF84FEB2FD0DFEF7FD2A" box="[293,347,702,721]" name="Iran" pageId="16" pageNumber="149">IRAN</collectingCountry>
</emphasis>
: 
<emphasis id="B9311E6CFFD8FF84FEF3FD0DFE51FD2A" bold="true" box="[356,509,702,721]" pageId="16" pageNumber="149">Gඈඅൾඌඍൺඇ Pඋඈඏ.:</emphasis>
Shahkuh Range near Shahrud, 
<geoCoordinate id="EE71A4B9FFD8FF84FF00FD6AFEA1FD17" box="[151,269,729,748]" degrees="36" direction="north" minutes="30" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="00.9" value="36.50025">36°30′00.9″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FE84FD6AFE24FD17" box="[275,392,729,748]" degrees="54" direction="east" minutes="24" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="21.3" value="54.40592">54°24′21.3″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FE06FD6AFE45FD17" box="[401,489,729,748]" pageId="16" pageNumber="149" value="1914-07-05">5.vii.1914</date>
, A-r Kiritshenko, 1* J. 
<emphasis id="B9311E6CFFD8FF84FD2BFD6AFED9FCFC" bold="true" pageId="16" pageNumber="149">Sංඌඍൺඇ ൺඇൽ BൺඅඎർΗൾඌඍൺඇ Pඋඈඏ.</emphasis>
, Taftan volcano, 
<geoCoordinate id="EE71A4B9FFD8FF84FD92FD47FDD7FCFC" box="[517,635,756,775]" degrees="28" direction="north" minutes="36" orientation="latitude" pageId="16" pageNumber="149" precision="1" seconds="00.0" value="28.6">28°36′00.0″N</geoCoordinate>
<geoCoordinate id="EE71A4B9FFD8FF84FDE8FD47FD59FCFC" box="[639,757,756,775]" degrees="61" direction="east" minutes="07" orientation="longitude" pageId="16" pageNumber="149" precision="1" seconds="57.0" value="61.1325">61°07′57.0″E</geoCoordinate>
, 
<date id="FFFBE4BEFFD8FF84FF00FCBDFF41FCDA" box="[151,237,782,801]" pageId="16" pageNumber="149" value="1901-05-13">13.v.1901</date>
, A. Zarudny, 1* J, 1* 
<specimenCount id="9D4309F7FFD8FF84FE27FCBBFE6CFCDA" box="[432,448,776,801]" pageId="16" pageNumber="149" type="female">♀</specimenCount>
.
</paragraph>
<paragraph id="8BFAC27EFFD8FF84FF00FC89FE6EFBEA" blockId="16.[151,779,826,2065]" pageId="16" pageNumber="149">
<emphasis id="B9311E6CFFD8FF84FF00FC89FF7FFCAA" bold="true" box="[151,211,826,849]" pageId="16" pageNumber="149">Note.</emphasis>
Hඈൻൾ*©අൺඇൽඍ (1961) redescribed the external morphology of this species; for this reason, and because the external structure of the species almost completely corresponds to the description of the genus, I provide here only diagnostic characters necessary for comparison with other species of the genus and provide the first description of the male and female terminalia.
</paragraph>
</subSubSection>
<subSubSection id="C35F91F5FFD8FF86FF00FBA9FE44FECE" lastPageId="18" lastPageNumber="151" pageId="16" pageNumber="149" type="diagnosis">
<paragraph id="8BFAC27EFFD8FF84FF00FBA9FDD2F9CA" blockId="16.[151,779,826,2065]" pageId="16" pageNumber="149">
<emphasis id="B9311E6CFFD8FF84FF00FBA9FEABFBCA" bold="true" box="[151,263,1050,1073]" pageId="16" pageNumber="149">Diagnosis.</emphasis>
Body usually dark brown, with dull sheen, but head, pronotum and scutellum in some specimens blackish, hemelytra blackish-brown, less often entire dorsum almost black; some specimens with fairly strong sheen. Punctation on body as a whole slightly smaller than in other congeners, sometimes punctures rather large and deep. Pale stripes on lateral margins of hemelytra broad anteriorly, narrow in posterior about two-thirds or two-fifths, almost always reaching posterior margins of coria (
<figureCitation id="137EDEFBFFD8FF84FE25FAA9FDABFACA" box="[434,519,1306,1329]" captionStart="Fig" captionStartId="4.[151,182,1834,1853]" captionTargetBox="[164,1391,218,1805]" captionTargetId="figure-13@4.[151,1436,189,1820]" captionTargetPageId="4" captionText="Fig. 2. Exosehirus species, general appearance from dorsal side and type labels. A – E. validus (Jakovlev, 1877), holotype; B – E. elamensis sp. nov., holotype; C – E. essedonius sp. nov., holotype; D – E. marginatus (Signoret, 1881), without abdomen, from Turkey." figureDoi="http://doi.org/10.5281/zenodo.5037288" httpUri="https://zenodo.org/record/5037288/files/figure.png" pageId="16" pageNumber="149">Fig. 2A</figureCitation>
). Posterior part of each stripe relatively wide like in 
<taxonomicName id="4C45B9FDFFD8FF84FE2CFA89FDEBFAAA" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" box="[443,583,1338,1361]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="149" phylum="Arthropoda" rank="species" species="essedonius" status="sp. nov.">
<emphasis id="B9311E6CFFD8FF84FE2CFA89FDEBFAAA" box="[443,583,1338,1361]" italics="true" pageId="16" pageNumber="149">E. essedonius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFD8FF84FDDBFA89FD30FAAA" box="[588,668,1338,1361]" pageId="16" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
or extremely narrow, disappearing like in 
<taxonomicName id="4C45B9FDFFD8FF84FE79FAE9FDC3FA8A" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" box="[494,623,1370,1393]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="149" phylum="Arthropoda" rank="species" species="elamensis" status="sp. nov.">
<emphasis id="B9311E6CFFD8FF84FE79FAE9FDC3FA8A" box="[494,623,1370,1393]" italics="true" pageId="16" pageNumber="149">E. elamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFD8FF84FDE4FAE9FD6EFA8A" box="[627,706,1370,1393]" pageId="16" pageNumber="149" rank="species">sp. nov.</taxonomicNameLabel>
Eyes relatively large. Preocular part of head usually rather smoothly rounded (
<figureCitation id="137EDEFBFFD8FF84FE8CFA29FEDDFA4A" box="[283,369,1434,1457]" captionStart="Fig" captionStartId="5.[150,181,2008,2027]" captionTargetBox="[150,1435,204,1928]" captionTargetId="figure-11@5.[150,1435,189,1928]" captionTargetPageId="5" captionText="Fig. 3. Head (A–F), pronotum (G–K) and external scent efferent system (L) of Exosehirus species.A, G, L – E. validus (Jakovlev, 1877); B, H – E. sargon Linnavuori, 1984; C, I – E. elamensis sp. nov.; D, E, J – E. essedonius sp. nov.; F, K – E. marginatus (Signoret, 1881). Scale bars: 0.25 mm (A, C–F, L) and 0.5 mm (G, I–K). B and H after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037290" httpUri="https://zenodo.org/record/5037290/files/figure.png" pageId="16" pageNumber="149">Fig. 3A</figureCitation>
). Anterolateral margins of pronotum moderately diverging posteraid (
<figureCitation id="137EDEFBFFD8FF84FE48FA09FD83FA2A" box="[479,559,1466,1489]" captionStart="Fig" captionStartId="5.[150,181,2008,2027]" captionTargetBox="[150,1435,204,1928]" captionTargetId="figure-11@5.[150,1435,189,1928]" captionTargetPageId="5" captionText="Fig. 3. Head (A–F), pronotum (G–K) and external scent efferent system (L) of Exosehirus species.A, G, L – E. validus (Jakovlev, 1877); B, H – E. sargon Linnavuori, 1984; C, I – E. elamensis sp. nov.; D, E, J – E. essedonius sp. nov.; F, K – E. marginatus (Signoret, 1881). Scale bars: 0.25 mm (A, C–F, L) and 0.5 mm (G, I–K). B and H after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037290" httpUri="https://zenodo.org/record/5037290/files/figure.png" pageId="16" pageNumber="149">Fig. 3G</figureCitation>
). Apex of scutellum narrower than in 
<taxonomicName id="4C45B9FDFFD8FF84FEDDFA69FE71FA0A" baseAuthorityName="Signoret" baseAuthorityYear="1881" box="[330,477,1498,1521]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="149" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B9311E6CFFD8FF84FEDDFA69FE71FA0A" box="[330,477,1498,1521]" italics="true" pageId="16" pageNumber="149">E. marginatus</emphasis>
</taxonomicName>
. Peritreme of metathoracic scent glands slightly widened on mesal part. Evaporatorium with only several scattered punctures (
<figureCitation id="137EDEFBFFD8FF84FDB5F9A9FDDDF9CA" box="[546,625,1562,1585]" captionStart="Fig" captionStartId="5.[150,181,2008,2027]" captionTargetBox="[150,1435,204,1928]" captionTargetId="figure-11@5.[150,1435,189,1928]" captionTargetPageId="5" captionText="Fig. 3. Head (A–F), pronotum (G–K) and external scent efferent system (L) of Exosehirus species.A, G, L – E. validus (Jakovlev, 1877); B, H – E. sargon Linnavuori, 1984; C, I – E. elamensis sp. nov.; D, E, J – E. essedonius sp. nov.; F, K – E. marginatus (Signoret, 1881). Scale bars: 0.25 mm (A, C–F, L) and 0.5 mm (G, I–K). B and H after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037290" httpUri="https://zenodo.org/record/5037290/files/figure.png" pageId="16" pageNumber="149">Fig. 3L</figureCitation>
).
</paragraph>
<paragraph id="8BFAC27EFFD8FF84FF22F989FDAEF8AA" blockId="16.[151,779,826,2065]" pageId="16" pageNumber="149">
<emphasis id="B9311E6CFFD8FF84FF22F989FE86F9AA" bold="true" box="[181,298,1594,1617]" italics="true" pageId="16" pageNumber="149">Pygophore</emphasis>
(
<figureCitation id="137EDEFBFFD8FF84FEA1F989FE0AF9AA" box="[310,422,1594,1617]" captionStart="Fig" captionStartId="7.[151,182,1951,1970]" captionTargetBox="[153,1433,194,1836]" captionTargetId="figure-11@7.[150,1435,189,1852]" captionTargetPageId="7" captionText="Fig. 4. Pygophore, paramere and aedeagus of Exosehirus species.A, F, G, H – E. validus (Jakovlev, 1877); B, K – E. sargon Linnavuori, 1984; C – E. elamensis sp. nov.; D, I – E. essedonius sp. nov.; E, J – E. marginatus (Signoret, 1881). Pygophore in ventrocaudal (A–E) and dorsal (F) view; paramere in lateral (G) and dorsal (H–J) view; aedeagus in lateral view (K). Abbreviations: d.inf – dorsal infolding of pygophore; l.inf – lateral infolding of pygophore; l.r – lateral ridge of subgenital plate; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding of pygophore. Scale bars: 0.25 mm (A, C, D, E, F) and 0.14 mm (G–J). B and K after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037292" httpUri="https://zenodo.org/record/5037292/files/figure.png" pageId="16" pageNumber="149">Figs 4A, F</figureCitation>
). Notch in middle of posteroventral margin of pygophore slightly deeper and wider than in other congeners. Posterodorsal margin and mesal parts of posterolateral margins of pygophore and dorsal infolding along midline narrowly desclerotised. Dorsal infolding without depressions. Narrow depressed areas of lateral infoldings near margin of genital opening shallower than in 
<taxonomicName id="4C45B9FDFFD8FF84FF26F8A9FEE9F8CA" baseAuthorityName="Signoret" baseAuthorityYear="1881" box="[177,325,1818,1841]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="16" pageNumber="149" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B9311E6CFFD8FF84FF26F8A9FEE9F8CA" box="[177,325,1818,1841]" italics="true" pageId="16" pageNumber="149">E. marginatus</emphasis>
</taxonomicName>
and strongly sclerotised. Genital opening with very weak paramere sockets.
</paragraph>
<paragraph id="8BFAC27EFFD8FF84FF22F8E9FD2DF7EA" blockId="16.[151,779,826,2065]" pageId="16" pageNumber="149">
<emphasis id="B9311E6CFFD8FF84FF22F8E9FE80F88A" bold="true" box="[181,300,1882,1905]" italics="true" pageId="16" pageNumber="149">Parameres</emphasis>
(
<figureCitation id="137EDEFBFFD8FF84FEAEF8E9FE1DF88A" box="[313,433,1882,1905]" captionStart="Fig" captionStartId="7.[151,182,1951,1970]" captionTargetBox="[153,1433,194,1836]" captionTargetId="figure-11@7.[150,1435,189,1852]" captionTargetPageId="7" captionText="Fig. 4. Pygophore, paramere and aedeagus of Exosehirus species.A, F, G, H – E. validus (Jakovlev, 1877); B, K – E. sargon Linnavuori, 1984; C – E. elamensis sp. nov.; D, I – E. essedonius sp. nov.; E, J – E. marginatus (Signoret, 1881). Pygophore in ventrocaudal (A–E) and dorsal (F) view; paramere in lateral (G) and dorsal (H–J) view; aedeagus in lateral view (K). Abbreviations: d.inf – dorsal infolding of pygophore; l.inf – lateral infolding of pygophore; l.r – lateral ridge of subgenital plate; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding of pygophore. Scale bars: 0.25 mm (A, C, D, E, F) and 0.14 mm (G–J). B and K after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037292" httpUri="https://zenodo.org/record/5037292/files/figure.png" pageId="16" pageNumber="149">Figs 4G, H</figureCitation>
). Posterior margin of paramere corpus relatively deeply concave. Dorsal wall of paramere corpus at base of hypophysis with longitudinal, strongly sclerotised tubercle (clearly visible dorsally, especially so from ventral side). Hypophysis long; its extreme apex strongly narrowed and curved anteriad, sharp.
</paragraph>
<paragraph id="8BFAC27EFFD8FF84FCCDFF0DFB30FEAE" blockId="16.[828,1437,190,2069]" pageId="16" pageNumber="149">
<emphasis id="B9311E6CFFD8FF84FCCDFF0DFC69FF2E" bold="true" box="[858,965,190,213]" italics="true" pageId="16" pageNumber="149">Aedeagus</emphasis>
(
<figureCitation id="137EDEFBFFD8FF84FC43FF0DFBE2FF2E" box="[980,1102,190,213]" captionStart="Fig" captionStartId="8.[151,182,2030,2049]" captionTargetBox="[203,1427,417,1998]" captionTargetId="figure-13@8.[151,1436,411,2018]" captionTargetPageId="8" captionText="Fig. 5. Uninflated aedeagi of Exosehirus species.A, B – E. validus (Jakovlev, 1877); C, D – E. essedonius sp. nov. Lateral (A, C) and ventral (B, D) view. Abbreviations: vl.l – ventrolateral lobe of conjunctiva; dl.l – dorsolateral lobe; da.l – dorsoapical lobe. Scale bar: 0.14 mm." figureDoi="http://doi.org/10.5281/zenodo.5037294" httpUri="https://zenodo.org/record/5037294/files/figure.png" pageId="16" pageNumber="149">Figs 5A, B</figureCitation>
, 
<figureCitation id="137EDEFBFFD8FF84FBCDFF0DFB0FFF2E" box="[1114,1187,190,213]" captionStart="Fig" captionStartId="9.[151,183,1535,1554]" captionTargetBox="[154,1434,208,1514]" captionTargetId="figure-270@9.[151,1436,189,1518]" captionTargetPageId="9" captionText="Fig. 6. Wet preparations of conjunctival lobes of Exosehirus species. A–C – E. validus (Jakovlev, 1877); D–F – E. sargon Linnavuori, 1984; G–I – E. elamensis sp. nov.; J–L – E. essedonius sp. nov. Ventrolateral lobe (A, D, G, J); dorsolateral lobe (B, E, H, K); dorsoapical lobe (C, F, I, L). Scale bar: 0.14 mm. D–F after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037296" httpUri="https://zenodo.org/record/5037296/files/figure.png" pageId="16" pageNumber="149">6 A–C</figureCitation>
, 
<figureCitation id="137EDEFBFFD8FF84FB39FF0DFB11FF2E" box="[1198,1213,190,213]" captionStart="Fig" captionStartId="12.[943,974,1953,1972]" captionTargetBox="[156,1438,197,2079]" captionTargetId="figure-13@12.[151,1436,189,2079]" captionTargetPageId="12" captionText="Fig. 8. Completely inflated aedeagus of Exosehirus vali- dus (Jakovlev, 1877). A – ventral view; B – lateral view; C – dorsal view.Abbreviations: vl.l – ventrolateral lobe of conjunctiva;dl.l – dorsolateral lobe; da.l – dorsoapical lobe. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5037300" httpUri="https://zenodo.org/record/5037300/files/figure.png" pageId="16" pageNumber="149">8</figureCitation>
). Ventrolateral walls of theca with a pair of small conical tubercles at border of sclerotised and narrow apical membranous parts of walls. Ventral side of apical thecal margin medially with a small transverse membranous tubercle.
</paragraph>
<paragraph id="8BFAC27EFFD8FF84FCCDFEEDFBD9F9EE" blockId="16.[828,1437,190,2069]" pageId="16" pageNumber="149">Conjunctiva slightly shorter than theca and phallobase combined, tapering apically, slightly curved ventrally. Basal part of conjunctiva wide, long; apical part narrow, short. Basal part of conjunctiva ventrally with ventrolateral lobes; their bases very wide, long, occupying most of ventral and lateral walls of basal part of conjunctiva. On ventral side, these lobes rather wide, with rounded apices, bordered by thin, highly sclerotised band with denticles directed inward; mesal margins of lobes with wider, smooth, slightly less sclerotised band nearly reaching base of apical part of conjunctiva; both left and right bands slightly converging and tapering towards apex of conjunctiva. In lateral view, ventrolateral lobes triangularly tapering, directed ventrally and towards base of conjunctiva; anterior (facing base of conjunctiva) wall of each lobe with thin sclerotised band passing into serrate band bordering apex of lobe. Dorsolateral lobes located distally on basal part of conjunctiva, rather long, finger-shaped, curved, directed anteriad (towards base of conjunctiva) and slightly outward. Mesal wall of each lobe with longitudinal, long, wide sclerite protruding far beyond apex of lobe; dorsal surface and mesal margin of each sclerite covered with extremely small denticles; apex of sclerite broadly rounded. Dorsomesal wall of each dorsolateral lobe basally with short, wide, weakly sclerotised band, strongly narrowing proximally, dilating distally. Dorsoapical lobes lying on dorsolateral walls of apical part of conjunctiva subapically. Membranous parts of these lobes short, conical, directed dorsolaterally; posterior (facing apex of conjunctiva) wall of each lobe with spicule, that being long, thin strongly curved in middle, directed anteriorly (to base of conjunctiva) and dorsally; apices of both spicules almost parallel. Vesica lying on ventral wall of apical part of conjunctiva distal to bases of dorsoapical lobes. Its walls membranous, ventral wall at base with membranous tubercle having rectangular shape in ventral view, located at acute angle to conjunctival wall. Vesica gradually narrowing apically; its extreme apex elongated into short filament.</paragraph>
<paragraph id="8BFAC27EFFD8FF84FCCDF9ADFB17F86E" blockId="16.[828,1437,190,2069]" pageId="16" pageNumber="149">
<emphasis id="B9311E6CFFD8FF84FCCDF9ADFB85F9CE" bold="true" box="[858,1065,1566,1589]" italics="true" pageId="16" pageNumber="149">Female terminalia</emphasis>
(
<figureCitation id="137EDEFBFFD8FF84FBADF9ADFB3CF9CE" box="[1082,1168,1566,1589]" captionStart="Fig" captionStartId="19.[151,182,1978,1997]" captionTargetBox="[162,1434,196,1904]" captionTargetId="figure-10@19.[150,1435,189,1915]" captionTargetPageId="19" captionText="Fig. 12. Female terminalia of Exosehirus validus (Jakovlev, 1877).A – dorsal view; B – ventral view.Abbreviations: a.s – arcuate sclerites; a.l.g – anterior limb of gonangulum; f.r – first rami; g.c – gynatrial cone; gc.I – gonocoxites I; gc.II – gonocoxites II; gp.I – gonapophyses I; gp.II – gonapophyses II; p.i.m – posterior intervalvular membrane; p.l.g – posterior limb of gonangulum; p.p – pouches of posterior part of gynatrial sac; pt.IX – patatergites IX; r.s – ring sclerites; s.r – second rami; v.f – ventral fold of posterior intervalvular membrane. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5037312" httpUri="https://zenodo.org/record/5037312/files/figure.png" pageId="16" pageNumber="149">Figs 12</figureCitation>
, 
<figureCitation id="137EDEFBFFD8FF84FB09F9ADFB17F9CE" box="[1182,1211,1566,1589]" captionStart="Fig" captionStartId="20.[151,182,2032,2051]" captionTargetBox="[206,1365,190,1934]" captionTargetId="figure-13@20.[186,1402,165,1953]" captionTargetPageId="20" captionText="Fig. 13. Posterior part of gynatrial sac of Exosehirus validus (Jakovlev, 1877). A – dorsoposterior view; B – dorsolateral view. Abbreviation: g.c – gynatrial cone. Scale bar: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5037314" httpUri="https://zenodo.org/record/5037314/files/figure.png" pageId="16" pageNumber="149">13</figureCitation>
). Notch of posterior margin of sternite VII parabolic, slightly tapering anteriorly. Common posterior margin of paratergites VIII slightly convex in middle, without notch. Paratergites IX large.Anterior limb of gonangulum long, very thin anteriorly. Gonocoxites I with straight posterior margins lying in one line, with acute, non-rounded lateral angles; their ventral surfaces weakly, evenly convex, finely rugose, without rib along mesal margins. Mesal parts of gonocoxites II relatively narrow, longer than their width, with sharp, long anteromesal angles; lateral parts thin, very long, slightly widened at extreme lateral ends, those being not curved inward.
</paragraph>
<paragraph id="8BFAC27EFFD8FF84FCCDF82DFA30F7EE" blockId="16.[828,1437,190,2069]" pageId="16" pageNumber="149">Posterior intervalvular membrane very long; transverse fold on its ventral surface located behind its anterior margin, with middle part forming long wide triangular projection directed posteriad. Gonapophyses II overlying</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FF0DFA30FE4E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
and almost completely covering posterior intervalvular Aඐൺඅ 2008: Khorasan Razavi Province, Shirvan), southern membrane; their posterior margins concave, anterior 
<collectingCountry id="F35282EEFFD9FF85FCABFF6DFC12FF0E" box="[828,958,222,245]" name="Kazakhstan" pageId="17" pageNumber="150">Kazakhstan</collectingCountry>
, Kirgizia (Oඌ*üൺඇංඇ 1910: Osh; Pඎඍඌ*üκඈඏ margins straight laterally, strongly convex mesally, mesal 1965: Fergana Range, Chatkal Range), 
<collectingCountry id="F35282EEFFD9FF85FB77FF4DFAE5FEEE" box="[1248,1353,254,277]" name="Tajikistan" pageId="17" pageNumber="150">Tajikistan</collectingCountry>
(Oඌ*üൺmargins straight, posteriomesal angles elongate into acute ඇංඇ 1910: “Ura-Tyube” [Istaravshan]; Kං*©ංඍඌ*üൾඇκඈ 1964: apices directed posteromesad. Gonapophyses I wide, with “Kshtut” [tributary of Zeravshan River”]), 
<collectingCountry id="F35282EEFFD9FF85FA9EFE8DFA30FEAE" box="[1289,1436,318,341]" name="Turkmenistan" pageId="17" pageNumber="150">Turkmenistan</collectingCountry>
contiguous, short, triangular apices; their ventral margins (Pඎඍඌ*üκඈඏ 1965), 
<collectingCountry id="F35282EEFFD9FF85FB99FEEDFB2AFE8E" box="[1038,1158,350,373]" name="Uzbekistan" pageId="17" pageNumber="150">Uzbekistan</collectingCountry>
(Oඌ*üൺඇංඇ 1910: Andizhan, armed with thin sclerotised band with lateral end being Dzhizak; Kං*©ංඍඌ*üൾඇκඈ 1964: Tashkent, Changir near connected to anterior angle gonocoxite I. Posterior ends “Khatyrchi [Yangirabad]”, Samarkand, Guzar, Yargak,
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FE0DFBD2FE2E" blockId="17.[151,1468,190,2069]" box="[151,1150,446,469]" pageId="17" pageNumber="150">of first rami fused with anterior limbs of gonangula; inner Ispaskan [Izboskan], Termez).</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FE6DFA30FD8E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
margin of each first ramus with thin groove clamping The record for northern 
<collectingCountry id="F35282EEFFD9FF85FBEEFE6DFB0AFE0E" box="[1145,1190,478,501]" name="Iraq" pageId="17" pageNumber="150">Iraq</collectingCountry>
(Hඈൻൾ*©අൺඇൽඍ 1961) second ramus along its entire length. probably concerns 
<taxonomicName id="4C45B9FDFFD9FF85FB9CFE4DFBD9FDEE" authorityName="Linnavuori" authorityYear="1984" box="[1035,1141,510,533]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="sargon">
<emphasis id="B9311E6CFFD9FF85FB9CFE4DFBD9FDEE" box="[1035,1141,510,533]" italics="true" pageId="17" pageNumber="150">E. sargon</emphasis>
</taxonomicName>
; for Kazakhstanian Atyrau Anterior part of gynatrial sac large, with convex anterior Province, Lake Inder (Kං*©ංඍඌ*üൾඇκඈ 1954) and Kyzylorda and lateral margins, tapering posteriorly, there its lateral Provence, “Yani-Kyrgan” [Zhanakorgan] (Oඌ*üൺඇංඇ 1910), margins almost straight. Arcuate sclerites reinforcing base to 
<taxonomicName id="4C45B9FDFFD9FF85FCC2FDEDFC49FD8E" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" box="[853,997,606,629]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="essedonius" status="sp. nov.">
<emphasis id="B9311E6CFFD9FF85FCC2FDEDFC49FD8E" box="[853,997,606,629]" italics="true" pageId="17" pageNumber="150">E. essedonius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFD9FF85FC7DFDEDFB97FD8E" box="[1002,1083,606,629]" pageId="17" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
The record for Lorestan Province
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FDCDFAC0FD6E" blockId="17.[151,1468,190,2069]" box="[151,1388,638,661]" pageId="17" pageNumber="150">
of gynatrial cone relatively small, narrow, without lamellar of 
<collectingCountry id="F35282EEFFD9FF85FCCEFDCDFC28FD6E" box="[857,900,638,661]" name="Iran" pageId="17" pageNumber="150">Iran</collectingCountry>
(G*üൺ*üൺ*©ං et al. 2009) raise some doubts.
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FD2DFD54FD4E" blockId="17.[151,1468,190,2069]" box="[151,760,670,693]" pageId="17" pageNumber="150">outgrowths. Gynatrial cone posteriorly transverse, short,</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FD0DFA94FD2E" blockId="17.[151,1468,190,2069]" box="[151,1336,702,725]" pageId="17" pageNumber="150">
wide, with two wide triangular pouches on sides; dorsal 
<taxonomicName id="4C45B9FDFFD9FF85FC08FD0DFA94FD2E" authority="Linnavuori, 1984" authorityName="Linnavuori" authorityYear="1984" box="[927,1336,702,725]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="sargon">
<emphasis id="B9311E6CFFD9FF85FC08FD0DFA94FD2E" bold="true" box="[927,1336,702,725]" pageId="17" pageNumber="150">
<emphasis id="B9311E6CFFD9FF85FC08FD0DFBC3FD2E" bold="true" box="[927,1135,702,725]" italics="true" pageId="17" pageNumber="150">Exosehirus sargon</emphasis>
<bibRefCitation id="EFD4BF8FFFD9FF85FBE2FD0DFA94FD2E" author="LINNAVUORI R. E." box="[1141,1336,702,725]" pageId="17" pageNumber="150" pagination="1 - 59" refId="ref19171" refString="LINNAVUORI R. E. 1984: New species of Hemiptera Heteroptera from Iraq and the adjacent countries. Acta Entomologica Fennica 44: 1 - 59." type="journal article" year="1984">Linnavuori, 1984</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FD6DFB72FD14" blockId="17.[151,1468,190,2069]" box="[151,1246,732,757]" pageId="17" pageNumber="150">
margin of each pouch posteriorly with small rounded pro- (
<figureCitation id="137EDEFBFFD9FF85FB97FD6FFBCCFD14" box="[1024,1120,732,751]" captionStart="Fig" captionStartId="5.[150,181,2008,2027]" captionTargetBox="[150,1435,204,1928]" captionTargetId="figure-11@5.[150,1435,189,1928]" captionTargetPageId="5" captionText="Fig. 3. Head (A–F), pronotum (G–K) and external scent efferent system (L) of Exosehirus species.A, G, L – E. validus (Jakovlev, 1877); B, H – E. sargon Linnavuori, 1984; C, I – E. elamensis sp. nov.; D, E, J – E. essedonius sp. nov.; F, K – E. marginatus (Signoret, 1881). Scale bars: 0.25 mm (A, C–F, L) and 0.5 mm (G, I–K). B and H after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037290" httpUri="https://zenodo.org/record/5037290/files/figure.png" pageId="17" pageNumber="150">Figs 3B, H</figureCitation>
, 
<figureCitation id="137EDEFBFFD9FF85FBFEFD6FFB31FD14" box="[1129,1181,732,751]" captionStart="Fig" captionStartId="7.[151,182,1951,1970]" captionTargetBox="[153,1433,194,1836]" captionTargetId="figure-11@7.[150,1435,189,1852]" captionTargetPageId="7" captionText="Fig. 4. Pygophore, paramere and aedeagus of Exosehirus species.A, F, G, H – E. validus (Jakovlev, 1877); B, K – E. sargon Linnavuori, 1984; C – E. elamensis sp. nov.; D, I – E. essedonius sp. nov.; E, J – E. marginatus (Signoret, 1881). Pygophore in ventrocaudal (A–E) and dorsal (F) view; paramere in lateral (G) and dorsal (H–J) view; aedeagus in lateral view (K). Abbreviations: d.inf – dorsal infolding of pygophore; l.inf – lateral infolding of pygophore; l.r – lateral ridge of subgenital plate; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding of pygophore. Scale bars: 0.25 mm (A, C, D, E, F) and 0.14 mm (G–J). B and K after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037292" httpUri="https://zenodo.org/record/5037292/files/figure.png" pageId="17" pageNumber="150">4B, K</figureCitation>
, 
<figureCitation id="137EDEFBFFD9FF85FB31FD6FFB74FD14" box="[1190,1240,732,751]" captionStart="Fig" captionStartId="9.[151,183,1535,1554]" captionTargetBox="[154,1434,208,1514]" captionTargetId="figure-270@9.[151,1436,189,1518]" captionTargetPageId="9" captionText="Fig. 6. Wet preparations of conjunctival lobes of Exosehirus species. A–C – E. validus (Jakovlev, 1877); D–F – E. sargon Linnavuori, 1984; G–I – E. elamensis sp. nov.; J–L – E. essedonius sp. nov. Ventrolateral lobe (A, D, G, J); dorsolateral lobe (B, E, H, K); dorsoapical lobe (C, F, I, L). Scale bar: 0.14 mm. D–F after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037296" httpUri="https://zenodo.org/record/5037296/files/figure.png" pageId="17" pageNumber="150">6D–F</figureCitation>
)
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FD4DFA2DFCE2" blockId="17.[151,1468,190,2069]" box="[151,1409,766,793]" pageId="17" pageNumber="150">
jection; anterior part of gynatrial cone longitudinal, short, 
<taxonomicName id="4C45B9FDFFD9FF85FCABFCB5FB20FCE2" authority="Linnavuori, 1984: 2" authorityName="Linnavuori" authorityPageNumber="2" authorityYear="1984" box="[828,1164,774,793]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="sargon">
<emphasis id="B9311E6CFFD9FF85FCABFCB5FC76FCE2" box="[828,986,774,793]" italics="true" pageId="17" pageNumber="150">Exosehirus sargon</emphasis>
<bibRefCitation id="EFD4BF8FFFD9FF85FC77FCB5FB20FCE2" author="LINNAVUORI R. E." box="[992,1164,774,793]" pageId="17" pageNumber="150" pagination="1 - 59" refId="ref19171" refString="LINNAVUORI R. E. 1984: New species of Hemiptera Heteroptera from Iraq and the adjacent countries. Acta Entomologica Fennica 44: 1 - 59." type="journal article" year="1984">Linnavuori, 1984: 2</bibRefCitation>
</taxonomicName>
; Lංඌ (1999: 223; 2006: 142).
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FCADFD54FCCE" blockId="17.[151,1468,190,2069]" box="[151,760,798,821]" pageId="17" pageNumber="150">narrow, tapering conically distad. Spermatheca attached to</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFC81FD54FCAE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
<emphasis id="B9311E6CFFD9FF85FCABFC81FBEFFCB2" bold="true" box="[828,1091,818,841]" pageId="17" pageNumber="150">Notes on morphology.</emphasis>
The only known specimen of gynatrial cone at its apex. Proximal part of spermathecal
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFCE1FD54FC8E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">this species, stored in the American Museum of Natural duct relatively short, wide, widening towards middle, with</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFCC1FD54FC6E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">History, was not available to me for study, and therefore annular folds on usually weakly sclerotised walls; middle</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFC21FD54FC4E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">here I list the characters of this species based on the oripart longest, slightly protruding beyond anterior margin</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFC01FC5AFC32" blockId="17.[151,1468,190,2069]" box="[828,1014,946,969]" pageId="17" pageNumber="150">ginal description.</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FC0DFD54FC2E" blockId="17.[151,1468,190,2069]" box="[151,760,958,981]" pageId="17" pageNumber="150">of gynatrial sac, with membranous thickened walls; distal</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCCDFC61FD54FC0E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">Body black, very shiny and very convex. Eyes smaller part very narrow, rather longer then proximal part of duct,</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFC41FD5BFBEE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
then in 
<taxonomicName id="4C45B9FDFFD9FF85FC10FC41FC43FBF2" box="[903,1007,1010,1033]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="B9311E6CFFD9FF85FC10FC41FC43FBF2" box="[903,1007,1010,1033]" italics="true" pageId="17" pageNumber="150">E. validus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C45B9FDFFD9FF85FC6DFC41FBD2FBF2" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" box="[1018,1150,1010,1033]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="elamensis" status="sp. nov.">
<emphasis id="B9311E6CFFD9FF85FC6DFC41FBA6FBF2" box="[1018,1034,1010,1033]" italics="true" pageId="17" pageNumber="150">E</emphasis>
. 
<emphasis id="B9311E6CFFD9FF85FB81FC41FBD2FBF2" box="[1046,1150,1010,1033]" italics="true" pageId="17" pageNumber="150">elamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFD9FF85FB14FC41FB7FFBF2" box="[1155,1235,1010,1033]" pageId="17" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C45B9FDFFD9FF85FA95FC41FA39FBF2" baseAuthorityName="Signoret" baseAuthorityYear="1881" box="[1282,1429,1010,1033]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="B9311E6CFFD9FF85FA95FC41FA39FBF2" box="[1282,1429,1010,1033]" italics="true" pageId="17" pageNumber="150">E. marginatus</emphasis>
</taxonomicName>
; with membranous thin walls. Proximal convoluted part of
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFBA1FD54FBCE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">preocular part of head longer and more sharply rounded spermathecal capsule relatively short, several times longer</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFB81FD54FBAE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
anteriorly then in other congeners (
<figureCitation id="137EDEFBFFD9FF85FB3BFB81FB52FBB2" box="[1196,1278,1074,1097]" captionStart="Fig" captionStartId="5.[150,181,2008,2027]" captionTargetBox="[150,1435,204,1928]" captionTargetId="figure-11@5.[150,1435,189,1928]" captionTargetPageId="5" captionText="Fig. 3. Head (A–F), pronotum (G–K) and external scent efferent system (L) of Exosehirus species.A, G, L – E. validus (Jakovlev, 1877); B, H – E. sargon Linnavuori, 1984; C, I – E. elamensis sp. nov.; D, E, J – E. essedonius sp. nov.; F, K – E. marginatus (Signoret, 1881). Scale bars: 0.25 mm (A, C–F, L) and 0.5 mm (G, I–K). B and H after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037290" httpUri="https://zenodo.org/record/5037290/files/figure.png" pageId="17" pageNumber="150">Fig. 3B</figureCitation>
). Anterolateral than diameter of spherical part. Dorsal wall of posterior part
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFBE1FA30FB92" blockId="17.[151,1468,190,2069]" box="[828,1436,1106,1129]" pageId="17" pageNumber="150">margins of pronotum moderately diverging posteriad,</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FBEDFD5BFB8E" blockId="17.[151,1468,190,2069]" box="[151,759,1118,1141]" pageId="17" pageNumber="150">of gynatrial sac flat, with deep transverse fold extending</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFBC1FD5BFB6E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">roundedly curvate, anterolateral angles broadly rounded along its entire width, lying at posterior margin of sac me-</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFB21FD54FB4E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
(
<figureCitation id="137EDEFBFFD9FF85FCD3FB21FC36FB52" box="[836,922,1170,1193]" captionStart="Fig" captionStartId="5.[150,181,2008,2027]" captionTargetBox="[150,1435,204,1928]" captionTargetId="figure-11@5.[150,1435,189,1928]" captionTargetPageId="5" captionText="Fig. 3. Head (A–F), pronotum (G–K) and external scent efferent system (L) of Exosehirus species.A, G, L – E. validus (Jakovlev, 1877); B, H – E. sargon Linnavuori, 1984; C, I – E. elamensis sp. nov.; D, E, J – E. essedonius sp. nov.; F, K – E. marginatus (Signoret, 1881). Scale bars: 0.25 mm (A, C–F, L) and 0.5 mm (G, I–K). B and H after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037290" httpUri="https://zenodo.org/record/5037290/files/figure.png" pageId="17" pageNumber="150">Fig. 3H</figureCitation>
). Disk of pronotum very convex, calli elevated; dially and at bases of its posterior pouches laterally, having
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFB01FD5BFB2E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">anterior margin, sides and disk behind calli densely and obtuse margin directed anteriad, overlying dorsal wall of</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFB61FD54FB0E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">coarsely punctate. Disc of scutellum densely and strongly sac. Posterior pouches located rather close to each other,</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFB41FD5BFAEE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">punctate and slightly rugose. Elytra densely and distinctly their posterior margins rounded, without membranous</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFAA1FD55FACE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">punctate. Pale stripes on lateral margins of hemelytra wide protrusion on mesal part, slightly concave on lateral part.</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFA81FD54FAAE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
in anterior two-thirds and very narrow posteriorly, reaching 
<emphasis id="B9311E6CFFD9FF85FF22FA8DFEFAFAAE" bold="true" box="[181,342,1342,1365]" italics="true" pageId="17" pageNumber="150">Measurements</emphasis>
(n = 12; mm). Body: length 4.25–5.75,
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFAE1FBFBFA92" blockId="17.[151,1468,190,2069]" box="[828,1111,1362,1385]" pageId="17" pageNumber="150">posterior margins of coria.</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FAEDFD5BFA8E" blockId="17.[151,1468,190,2069]" box="[151,759,1374,1397]" pageId="17" pageNumber="150">width 2.60–3.35; ratio body length / pronotum width</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCCDFAC1FD5BFA6E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">The brief description and drawings of the male termi- 1.78–1.98; head: length 0.90–1.12, width 1.08–1.36,</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFA21FD5BFA4E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">nalia provide little information. Judging by these data, the ratio width / length 1.20–1.29; synthlipsis 0.70–0.87;</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFA01FD54FA2E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
pygophore (
<figureCitation id="137EDEFBFFD9FF85FC2CFA01FBA1FA32" box="[955,1037,1458,1481]" captionStart="Fig" captionStartId="7.[151,182,1951,1970]" captionTargetBox="[153,1433,194,1836]" captionTargetId="figure-11@7.[150,1435,189,1852]" captionTargetPageId="7" captionText="Fig. 4. Pygophore, paramere and aedeagus of Exosehirus species.A, F, G, H – E. validus (Jakovlev, 1877); B, K – E. sargon Linnavuori, 1984; C – E. elamensis sp. nov.; D, I – E. essedonius sp. nov.; E, J – E. marginatus (Signoret, 1881). Pygophore in ventrocaudal (A–E) and dorsal (F) view; paramere in lateral (G) and dorsal (H–J) view; aedeagus in lateral view (K). Abbreviations: d.inf – dorsal infolding of pygophore; l.inf – lateral infolding of pygophore; l.r – lateral ridge of subgenital plate; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding of pygophore. Scale bars: 0.25 mm (A, C, D, E, F) and 0.14 mm (G–J). B and K after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037292" httpUri="https://zenodo.org/record/5037292/files/figure.png" pageId="17" pageNumber="150">Fig. 4B</figureCitation>
) has its ventral wall more convex and ocular index 3.31–3.70; length of antennal segments I–V:
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFA61FD54FA0E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">its posteroventral margin is less broadly notched than in 0.22–0.32: 0.31–0.43: 0.29–0.41: 0.45–0.59: 0.59–0.67;</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABFA41FA31F9F2" blockId="17.[151,1468,190,2069]" box="[828,1437,1522,1545]" pageId="17" pageNumber="150">
<taxonomicName id="4C45B9FDFFD9FF85FCABFA41FC0BF9F2" box="[828,935,1522,1545]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="B9311E6CFFD9FF85FCABFA41FC0BF9F2" box="[828,935,1522,1545]" italics="true" pageId="17" pageNumber="150">E. validus</emphasis>
</taxonomicName>
(
<figureCitation id="137EDEFBFFD9FF85FC20FA41FBA7F9F2" box="[951,1035,1522,1545]" captionStart="Fig" captionStartId="7.[151,182,1951,1970]" captionTargetBox="[153,1433,194,1836]" captionTargetId="figure-11@7.[150,1435,189,1852]" captionTargetPageId="7" captionText="Fig. 4. Pygophore, paramere and aedeagus of Exosehirus species.A, F, G, H – E. validus (Jakovlev, 1877); B, K – E. sargon Linnavuori, 1984; C – E. elamensis sp. nov.; D, I – E. essedonius sp. nov.; E, J – E. marginatus (Signoret, 1881). Pygophore in ventrocaudal (A–E) and dorsal (F) view; paramere in lateral (G) and dorsal (H–J) view; aedeagus in lateral view (K). Abbreviations: d.inf – dorsal infolding of pygophore; l.inf – lateral infolding of pygophore; l.r – lateral ridge of subgenital plate; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding of pygophore. Scale bars: 0.25 mm (A, C, D, E, F) and 0.14 mm (G–J). B and K after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037292" httpUri="https://zenodo.org/record/5037292/files/figure.png" pageId="17" pageNumber="150">Fig. 4A</figureCitation>
) and 
<taxonomicName id="4C45B9FDFFD9FF85FBDEFA41FB7CF9F2" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" box="[1097,1232,1522,1545]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="elamensis" status="sp. nov.">
<emphasis id="B9311E6CFFD9FF85FBDEFA41FB7CF9F2" box="[1097,1232,1522,1545]" italics="true" pageId="17" pageNumber="150">E. elamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFD9FF85FB4FFA41FA80F9F2" box="[1240,1324,1522,1545]" pageId="17" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="137EDEFBFFD9FF85FAACFA41FA3CF9F2" box="[1339,1424,1522,1545]" captionStart="Fig" captionStartId="7.[151,182,1951,1970]" captionTargetBox="[153,1433,194,1836]" captionTargetId="figure-11@7.[150,1435,189,1852]" captionTargetPageId="7" captionText="Fig. 4. Pygophore, paramere and aedeagus of Exosehirus species.A, F, G, H – E. validus (Jakovlev, 1877); B, K – E. sargon Linnavuori, 1984; C – E. elamensis sp. nov.; D, I – E. essedonius sp. nov.; E, J – E. marginatus (Signoret, 1881). Pygophore in ventrocaudal (A–E) and dorsal (F) view; paramere in lateral (G) and dorsal (H–J) view; aedeagus in lateral view (K). Abbreviations: d.inf – dorsal infolding of pygophore; l.inf – lateral infolding of pygophore; l.r – lateral ridge of subgenital plate; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding of pygophore. Scale bars: 0.25 mm (A, C, D, E, F) and 0.14 mm (G–J). B and K after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037292" httpUri="https://zenodo.org/record/5037292/files/figure.png" pageId="17" pageNumber="150">Fig. 4C</figureCitation>
).
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FF00FA4DFD5BF9EE" blockId="17.[151,1468,190,2069]" box="[151,759,1534,1557]" pageId="17" pageNumber="150">pronotum: length 1.10–1.53, width 2.33–3.03, ratio width /</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF9A1FD5BF9CE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
Parameres as in 
<taxonomicName id="4C45B9FDFFD9FF85FC64F9A1FBCDF9D2" box="[1011,1121,1554,1577]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="B9311E6CFFD9FF85FC64F9A1FBCDF9D2" box="[1011,1121,1554,1577]" italics="true" pageId="17" pageNumber="150">E. validus</emphasis>
</taxonomicName>
[i.e., probably with tubercle length 1.97–2.11; scutellum: length 1.78–2.40, width
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF981FDFFF9AE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
near base of hypophysis]. Vesica (
<figureCitation id="137EDEFBFFD9FF85FB30F981FB57F9B2" box="[1191,1275,1586,1609]" captionStart="Fig" captionStartId="7.[151,182,1951,1970]" captionTargetBox="[153,1433,194,1836]" captionTargetId="figure-11@7.[150,1435,189,1852]" captionTargetPageId="7" captionText="Fig. 4. Pygophore, paramere and aedeagus of Exosehirus species.A, F, G, H – E. validus (Jakovlev, 1877); B, K – E. sargon Linnavuori, 1984; C – E. elamensis sp. nov.; D, I – E. essedonius sp. nov.; E, J – E. marginatus (Signoret, 1881). Pygophore in ventrocaudal (A–E) and dorsal (F) view; paramere in lateral (G) and dorsal (H–J) view; aedeagus in lateral view (K). Abbreviations: d.inf – dorsal infolding of pygophore; l.inf – lateral infolding of pygophore; l.r – lateral ridge of subgenital plate; p.r – posterior ridge of subgenital plate; v.inf – ventral infolding of pygophore. Scale bars: 0.25 mm (A, C, D, E, F) and 0.14 mm (G–J). B and K after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037292" httpUri="https://zenodo.org/record/5037292/files/figure.png" pageId="17" pageNumber="150">Fig. 4K</figureCitation>
) very long and 1.53–2.05, ratio length / width 1.13–1.19.
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF9E1FD54F98E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
curved [assuming that it protrudes strongly from the theca 
<emphasis id="B9311E6CFFD9FF85FF22F9EDFE9CF98E" bold="true" box="[181,304,1630,1653]" pageId="17" pageNumber="150">
<emphasis id="B9311E6CFFD9FF85FF22F9EDFE84F98E" bold="true" box="[181,296,1630,1653]" italics="true" pageId="17" pageNumber="150">Variability</emphasis>
.
</emphasis>
In two examined females from Turkme-
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF9C1FD5BF96E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">not as a consequence of damage and incomplete retraction nistan (Kyzyl-Arvat and Repetek), the margins of the</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF921FD54F94E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">of the vesica; it is also important to note that the filiform ring sclerites are very wide and the middle part of the</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF901FD5BF92E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">distal part of the vesica is not shown in the figure]; each spermathecal duct is weakly sclerotised; one of them</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF961FD5BF90E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">ventrolateral lobe (“conjunctival appendage”, probably has the pouches of gynatrial cone with internal areas of</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF941FD5BF8EE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
denoted by the letter e in the original drawing; 
<figureCitation id="137EDEFBFFD9FF85FAD7F941FA38F8F2" box="[1344,1428,1778,1801]" captionStart="Fig" captionStartId="9.[151,183,1535,1554]" captionTargetBox="[154,1434,208,1514]" captionTargetId="figure-270@9.[151,1436,189,1518]" captionTargetPageId="9" captionText="Fig. 6. Wet preparations of conjunctival lobes of Exosehirus species. A–C – E. validus (Jakovlev, 1877); D–F – E. sargon Linnavuori, 1984; G–I – E. elamensis sp. nov.; J–L – E. essedonius sp. nov. Ventrolateral lobe (A, D, G, J); dorsolateral lobe (B, E, H, K); dorsoapical lobe (C, F, I, L). Scale bar: 0.14 mm. D–F after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037296" httpUri="https://zenodo.org/record/5037296/files/figure.png" pageId="17" pageNumber="150">Fig. 6D</figureCitation>
) sclerotisation; in another specimen, the middle part of
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF8A1FD54F8CE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">and dorsolateral lobe (“conjunctival appendage”, prothe spermathecal duct is somewhat shortened and slightly</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF881FE39F8AE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
bably denoted by the letter d; 
<figureCitation id="137EDEFBFFD9FF85FB1AF881FB4CF8B2" box="[1165,1248,1842,1865]" captionStart="Fig" captionStartId="9.[151,183,1535,1554]" captionTargetBox="[154,1434,208,1514]" captionTargetId="figure-270@9.[151,1436,189,1518]" captionTargetPageId="9" captionText="Fig. 6. Wet preparations of conjunctival lobes of Exosehirus species. A–C – E. validus (Jakovlev, 1877); D–F – E. sargon Linnavuori, 1984; G–I – E. elamensis sp. nov.; J–L – E. essedonius sp. nov. Ventrolateral lobe (A, D, G, J); dorsolateral lobe (B, E, H, K); dorsoapical lobe (C, F, I, L). Scale bar: 0.14 mm. D–F after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037296" httpUri="https://zenodo.org/record/5037296/files/figure.png" pageId="17" pageNumber="150">Fig. 6E</figureCitation>
) with sclerotised widening in the middle.
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF8E1FCAFF88E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
denticles on some common sclerotised base. In the latter 
<emphasis id="B9311E6CFFD9FF85FF00F8EDFEB9F88E" bold="true" box="[151,277,1886,1909]" pageId="17" pageNumber="150">Bionomics.</emphasis>
According to Pඎඍඌ*üκඈඏ (1965), the species
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF8C1FD5BF86E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
case, three sharp denticles are depicted, namely two large lives on 
<taxonomicName id="4C45B9FDFFD9FF85FF67F8CDFD82F86E" box="[240,558,1918,1941]" class="Magnoliopsida" family="Amaranthaceae" genus="Krascheninnikovia" kingdom="Plantae" order="Caryophyllales" pageId="17" pageNumber="150" phylum="Tracheophyta" rank="species" species="ceratoides">
<emphasis id="B9311E6CFFD9FF85FF67F8CDFD82F86E" box="[240,558,1918,1941]" italics="true" pageId="17" pageNumber="150">Krascheninnikovia ceratoides</emphasis>
</taxonomicName>
, less often on 
<emphasis id="B9311E6CFFD9FF85FD52F8CDFD5BF86E" box="[709,759,1918,1941]" italics="true" pageId="17" pageNumber="150">Atri-</emphasis>
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF821FD5BF84E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
curved and one small straight one. Spicules of 
<emphasis id="B9311E6CFFD9FF85FAA0F821FF52F84E" italics="true" pageId="17" pageNumber="150">
dorsoapiplex 
<taxonomicName id="4C45B9FDFFD9FF85FF5DF82DFF52F84E" box="[202,254,1950,1973]" class="Magnoliopsida" family="Amaranthaceae" genus="Atriplex" kingdom="Plantae" order="Caryophyllales" pageId="17" pageNumber="150" phylum="Tracheophyta" rank="species" species="cana">cana</taxonomicName>
</emphasis>
(both from 
<taxonomicName id="4C45B9FDFFD9FF85FEEAF82DFD8BF84E" box="[381,551,1950,1973]" class="Magnoliopsida" family="Amaranthaceae" kingdom="Plantae" order="Caryophyllales" pageId="17" pageNumber="150" phylum="Tracheophyta" rank="family">Amaranthaceae</taxonomicName>
), although this data
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF801FDA7F82E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
cal lobes (“aedeagal appendage”; 
<figureCitation id="137EDEFBFFD9FF85FB3DF801FB56F832" box="[1194,1274,1970,1993]" captionStart="Fig" captionStartId="9.[151,183,1535,1554]" captionTargetBox="[154,1434,208,1514]" captionTargetId="figure-270@9.[151,1436,189,1518]" captionTargetPageId="9" captionText="Fig. 6. Wet preparations of conjunctival lobes of Exosehirus species. A–C – E. validus (Jakovlev, 1877); D–F – E. sargon Linnavuori, 1984; G–I – E. elamensis sp. nov.; J–L – E. essedonius sp. nov. Ventrolateral lobe (A, D, G, J); dorsolateral lobe (B, E, H, K); dorsoapical lobe (C, F, I, L). Scale bar: 0.14 mm. D–F after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037296" httpUri="https://zenodo.org/record/5037296/files/figure.png" pageId="17" pageNumber="150">Fig. 6F</figureCitation>
) straight, wide may refer to 
<taxonomicName id="4C45B9FDFFD9FF85FEB7F80DFE1EF82E" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" box="[288,434,1982,2005]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="essedonius" status="sp. nov.">
<emphasis id="B9311E6CFFD9FF85FEB7F80DFE1EF82E" box="[288,434,1982,2005]" italics="true" pageId="17" pageNumber="150">E. essedonius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFD9FF85FE2FF80DFDA7F82E" box="[440,523,1982,2005]" pageId="17" pageNumber="150" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF861FCA0F80E" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
at base and narrowing distally, with sloping apex, longer 
<emphasis id="B9311E6CFFD9FF85FF00F86DFE8EF80E" bold="true" box="[151,290,2014,2037]" pageId="17" pageNumber="150">Distribution</emphasis>
(
<figureCitation id="137EDEFBFFD9FF85FEA3F86DFEDAF80E" box="[308,374,2014,2037]" captionStart="Fig" captionStartId="2.[151,182,802,821]" captionTargetBox="[151,1436,189,777]" captionTargetId="figure-712@2.[151,1436,189,777]" captionTargetPageId="2" captionText="Fig. 1. Distribution of Exosehirus species based on the examined material (coloured buttons) and literature (white buttons)." figureDoi="http://doi.org/10.5281/zenodo.5037284" httpUri="https://zenodo.org/record/5037284/files/figure.png" pageId="17" pageNumber="150">Fig. 1</figureCitation>
). 
<collectingCountry id="F35282EEFFD9FF85FE19F86DFDBCF80E" box="[398,528,2014,2037]" name="Afghanistan" pageId="17" pageNumber="150">Afghanistan</collectingCountry>
(Hඈൻൾ*©අൺඇൽඍ 1961:
</paragraph>
<paragraph id="8BFAC27EFFD9FF85FCABF841FC8CF7EE" blockId="17.[151,1468,190,2069]" pageId="17" pageNumber="150">
than in 
<taxonomicName id="4C45B9FDFFD9FF85FC1AF841FC55F7F2" box="[909,1017,2034,2057]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="17" pageNumber="150" phylum="Arthropoda" rank="species" species="validus">
<emphasis id="B9311E6CFFD9FF85FC1AF841FC55F7F2" box="[909,1017,2034,2057]" italics="true" pageId="17" pageNumber="150">E. validus</emphasis>
</taxonomicName>
(
<figureCitation id="137EDEFBFFD9FF85FB9FF841FBF1F7F2" box="[1032,1117,2034,2057]" captionStart="Fig" captionStartId="9.[151,183,1535,1554]" captionTargetBox="[154,1434,208,1514]" captionTargetId="figure-270@9.[151,1436,189,1518]" captionTargetPageId="9" captionText="Fig. 6. Wet preparations of conjunctival lobes of Exosehirus species. A–C – E. validus (Jakovlev, 1877); D–F – E. sargon Linnavuori, 1984; G–I – E. elamensis sp. nov.; J–L – E. essedonius sp. nov. Ventrolateral lobe (A, D, G, J); dorsolateral lobe (B, E, H, K); dorsoapical lobe (C, F, I, L). Scale bar: 0.14 mm. D–F after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037296" httpUri="https://zenodo.org/record/5037296/files/figure.png" pageId="17" pageNumber="150">Fig. 6C</figureCitation>
), but probably shorter than in Puli Khumri), northern 
<collectingCountry id="F35282EEFFD9FF85FE34F84DFE63F7EE" box="[419,463,2046,2069]" name="Iran" pageId="17" pageNumber="150">Iran</collectingCountry>
(
<typeStatus id="54FE7CDCFFD9FF85FE76F84DFDBDF7EE" box="[481,529,2046,2069]" pageId="17" pageNumber="150">type</typeStatus>
locality; Mඈൽൺ*©*©ൾඌ
</paragraph>
<paragraph id="8BFAC27EFFDAFF86FF00FF0DFEC6FEEE" blockId="18.[151,760,190,501]" pageId="18" pageNumber="151">
<taxonomicName id="4C45B9FDFFDAFF86FF00FF0DFEB2FF2E" authority="Gapon, 2021" authorityName="Gapon" authorityYear="2021" box="[151,286,190,213]" class="Insecta" family="Cydnidae" genus="Exosehirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="151" phylum="Arthropoda" rank="species" species="elamensis" status="sp. nov.">
<emphasis id="B9311E6CFFDAFF86FF00FF0DFEB2FF2E" box="[151,286,190,213]" italics="true" pageId="18" pageNumber="151">E. elamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A202A317FFDAFF86FEB2FF0DFED4FF2E" box="[293,376,190,213]" pageId="18" pageNumber="151" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="137EDEFBFFDAFF86FE11FF0DFE63FF2E" box="[390,463,190,213]" captionStart="Fig" captionStartId="9.[151,183,1535,1554]" captionTargetBox="[154,1434,208,1514]" captionTargetId="figure-270@9.[151,1436,189,1518]" captionTargetPageId="9" captionText="Fig. 6. Wet preparations of conjunctival lobes of Exosehirus species. A–C – E. validus (Jakovlev, 1877); D–F – E. sargon Linnavuori, 1984; G–I – E. elamensis sp. nov.; J–L – E. essedonius sp. nov. Ventrolateral lobe (A, D, G, J); dorsolateral lobe (B, E, H, K); dorsoapical lobe (C, F, I, L). Scale bar: 0.14 mm. D–F after Lංඇඇൺඏඎඈ*©ං (1984)." figureDoi="http://doi.org/10.5281/zenodo.5037296" httpUri="https://zenodo.org/record/5037296/files/figure.png" pageId="18" pageNumber="151">Fig. 6I</figureCitation>
) [judging by the size of the figure of the spicule relative to the figures of parts of other conjunctival lobes].
</paragraph>
<paragraph id="8BFAC27EFFDAFF86FF22FEADFE44FECE" blockId="18.[151,760,190,501]" box="[181,488,286,309]" pageId="18" pageNumber="151">Female terminalia unknown.</paragraph>
</subSubSection>
<subSubSection id="C35F91F5FFDAFF86FF22FE8DFD7EFE2E" pageId="18" pageNumber="151" type="description">
<paragraph id="8BFAC27EFFDAFF86FF22FE8DFD7EFE2E" blockId="18.[151,760,190,501]" pageId="18" pageNumber="151">
<emphasis id="B9311E6CFFDAFF86FF22FE8DFEFAFEAE" bold="true" box="[181,342,318,341]" italics="true" pageId="18" pageNumber="151">Measurements</emphasis>
(n = 1; mm). Body length 6.75; ratio body length / pronotum width 1.70; ratio head width / head length 1.22; ocular index 3.60; proportions between antennal segments I–V: 8: 12: 10: 16: 18; ratio width / length c. 2.00; ratio scutellum length / scutellum width 1.20.
</paragraph>
</subSubSection>
<subSubSection id="C35F91F5FFDAFF86FF00FE6DFD1EFE0E" box="[151,690,478,501]" pageId="18" pageNumber="151" type="distribution">
<paragraph id="8BFAC27EFFDAFF86FF00FE6DFD1EFE0E" blockId="18.[151,760,190,501]" box="[151,690,478,501]" pageId="18" pageNumber="151">
<emphasis id="B9311E6CFFDAFF86FF00FE6DFE86FE0E" bold="true" box="[151,298,478,501]" pageId="18" pageNumber="151">Distribution.</emphasis>
Northern 
<collectingCountry id="F35282EEFFDAFF86FE00FE6DFE6EFE0E" box="[407,450,478,501]" name="Iraq" pageId="18" pageNumber="151">Iraq</collectingCountry>
(Lංඇඇൺඏඎඈ*©ං 1984).
</paragraph>
</subSubSection>
</treatment>
</document>