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<document id="949B89F6BC3FE8A06B8D2574C0A526DD" ID-CLB-Dataset="26743" ID-DOI="10.11646/zootaxa.4608.3.5" ID-GBIF-Dataset="1c5094f8-baa5-479b-9a86-b05da2fb210f" ID-ISSN="1175-5326" ID-Zenodo-Dep="3066723" ID-ZooBank="5B31798B-FFA3-42F3-9F3B-DE81B1C318EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1558416183313" checkinUser="plazi" docAuthor="Kovačić, Marcelo, Bogorodsky, Sergey V. &amp; Mal, Ahmad O." docDate="2019" docId="03C987875F7FFFB7FF78F8E6FC51FC41" docLanguage="en" docName="zootaxa.4608.3.5.pdf" docOrigin="Zootaxa 4608 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hetereleotris aurantiaca Kovačić &amp; Bogorodsky &amp; Mal 2019, sp. nov." docType="treatment" docVersion="10" lastPageNumber="508" masterDocId="FFF0FFFF5F7CFFB0FFEFFFE9FFA9FFAE" masterDocTitle="Two new species of Hetereleotris (Perciformes: Gobiidae) from the Red Sea" masterLastPageNumber="516" masterPageNumber="501" pageNumber="504" updateTime="1698731353378" updateUser="ExternalLinkService">
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<mods:title id="57F82A884740E746976F9B4D98754117">Two new species of Hetereleotris (Perciformes: Gobiidae) from the Red Sea</mods:title>
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<mods:namePart id="4885A4AFCF50340B6F151203308D6696">Kovačić, Marcelo</mods:namePart>
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<mods:namePart id="F3EF169DC294064D1F31D7A0821B5EB8">Bogorodsky, Sergey V.</mods:namePart>
</mods:name>
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<mods:role id="86826532F12B003CBCBDBBF23D4DF41F">
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<mods:namePart id="93741E2E86FCEC877495ADD12056CC26">Mal, Ahmad O.</mods:namePart>
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<mods:typeOfResource id="1A5AC77168C874681D2060F4F2CCF983">text</mods:typeOfResource>
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<mods:title id="8AE42A8204D77390C4D0B5269AAF944B">Zootaxa</mods:title>
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<mods:part id="F1B9CF85D45378CAB5B896797FD3B6E0">
<mods:date id="6671A2E350F5B1A1D1055A80C454D69F">2019</mods:date>
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<mods:number id="EFC3C4DCBB07DF4C7DF922105AAD3129">2019-05-21</mods:number>
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<mods:number id="1BDDE95C67F6E33FB35B1C8E88B4A9B6">4608</mods:number>
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<mods:identifier id="4122FB1F960A2548EA564592D366784D" type="DOI">10.11646/zootaxa.4608.3.5</mods:identifier>
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<treatment id="03C987875F7FFFB7FF78F8E6FC51FC41" ID-DOI="http://doi.org/10.5281/zenodo.5696368" ID-GBIF-Taxon="157232717" ID-Zenodo-Dep="5696368" LSID="urn:lsid:plazi:treatment:03C987875F7FFFB7FF78F8E6FC51FC41" httpUri="http://treatment.plazi.org/id/03C987875F7FFFB7FF78F8E6FC51FC41" lastPageId="7" lastPageNumber="508" pageId="3" pageNumber="504">
<subSubSection id="C37A651A5F7FFFB3FF78F8E6FE2BF8E3" pageId="3" pageNumber="504" type="nomenclature">
<paragraph id="8BDF36915F7FFFB3FF78F8E6FCE4F884" blockId="3.[151,845,1807,1906]" box="[151,845,1807,1834]" pageId="3" pageNumber="504">
<heading id="D09781FD5F7FFFB3FF78F8E6FCE4F884" bold="true" box="[151,845,1807,1834]" fontSize="11" level="1" pageId="3" pageNumber="504" reason="1">
<emphasis id="B914EA835F7FFFB3FF78F8E6FCE4F884" bold="true" box="[151,845,1807,1834]" pageId="3" pageNumber="504">
<taxonomicName id="4C604D125F7FFFB3FF78F8E6FD4EF884" authority="Kovacic &amp; Bogorodsky" authorityName="Kovačić &amp; Bogorodsky &amp; Mal" authorityYear="2019" box="[151,743,1807,1834]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="504" phylum="Chordata" rank="species" species="aurantiaca" status="sp. nov.">
<emphasis id="B914EA835F7FFFB3FF78F8E6FE16F884" bold="true" box="[151,447,1807,1834]" italics="true" pageId="3" pageNumber="504">Hetereleotris aurantiaca</emphasis>
Kovačić &amp; Bogorodsky
</taxonomicName>
<taxonomicNameLabel id="A22757F85F7FFFB3FD00F8F9FCE4F884" box="[751,845,1808,1834]" pageId="3" pageNumber="504" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36915F7FFFB3FF78F8DDFE2BF8E3" blockId="3.[151,845,1807,1906]" box="[151,386,1843,1870]" pageId="3" pageNumber="504">Orange Goggle goby</paragraph>
</subSubSection>
<subSubSection id="C37A651A5F7FFFB3FF78F8BEFEECF8DC" box="[151,325,1879,1906]" pageId="3" pageNumber="504" type="description">
<paragraph id="8BDF36915F7FFFB3FF78F8BEFEECF8DC" blockId="3.[151,845,1807,1906]" box="[151,325,1879,1906]" pageId="3" pageNumber="504">
(
<figureCitation id="135B2A145F7FFFB3FF70F8BEFEA3F8DC" box="[159,266,1879,1906]" captionStart="FIGURE 1" captionStartId="6.[151,250,1898,1923]" captionTargetBox="[151,1436,251,1865]" captionTargetId="figure@6.[151,1436,269,1865]" captionTargetPageId="6" captionText="FIGURE 1. Hetereleotris aurantiaca sp. nov. SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). A: freshly collected specimen; B: preserved specimen; C: three peduncular scales marked with arrows. Photos by S.V. Bogorodsky (A), M. Kovačić (B, C)." httpUri="https://zenodo.org/record/3066725/files/figure.png" pageId="3" pageNumber="504">Figures 1</figureCitation>
&amp;
<figureCitation id="135B2A145F7FFFB3FEC2F8B1FE95F8DC" box="[301,316,1880,1906]" captionStart="FIGURE 2" captionStartId="7.[151,250,1923,1948]" captionTargetBox="[202,1436,1116,1855]" captionTargetId="figure@7.[151,1436,1047,1888]" captionTargetPageId="7" captionText="FIGURE 2. Hetereleotris aurantiaca sp. nov. Cephalic sensory papillae: SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/3066727/files/figure.png" pageId="3" pageNumber="504">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37A651A5F7FFFB3FF78F875FF7EF850" pageId="3" pageNumber="504" type="materials_examined">
<paragraph id="8BDF36915F7FFFB3FF78F875FF7EF850" blockId="3.[151,1437,1947,2046]" pageId="3" pageNumber="504">
<materialsCitation id="3B083CCC5F7FFFB3FF78F875FF7EF850" ID-GBIF-Occurrence="2243028727" collectingDate="2013-07-01" collectionCode="SMF" collectorName="T. J. Alpermann &amp; S. V. Bogorodsky" country="Saudi Arabia" county="Obhur creek" elevation="14" latitude="21.708612" location="Red Sea" longLatPrecision="1" longitude="39.095833" municipality="Jeddah" pageId="3" pageNumber="504" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DB88335F7FFFB3FF78F875FEADF818" box="[151,260,1948,1974]" pageId="3" pageNumber="504" type="holotype">
<emphasis id="B914EA835F7FFFB3FF78F875FEADF818" bold="true" box="[151,260,1948,1974]" pageId="3" pageNumber="504">Holotype</emphasis>
</typeStatus>
.
<collectionCode id="ED71AE545F7FFFB3FEFEF875FEE0F818" box="[273,329,1948,1974]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="504">SMF</collectionCode>
35966 (field number KAU13-37), female, 14.51 + 3.40 mm,
<location id="8EBF604A5F7FFFB3FC1CF872FBF8F818" LSID="urn:lsid:plazi:treatment:03C987875F7FFFB7FF78F8E6FC51FC41:8EBF604A5F7FFFB3FC1CF872FBF8F818" box="[1011,1105,1947,1974]" country="Saudi Arabia" county="Obhur creek" latitude="21.708612" longLatPrecision="1" longitude="39.095833" municipality="Jeddah" name="Red Sea" pageId="3" pageNumber="504">Red Sea</location>
,
<collectingCountry id="F37776015F7FFFB3FBB3F872FB46F81B" box="[1116,1263,1947,1973]" name="Saudi Arabia" pageId="3" pageNumber="504">Saudi Arabia</collectingCountry>
,
<collectingMunicipality id="6BBBACEB5F7FFFB3FB15F872FAE2F81B" box="[1274,1355,1947,1973]" pageId="3" pageNumber="504">Jeddah</collectingMunicipality>
,
<collectingCounty id="62BE4E1D5F7FFFB3FABAF872FF7DF877" pageId="3" pageNumber="504">Obhur creek</collectingCounty>
(Sharm Obhur),
<geoCoordinate id="EE5450565F7FFFB3FE74F829FDFDF874" box="[411,596,1984,2010]" degrees="21" direction="north" minutes="42" orientation="latitude" pageId="3" pageNumber="504" precision="1" seconds="31.00" value="21.708612">21°42’31.00” N</geoCoordinate>
,
<geoCoordinate id="EE5450565F7FFFB3FD8EF856FCA4F874" box="[609,781,1983,2010]" degrees="39" direction="east" minutes="05" orientation="longitude" pageId="3" pageNumber="504" precision="1" seconds="45.00" value="39.095833">39°05’45.00”E</geoCoordinate>
,
<elevation id="004DD1A25F7FFFB3FCF4F829FCF0F874" box="[795,857,1984,2010]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="504" unit="m" value="14.0">
<quantity id="4C989B745F7FFFB3FCF4F829FCF0F874" box="[795,857,1984,2010]" metricMagnitude="1" metricUnit="m" metricValue="1.4" pageId="3" pageNumber="504" unit="m" value="14.0">14 m</quantity>
</elevation>
, coll.
<collectorName id="269553475F7FFFB3FC4EF829FBE6F877" box="[929,1103,1983,2010]" pageId="3" pageNumber="504">T.J. Alpermann</collectorName>
&amp;
<collectorName id="269553475F7FFFB3FB97F829FA93F877" box="[1144,1338,1983,2010]" pageId="3" pageNumber="504">S.V. Bogorodsky</collectorName>
,
<collectingDate id="EF9AE9B95F7FFFB3FAA7F829FF7AF850" pageId="3" pageNumber="504" value="2013-07-01">
<date id="FFDE10515F7FFFB3FAA7F829FF7AF850" pageId="3" pageNumber="504" value="2013-07-01">01 July 2013</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37A651A5F78FFB4FF28FF71FE34FD8E" pageId="4" pageNumber="505" type="diagnosis">
<paragraph id="8BDF36915F78FFB4FF28FF71FE34FD8E" blockId="4.[151,1437,152,2017]" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FF28FF71FE93FF1D" bold="true" box="[199,314,152,179]" pageId="4" pageNumber="505">Diagnosis</emphasis>
. Dorsal-fin rays VI + I,10; anal-fin rays I,9; pectoral-fin rays 14, all rays branched; pelvic-fin rays I,5, the fin separated and without frenum, fifth ray unbranched; snout short, its length 0.7 of eye diameter; anterior nostril with a long nasal tube without process from the rim, posterior nostril pore with slightly erected rim; no tentacle above eye; posterior angle of jaws extending posteriorly to below posterior edge of pupil; no opercular spine; no mental frenum; pelvic fins ending at anus, longer than pectoral fin; squamation reduced to a few ctenoid scales on caudal peduncle at caudal-fin base; no head canals; suborbital rows of papillae with four transverse rows; three distinct preorbital rows
<emphasis id="B914EA835F78FFB4FE0BFE9DFE46FE23" box="[484,495,372,397]" italics="true" pageId="4" pageNumber="505">s</emphasis>
present from posterior nostril to above upper lip; suborbital row
<emphasis id="B914EA835F78FFB4FB3EFE9AFB76FE23" box="[1233,1247,371,397]" italics="true" pageId="4" pageNumber="505">3</emphasis>
short, extending from eye ending distantly from row
<emphasis id="B914EA835F78FFB4FDC1FE71FDECFE1C" box="[558,581,408,434]" italics="true" pageId="4" pageNumber="505">d;</emphasis>
suborbital row
<emphasis id="B914EA835F78FFB4FD19FE71FCADFE1C" box="[758,772,408,434]" italics="true" pageId="4" pageNumber="505">4</emphasis>
long, curving at ventroposterior edge of eye and extending obliquely below of level of row
<emphasis id="B914EA835F78FFB4FDEFFE55FDA7FE78" box="[512,526,444,470]" italics="true" pageId="4" pageNumber="505">d</emphasis>
; suborbital row
<emphasis id="B914EA835F78FFB4FD28FE55FD7CFE78" box="[711,725,444,470]" italics="true" pageId="4" pageNumber="505">b</emphasis>
anteriorly not reaching to below posterior edge of eye; anterior dorsal rows
<emphasis id="B914EA835F78FFB4FECCFE0BFE98FE55" box="[291,305,482,507]" italics="true" pageId="4" pageNumber="505">o</emphasis>
and
<emphasis id="B914EA835F78FFB4FE84FE0BFED6FE55" box="[363,383,482,507]" italics="true" pageId="4" pageNumber="505">m</emphasis>
present; head of alive and fresh fish orange, body yellowish orange with five faint, vertical, brown bands (
<figureCitation id="135B2A145F78FFB4FED9FDEFFE26FD8E" box="[310,399,518,544]" captionStart="FIGURE 1" captionStartId="6.[151,250,1898,1923]" captionTargetBox="[151,1436,251,1865]" captionTargetId="figure@6.[151,1436,269,1865]" captionTargetPageId="6" captionText="FIGURE 1. Hetereleotris aurantiaca sp. nov. SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). A: freshly collected specimen; B: preserved specimen; C: three peduncular scales marked with arrows. Photos by S.V. Bogorodsky (A), M. Kovačić (B, C)." httpUri="https://zenodo.org/record/3066725/files/figure.png" pageId="4" pageNumber="505">Fig. 1A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37A651A5F78FFB5FF28FDC3FB0CFAAD" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505" type="description">
<paragraph id="8BDF36915F78FFB4FF28FDC3FE52FB8D" blockId="4.[151,1437,152,2017]" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FF28FDC3FEF8FDEB" bold="true" box="[199,337,554,581]" pageId="4" pageNumber="505">Description</emphasis>
. Body moderately deep, its depth at pelvic-fin origin 5.1 in SL, at anal-fin origin 5.6 in SL, laterally compressed posteriorly, caudal-peduncle depth 0.7 of body depth at anal-fin origin, with caudal peduncle moderately deep, caudal peduncle depth 0.6 of caudal peduncle length (
<figureCitation id="135B2A145F78FFB4FC9FFD9DFC18FD21" box="[880,945,628,655]" captionStart="FIGURE 1" captionStartId="6.[151,250,1898,1923]" captionTargetBox="[151,1436,251,1865]" captionTargetId="figure@6.[151,1436,269,1865]" captionTargetPageId="6" captionText="FIGURE 1. Hetereleotris aurantiaca sp. nov. SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). A: freshly collected specimen; B: preserved specimen; C: three peduncular scales marked with arrows. Photos by S.V. Bogorodsky (A), M. Kovačić (B, C)." httpUri="https://zenodo.org/record/3066725/files/figure.png" pageId="4" pageNumber="505">Fig. 1</figureCitation>
). Head large, its length 3.1 in SL, width 4.1 in SL, depth 5.1 in SL, and depressed, its depth 0.8 of width. Snout steep in profile from lateral view, and short, its length 0.7 of eye diameter, 5.0 in head length. Anterior nostril with a long nasal tube without process from the rim, posterior nostril pore with erected rim, height of rim less than 1/10 of anterior nasal tube length. Eyes dorsolateral, moderately large, eye diameter 3.7 in head length, orbit projecting above dorsal profile. Interorbital narrow, 6.0 in eye diameter. No tentacle above eye. Mouth terminal, oblique, jaws ending about equal anteriorly. Mouth moderately large, posterior angle of jaws extending to below posterior edge of pupil. Cheek narrow. Upper jaw with single row of recurved pointed teeth. Lower jaw with a single row of recurved pointed teeth anteriorly of similar size as in upper jaw, followed laterally by smaller, pointed, and irregularly spaced teeth among which one or two enlarged recurved canines are present. Tongue short, with rounded tip. No mental frenum. Branchiostegal membranes fused to isthmus, gill openings restricted to pectoral-fin base. Lower limb of first gill arch joined to gill cover by membrane. No spines on preopercle.
</paragraph>
<paragraph id="8BDF36915F78FFB4FF28FBC6FAC8FA3D" blockId="4.[151,1437,152,2017]" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FF28FBC6FF50FBE6" box="[199,249,1071,1096]" italics="true" pageId="4" pageNumber="505">Fins</emphasis>
. First dorsal fin VI, second dorsal fin I,10; anal fin I,9; branched caudal-fin rays 14, segmented 17. Pectoral-fin rays 14, all rays branched, upper rays not free at tips. Pectoral girdle without flaps on anterior edge. Pelvic fins I,5 + 5,I, left and right fin completely separated and without frenum, fifth ray unbranched, other rays branched, fourth ray the longest. Spines of first dorsal fin not elongate or filamentous, second to fourth spine of first dorsal fin longest, third and fourth only slightly longer than the second spine, sixth spine short; spines of first dorsal fin not reaching to origin of second dorsal fin when folded down. Origin of first dorsal fin behind vertical at pectoral-fin base. Fin membrane from last spine of first dorsal fin connected with second dorsal-spine base, no gap present. Origin of anal fin below vertical of first segmented ray of second dorsal fin. Pectoral fins not extending posteriorly to below origin of second dorsal fin. Pelvic fins ending at anus, about equal to distance between origin of fins and anus, longer than pectoral fins, 1.1 of pectoral-fin length. Caudal fin rounded, shorter than head, 1.4 in head length.
</paragraph>
<paragraph id="8BDF36915F78FFB4FF28FA77FF61F9AC" blockId="4.[151,1437,152,2017]" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FF28FA77FEE4FA16" box="[199,333,1438,1464]" italics="true" pageId="4" pageNumber="505">Squamation</emphasis>
. Squamation reduced to three ctenoid scales on caudal peduncle at caudal-fin base: one scale at midline, two scales on upper and on the lower lateral side slightly more backwards, forming midlateral triangle (
<figureCitation id="135B2A145F78FFB4FA80FA2AFF13F9AC" captionStart="FIGURE 1" captionStartId="6.[151,250,1898,1923]" captionTargetBox="[151,1436,251,1865]" captionTargetId="figure@6.[151,1436,269,1865]" captionTargetPageId="6" captionText="FIGURE 1. Hetereleotris aurantiaca sp. nov. SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). A: freshly collected specimen; B: preserved specimen; C: three peduncular scales marked with arrows. Photos by S.V. Bogorodsky (A), M. Kovačić (B, C)." httpUri="https://zenodo.org/record/3066725/files/figure.png" pageId="4" pageNumber="505">Fig. 1C</figureCitation>
).
</paragraph>
<paragraph id="8BDF36915F78FFB5FF28F9E5FB60FCAE" blockId="4.[151,1437,152,2017]" lastBlockId="5.[151,1437,152,2019]" lastPageId="5" lastPageNumber="506" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FF28F9E5FE4AF988" box="[199,483,1548,1574]" italics="true" pageId="4" pageNumber="505">Cephalic sensory systems</emphasis>
(
<figureCitation id="135B2A145F78FFB4FE1EF9E5FD9CF989" box="[497,565,1548,1575]" captionStart="FIGURE 2" captionStartId="7.[151,250,1923,1948]" captionTargetBox="[202,1436,1116,1855]" captionTargetId="figure@7.[151,1436,1047,1888]" captionTargetPageId="7" captionText="FIGURE 2. Hetereleotris aurantiaca sp. nov. Cephalic sensory papillae: SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/3066727/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
). No head canals. Rows of head sensory papillae were counted on the both side and presented (left, right). Only left side papillae presented, two papillae on the place of the missing preopercular canal visible only on the right side were transferred on the drawing on the left side (marked
<emphasis id="B914EA835F78FFB4FB4FF9BEFB13F9DE" box="[1184,1210,1623,1648]" italics="true" pageId="4" pageNumber="505">pc</emphasis>
on
<figureCitation id="135B2A145F78FFB4FB0AF9BFFA80F9DE" box="[1253,1321,1622,1648]" captionStart="FIGURE 2" captionStartId="7.[151,250,1923,1948]" captionTargetBox="[202,1436,1116,1855]" captionTargetId="figure@7.[151,1436,1047,1888]" captionTargetPageId="7" captionText="FIGURE 2. Hetereleotris aurantiaca sp. nov. Cephalic sensory papillae: SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/3066727/files/figure.png" pageId="4" pageNumber="505">Fig. 2</figureCitation>
). Rows of head sensory papillae reduced and additional individual larger papillae present on the position of absent head canals.
<emphasis id="B914EA835F78FFB4FF78F949FEEFF914" box="[151,326,1696,1722]" italics="true" pageId="4" pageNumber="505">Preorbital rows</emphasis>
: snout with four median preorbital series, upper row
<emphasis id="B914EA835F78FFB4FC7BF948FC36F914" box="[916,927,1697,1722]" italics="true" pageId="4" pageNumber="505">r</emphasis>
(2, 3) middorsally between eyes above posterior nostrils,
<emphasis id="B914EA835F78FFB4FEC8F92CFE9BF970" box="[295,306,1733,1758]" italics="true" pageId="4" pageNumber="505">s</emphasis>
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<emphasis id="B914EA835F78FFB4FEDDF92AFE93F97F" box="[306,314,1731,1745]" italics="true" pageId="4" pageNumber="505">1</emphasis>
</superScript>
(1, 1) at posterior nostril, row
<emphasis id="B914EA835F78FFB4FD7BF92CFD36F970" box="[660,671,1733,1758]" italics="true" pageId="4" pageNumber="505">s</emphasis>
<superScript id="7C159BD95F78FFB4FD71F92AFD0FF97F" attach="left" box="[670,678,1731,1745]" fontSize="6" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FD71F92AFD0FF97F" box="[670,678,1731,1745]" italics="true" pageId="4" pageNumber="505">2</emphasis>
</superScript>
(1, 1) below
<emphasis id="B914EA835F78FFB4FCD5F92CFCECF970" box="[826,837,1733,1758]" italics="true" pageId="4" pageNumber="505">s</emphasis>
<superScript id="7C159BD95F78FFB4FCAAF92AFCE4F97F" attach="left" box="[837,845,1731,1745]" fontSize="6" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FCAAF92AFCE4F97F" box="[837,845,1731,1745]" italics="true" pageId="4" pageNumber="505">1</emphasis>
</superScript>
and posterior nostril, and vertical row
<emphasis id="B914EA835F78FFB4FB11F92CFAA0F970" box="[1278,1289,1733,1758]" italics="true" pageId="4" pageNumber="505">s</emphasis>
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<emphasis id="B914EA835F78FFB4FAE6F92AFAB8F97F" box="[1289,1297,1731,1745]" italics="true" pageId="4" pageNumber="505">3</emphasis>
</superScript>
(2, 2) above upper lip. Lateral series
<emphasis id="B914EA835F78FFB4FE4AF903FE18F8AD" box="[421,433,1770,1795]" italics="true" pageId="4" pageNumber="505">c</emphasis>
in four parts: superior
<emphasis id="B914EA835F78FFB4FD5EF903FD14F8AD" box="[689,701,1770,1795]" italics="true" pageId="4" pageNumber="505">c</emphasis>
<superScript id="7C159BD95F78FFB4FD52F901FD6CF958" attach="left" box="[701,709,1768,1782]" fontSize="6" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FD52F901FD6CF958" box="[701,709,1768,1782]" italics="true" pageId="4" pageNumber="505">2</emphasis>
</superScript>
(1, 2) between posterior nostril and anterior nasal tube, middle
<emphasis id="B914EA835F78FFB4FA67F903FA3DF8AD" box="[1416,1428,1770,1795]" italics="true" pageId="4" pageNumber="505">c</emphasis>
<superScript id="7C159BD95F78FFB4FA7BF901FA35F958" attach="left" box="[1428,1436,1768,1782]" fontSize="6" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FA7BF901FA35F958" box="[1428,1436,1768,1782]" italics="true" pageId="4" pageNumber="505">1</emphasis>
</superScript>
(3, 3) just below base of anterior nasal tube, inferior rows: upper horizontal
<emphasis id="B914EA835F78FFB4FC0FF8E6FC45F886" box="[992,1004,1807,1832]" italics="true" pageId="4" pageNumber="505">c</emphasis>
<subScript id="17E434D45F78FFB4FC02F8C9FC5CF880" attach="left" box="[1005,1013,1824,1838]" fontSize="6" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FC02F8C9FC5CF880" box="[1005,1013,1824,1838]" italics="true" pageId="4" pageNumber="505">2</emphasis>
</subScript>
(3, 3) above upper lip and lower horizontal
<emphasis id="B914EA835F78FFB4FF0DF8DDFF47F8E3" box="[226,238,1844,1869]" italics="true" pageId="4" pageNumber="505">c</emphasis>
<subScript id="17E434D45F78FFB4FF00F8ADFF5EF8FC" attach="left" box="[239,247,1860,1874]" fontSize="6" pageId="4" pageNumber="505">
<emphasis id="B914EA835F78FFB4FF00F8ADFF5EF8FC" box="[239,247,1860,1874]" italics="true" pageId="4" pageNumber="505">1</emphasis>
</subScript>
(1, 1) between upper lip and row
<emphasis id="B914EA835F78FFB4FD9DF8DAFD29F8E3" box="[626,640,1843,1869]" italics="true" pageId="4" pageNumber="505">1</emphasis>
.
<emphasis id="B914EA835F78FFB4FD61F8DAFCE9F8E3" box="[654,832,1843,1869]" italics="true" pageId="4" pageNumber="505">Suborbital rows</emphasis>
: no row
<emphasis id="B914EA835F78FFB4FC4BF8DDFC1BF8E3" box="[932,946,1844,1869]" italics="true" pageId="4" pageNumber="505">a</emphasis>
. Row
<emphasis id="B914EA835F78FFB4FC13F8DAFBA3F8E3" box="[1020,1034,1843,1869]" italics="true" pageId="4" pageNumber="505">b</emphasis>
(3, 2) longitudinal, short, anteriorly beginning close to transverse row
<emphasis id="B914EA835F78FFB4FDFAF8B1FD8AF8DC" box="[533,547,1880,1906]" italics="true" pageId="4" pageNumber="505">4</emphasis>
and slightly behind vertical of posterior edge of eye. Four transverse suborbital rows (
<emphasis id="B914EA835F78FFB4FF32F894FEAAF839" box="[221,259,1917,1943]" italics="true" pageId="4" pageNumber="505">1-4</emphasis>
) of sensory papillae, row
<emphasis id="B914EA835F78FFB4FDCDF894FD99F839" box="[546,560,1917,1943]" italics="true" pageId="4" pageNumber="505">1</emphasis>
longer, below anterior edge of pupil reaching upper lip, row
<emphasis id="B914EA835F78FFB4FB3EF894FB76F839" box="[1233,1247,1917,1943]" italics="true" pageId="4" pageNumber="505">2</emphasis>
short and distant from eye, ending ventrally close to row
<emphasis id="B914EA835F78FFB4FDBCF848FDC8F815" box="[595,609,1953,1979]" italics="true" pageId="4" pageNumber="505">d</emphasis>
, row
<emphasis id="B914EA835F78FFB4FD4FF848FD07F815" box="[672,686,1953,1979]" italics="true" pageId="4" pageNumber="505">3</emphasis>
short, extending from eye ending distantly from row
<emphasis id="B914EA835F78FFB4FAEDF848FAB9F815" box="[1282,1296,1953,1979]" italics="true" pageId="4" pageNumber="505">d</emphasis>
, row
<emphasis id="B914EA835F78FFB4FAA0F848FAF4F815" box="[1359,1373,1953,1979]" italics="true" pageId="4" pageNumber="505">4</emphasis>
long, curving at ventroposterior edge of eye and then extending obliquely below of level of row
<emphasis id="B914EA835F78FFB4FB66F82FFB3EF84E" box="[1161,1175,1990,2016]" italics="true" pageId="4" pageNumber="505">d</emphasis>
(
<emphasis id="B914EA835F78FFB4FB48F82FFB1CF84E" box="[1191,1205,1990,2016]" italics="true" pageId="4" pageNumber="505">1</emphasis>
: 5, 5,
<emphasis id="B914EA835F78FFB4FB14F82FFAA0F84E" box="[1275,1289,1990,2016]" italics="true" pageId="4" pageNumber="505">2</emphasis>
: 3, 3,
<emphasis id="B914EA835F78FFB4FAA0F82FFAF4F84E" box="[1359,1373,1990,2016]" italics="true" pageId="4" pageNumber="505">3</emphasis>
: 4, 4,
<emphasis id="B914EA835F79FFB5FF78FF70FF0CFF1D" box="[151,165,153,179]" italics="true" pageId="5" pageNumber="506">4</emphasis>
: 14, 15). Row
<emphasis id="B914EA835F79FFB5FEA1FF70FEF5FF1D" box="[334,348,153,179]" italics="true" pageId="5" pageNumber="506">d</emphasis>
(11, 11) continuous above upper lip and on cheek, ending backwards below posterior part of eye and near transverse suborbital row
<emphasis id="B914EA835F79FFB5FDF0FF57FD84FF76" box="[543,557,190,216]" italics="true" pageId="5" pageNumber="506">4</emphasis>
.
<emphasis id="B914EA835F79FFB5FDD4FF57FC24FF76" box="[571,909,190,216]" italics="true" pageId="5" pageNumber="506">Preoperculo-mandibular rows</emphasis>
: external row
<emphasis id="B914EA835F79FFB5FBDFFF56FB95FF76" box="[1072,1084,191,216]" italics="true" pageId="5" pageNumber="506">e</emphasis>
(10 + 10, 10 + 11) longitudinal and uniserial, divided into anterior and posterior sections; internal row
<emphasis id="B914EA835F79FFB5FC48FF0AFC06FF52" box="[935,943,227,252]" italics="true" pageId="5" pageNumber="506">i</emphasis>
uniserial, divided into anterior and posterior sections (8 + 6, 8 + 6); mental row
<emphasis id="B914EA835F79FFB5FDDFFEEEFD91FE8F" box="[560,568,263,289]" italics="true" pageId="5" pageNumber="506">f</emphasis>
longitudinal (4, 5).
<emphasis id="B914EA835F79FFB5FCF0FEEEFBAAFE8F" box="[799,1027,263,289]" italics="true" pageId="5" pageNumber="506">Oculoscapular rows</emphasis>
: three larger papillae longitudinally arranged on position of missing anterior oculoscapular canal behind eye; two larger papillae longitudinally arranged on position of missing posterior oculoscapular canal above opercle (marked
<emphasis id="B914EA835F79FFB5FC14FEBBFB8AFEC5" box="[1019,1059,338,363]" italics="true" pageId="5" pageNumber="506">aoc</emphasis>
and
<emphasis id="B914EA835F79FFB5FBB2FEBBFB2CFEC5" box="[1117,1157,338,363]" italics="true" pageId="5" pageNumber="506">poc</emphasis>
on
<figureCitation id="135B2A145F79FFB5FB5DFEB8FB5EFEC5" box="[1202,1271,337,363]" captionStart="FIGURE 2" captionStartId="7.[151,250,1923,1948]" captionTargetBox="[202,1436,1116,1855]" captionTargetId="figure@7.[151,1436,1047,1888]" captionTargetPageId="7" captionText="FIGURE 2. Hetereleotris aurantiaca sp. nov. Cephalic sensory papillae: SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/3066727/files/figure.png" pageId="5" pageNumber="506">Fig. 2</figureCitation>
, respectively); longitudinal row
<date id="FFDE10515F79FFB5FEB8FE9EFEC2FE2D" box="[343,363,373,400]" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FEB8FE9EFECAFE3E" box="[343,355,375,400]" italics="true" pageId="5" pageNumber="506">x</emphasis>
<superScript id="7C159BD95F79FFB5FE8CFE9CFEC2FE2D" attach="left" box="[355,363,373,387]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FE8CFE9CFEC2FE2D" box="[355,363,373,387]" italics="true" pageId="5" pageNumber="506">1</emphasis>
</superScript>
</date>
(4, 6) short; posterior longitudinal row
<date id="FFDE10515F79FFB5FCCBFE9EFC91FE2D" box="[804,824,373,400]" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FCCBFE9EFC99FE3E" box="[804,816,375,400]" italics="true" pageId="5" pageNumber="506">x</emphasis>
<superScript id="7C159BD95F79FFB5FCDFFE9CFC91FE2D" attach="left" box="[816,824,373,387]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FCDFFE9CFC91FE2D" box="[816,824,373,387]" italics="true" pageId="5" pageNumber="506">2</emphasis>
</superScript>
</date>
(3, 4) above two papillae on position of missing posterior oculoscapular canal; anterior lower transverse row
<emphasis id="B914EA835F79FFB5FCEFFE72FCA2FE1A" box="[768,779,411,436]" italics="true" pageId="5" pageNumber="506">z</emphasis>
(5, 5) between row
<date id="FFDE10515F79FFB5FC05FE72FC56FE09" box="[1002,1023,409,436]" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FC05FE72FC5FFE1A" box="[1002,1014,411,436]" italics="true" pageId="5" pageNumber="506">x</emphasis>
<superScript id="7C159BD95F79FFB5FC18FE70FC56FE09" attach="left" box="[1015,1023,409,423]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FC18FE70FC56FE09" box="[1015,1023,409,423]" italics="true" pageId="5" pageNumber="506">1</emphasis>
</superScript>
</date>
and opercular transverse row
<emphasis id="B914EA835F79FFB5FAA2FE72FACAFE1A" box="[1357,1379,411,436]" italics="true" pageId="5" pageNumber="506">ot</emphasis>
; row
<emphasis id="B914EA835F79FFB5FF78FE29FF0CFE77" box="[151,165,448,473]" italics="true" pageId="5" pageNumber="506">q</emphasis>
(1, 1) below and behind posterior end of row
<date id="FFDE10515F79FFB5FD4DFE29FD1FFE62" box="[674,694,446,473]" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FD4DFE29FD07FE77" box="[674,686,448,473]" italics="true" pageId="5" pageNumber="506">x</emphasis>
<superScript id="7C159BD95F79FFB5FD41FE57FD1FFE62" attach="left" box="[686,694,446,460]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FD41FE57FD1FFE62" box="[686,694,446,460]" italics="true" pageId="5" pageNumber="506">1</emphasis>
</superScript>
</date>
; row
<emphasis id="B914EA835F79FFB5FD19FE29FCBEFE77" box="[758,791,448,473]" italics="true" pageId="5" pageNumber="506">trp</emphasis>
(3, 1) between row
<emphasis id="B914EA835F79FFB5FC1AFE29FBAAFE77" box="[1013,1027,448,473]" italics="true" pageId="5" pageNumber="506">q</emphasis>
and papillae on the position of missing posterior oculoscapular canal; row
<emphasis id="B914EA835F79FFB5FDA5FE0CFDFFFE50" box="[586,598,485,510]" italics="true" pageId="5" pageNumber="506">y</emphasis>
absent. Axillary transverse rows
<emphasis id="B914EA835F79FFB5FC24FE0CFC4DFE50" box="[971,996,485,510]" italics="true" pageId="5" pageNumber="506">as</emphasis>
<superScript id="7C159BD95F79FFB5FC0BFE0AFC45FE5F" attach="left" box="[996,1004,483,497]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FC0BFE0AFC45FE5F" box="[996,1004,483,497]" italics="true" pageId="5" pageNumber="506">1</emphasis>
</superScript>
(4, 3),
<emphasis id="B914EA835F79FFB5FBD5FE0CFBFAFE50" box="[1082,1107,485,510]" italics="true" pageId="5" pageNumber="506">as</emphasis>
<superScript id="7C159BD95F79FFB5FBBCFE0AFBF2FE5F" attach="left" box="[1107,1115,483,497]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FBBCFE0AFBF2FE5F" box="[1107,1115,483,497]" italics="true" pageId="5" pageNumber="506">2</emphasis>
</superScript>
(2, 3),
<emphasis id="B914EA835F79FFB5FB46FE0CFB6BFE50" box="[1193,1218,485,510]" italics="true" pageId="5" pageNumber="506">as</emphasis>
<superScript id="7C159BD95F79FFB5FB2DFE0AFB63FE5F" attach="left" box="[1218,1226,483,497]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FB2DFE0AFB63FE5F" box="[1218,1226,483,497]" italics="true" pageId="5" pageNumber="506">3</emphasis>
</superScript>
(3, 0) present; row
<emphasis id="B914EA835F79FFB5FF78FDE0FF04FD8D" box="[151,173,521,547]" italics="true" pageId="5" pageNumber="506">la</emphasis>
<superScript id="7C159BD95F79FFB5FF42FDE1FF1CFDB8" attach="left" box="[173,181,520,534]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FF42FDE1FF1CFDB8" box="[173,181,520,534]" italics="true" pageId="5" pageNumber="506">1</emphasis>
</superScript>
and row
<emphasis id="B914EA835F79FFB5FEF4FDE0FE98FD8D" box="[283,305,521,547]" italics="true" pageId="5" pageNumber="506">la</emphasis>
<superScript id="7C159BD95F79FFB5FEDEFDE1FE90FDB8" attach="left" box="[305,313,520,534]" fontSize="6" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FEDEFDE1FE90FDB8" box="[305,313,520,534]" italics="true" pageId="5" pageNumber="506">2</emphasis>
</superScript>
absent.
<emphasis id="B914EA835F79FFB5FE7BFDE0FDEBFD8D" box="[404,578,521,547]" italics="true" pageId="5" pageNumber="506">Opercular rows</emphasis>
: two larger papillae in the position of missing preopercular canal visible only on the right side; transverse row
<emphasis id="B914EA835F79FFB5FDE6FDC6FDB6FDE6" box="[521,543,559,584]" italics="true" pageId="5" pageNumber="506">ot</emphasis>
(10, 9); superior longitudinal row
<emphasis id="B914EA835F79FFB5FC49FDC6FC16FDE6" box="[934,959,559,584]" italics="true" pageId="5" pageNumber="506">os</emphasis>
(4, 4); and inferior longitudinal row
<emphasis id="B914EA835F79FFB5FA8EFDC6FADEFDE6" box="[1377,1399,559,584]" italics="true" pageId="5" pageNumber="506">oi</emphasis>
(3, 4).
<emphasis id="B914EA835F79FFB5FF52FDBAFE01FDC2" box="[189,424,595,621]" italics="true" pageId="5" pageNumber="506">Anterior dorsal rows</emphasis>
: single papilla behind eye on position of the missing pore
<emphasis id="B914EA835F79FFB5FBD1FDBAFBFBFDC3" bold="true" box="[1086,1106,595,621]" italics="true" pageId="5" pageNumber="506">ω</emphasis>
of the anterior oculoscapular canal (marked
<emphasis id="B914EA835F79FFB5FED2FD91FECCFD3F" box="[317,357,632,657]" italics="true" pageId="5" pageNumber="506">aoc</emphasis>
on
<figureCitation id="135B2A145F79FFB5FE60FD9EFE7BFD3F" box="[399,466,631,657]" captionStart="FIGURE 2" captionStartId="7.[151,250,1923,1948]" captionTargetBox="[202,1436,1116,1855]" captionTargetId="figure@7.[151,1436,1047,1888]" captionTargetPageId="7" captionText="FIGURE 2. Hetereleotris aurantiaca sp. nov. Cephalic sensory papillae: SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/3066727/files/figure.png" pageId="5" pageNumber="506">Fig. 2</figureCitation>
), transverse row
<emphasis id="B914EA835F79FFB5FD7DFD91FD09FD3F" box="[658,672,632,657]" italics="true" pageId="5" pageNumber="506">n</emphasis>
(4, 6) behind it, row
<emphasis id="B914EA835F79FFB5FC63FD91FC33FD3F" box="[908,922,632,657]" italics="true" pageId="5" pageNumber="506">o</emphasis>
(1, 1) slightly in front and middorsally to row
<emphasis id="B914EA835F79FFB5FF78FD74FF0CFD18" box="[151,165,669,694]" italics="true" pageId="5" pageNumber="506">g</emphasis>
, row
<emphasis id="B914EA835F79FFB5FF0AFD74FF5AFD18" box="[229,243,669,694]" italics="true" pageId="5" pageNumber="506">g</emphasis>
(4, 4) longitudinal; row
<emphasis id="B914EA835F79FFB5FDEBFD74FDB1FD18" box="[516,536,669,694]" italics="true" pageId="5" pageNumber="506">m</emphasis>
(1, 1) in the level of row
<emphasis id="B914EA835F79FFB5FCDCFD74FCE8FD18" box="[819,833,669,694]" italics="true" pageId="5" pageNumber="506">g</emphasis>
behind it, row
<emphasis id="B914EA835F79FFB5FC05FD75FC51FD18" box="[1002,1016,668,694]" italics="true" pageId="5" pageNumber="506">h</emphasis>
longitudinal (6, 6) in front of the first dorsal fin.
<emphasis id="B914EA835F79FFB5FEE0FD28FE64FD75" box="[271,461,705,731]" italics="true" pageId="5" pageNumber="506">Interorbital rows</emphasis>
: two pairs of larger papillae present in interorbital, one anteriorly on the position of the missing pore
<emphasis id="B914EA835F79FFB5FEC1FD0CFE93FCAE" bold="true" box="[302,314,741,768]" italics="true" pageId="5" pageNumber="506">λ</emphasis>
and one posteriorly on the position of the missing pore
<emphasis id="B914EA835F79FFB5FC43FD0FFC10FCAE" bold="true" box="[940,953,742,768]" italics="true" pageId="5" pageNumber="506">κ</emphasis>
(marked
<emphasis id="B914EA835F79FFB5FBCAFD0EFBE4FCAE" box="[1061,1101,743,768]" italics="true" pageId="5" pageNumber="506">aoc</emphasis>
on
<figureCitation id="135B2A145F79FFB5FB98FD0FFB15FCAE" box="[1143,1212,742,768]" captionStart="FIGURE 2" captionStartId="7.[151,250,1923,1948]" captionTargetBox="[202,1436,1116,1855]" captionTargetId="figure@7.[151,1436,1047,1888]" captionTargetPageId="7" captionText="FIGURE 2. Hetereleotris aurantiaca sp. nov. Cephalic sensory papillae: SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). Terminology in text. Drawing by M. Kovačić." httpUri="https://zenodo.org/record/3066727/files/figure.png" pageId="5" pageNumber="506">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BDF36915F79FFB5FF28FCE2FD33FB3B" blockId="5.[151,1437,152,2019]" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FF28FCE2FD3AFC8B" box="[199,659,779,805]" italics="true" pageId="5" pageNumber="506">Color of female immediately after death</emphasis>
(
<figureCitation id="135B2A145F79FFB5FD4BFCE3FD56FC8B" box="[676,767,778,805]" captionStart="FIGURE 1" captionStartId="6.[151,250,1898,1923]" captionTargetBox="[151,1436,251,1865]" captionTargetId="figure@6.[151,1436,269,1865]" captionTargetPageId="6" captionText="FIGURE 1. Hetereleotris aurantiaca sp. nov. SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). A: freshly collected specimen; B: preserved specimen; C: three peduncular scales marked with arrows. Photos by S.V. Bogorodsky (A), M. Kovačić (B, C)." httpUri="https://zenodo.org/record/3066725/files/figure.png" pageId="5" pageNumber="506">Fig. 1A</figureCitation>
). Head orange with scattered small yellowish grey to pale brown markings; eyes with brown-orange iris and black pupil; anterior nasal tube dark brown. Body yellowish orange, with five faint, vertical, brown bands, extending from dorsal to ventral body side, first two bands below first dorsal fin, remaining three bands below second dorsal fin, first band below second to fifth spines of first dorsal fin, is the most intensive. All bands dotted with brown. Small irregular brown-orange spot at base of second to fourth and of fifth to seventh ray of second dorsal fin. Caudal-fin base with narrow, diffuse, orange-brown bar. All fins similar in color, membranes of median fins with many tiny melanophores; first and second spine of the first dorsal fin and spine of second dorsal fin with alternating white and orange bands, third to sixth spines orange, dark brown streak basally on second membrane of first dorsal fin; rays of second dorsal, anal, and caudal fins orange with small white spots; pectoral fins with semi-translucent membranes and irregular dark orange bar at base, rays orange with white spots; pelvic fins pale brownish orange.
</paragraph>
<paragraph id="8BDF36915F79FFB5FF28FB76FB0CFAAD" blockId="5.[151,1437,152,2019]" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FF28FB76FE47FB17" box="[199,494,1183,1209]" italics="true" pageId="5" pageNumber="506">Color of preserved female</emphasis>
(
<figureCitation id="135B2A145F79FFB5FE12FB76FDFCFB17" box="[509,597,1183,1209]" captionStart="FIGURE 1" captionStartId="6.[151,250,1898,1923]" captionTargetBox="[151,1436,251,1865]" captionTargetId="figure@6.[151,1436,269,1865]" captionTargetPageId="6" captionText="FIGURE 1. Hetereleotris aurantiaca sp. nov. SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur). A: freshly collected specimen; B: preserved specimen; C: three peduncular scales marked with arrows. Photos by S.V. Bogorodsky (A), M. Kovačić (B, C)." httpUri="https://zenodo.org/record/3066725/files/figure.png" pageId="5" pageNumber="506">Fig. 1B</figureCitation>
). Body opaque white, lost orange and brownish coloration, no visible pigments on head and body. Eyes dark, including pupil. Dorsal and anal fins transparent, the rare dark pigmented dots visible on dorsal fins. Caudal, pectoral and pelvic fins transparent without any pigmentation.
</paragraph>
</subSubSection>
<subSubSection id="C37A651A5F79FFB5FF28FAE7FF7BFA4E" pageId="5" pageNumber="506" type="etymology">
<paragraph id="8BDF36915F79FFB5FF28FAE7FDACFAE2" blockId="5.[151,1437,152,2019]" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FF28FAE7FEE1FA86" bold="true" box="[199,328,1294,1320]" pageId="5" pageNumber="506">Etymology</emphasis>
. The specific epithet is from the Latin
<taxonomicName id="4C604D125F79FFB5FCE8FAE6FC2BFA86" authorityName="Kovačić &amp; Bogorodsky &amp; Mal" authorityYear="2019" box="[775,898,1295,1320]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="aurantiaca">
<emphasis id="B914EA835F79FFB5FCE8FAE6FC2BFA86" box="[775,898,1295,1320]" italics="true" pageId="5" pageNumber="506">aurantiaca</emphasis>
</taxonomicName>
, orange, and refers to the prevalence of orange color in head, body and fin-rays.
</paragraph>
<paragraph id="8BDF36915F79FFB5FF28FABFFF7BFA4E" blockId="5.[151,1437,152,2019]" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FF28FABFFE8AFADF" bold="true" box="[199,291,1366,1393]" pageId="5" pageNumber="506">Habitat</emphasis>
. Known only on the basis of the
<typeStatus id="54DB88335F79FFB5FD77FABEFD53FADF" box="[664,762,1367,1393]" pageId="5" pageNumber="506" type="holotype">holotype</typeStatus>
collected at
<quantity id="4C989B745F79FFB5FC66FABEFC45FADC" box="[905,1004,1367,1394]" metricMagnitude="1" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="5" pageNumber="506" unit="m" value="15.0" valueMax="16.0" valueMin="14.0">14–16 m</quantity>
depth, at Sharm Obhur, Jeddah,
<collectingCountry id="F37776015F79FFB5FAB3FABEFF4EFA38" name="Saudi Arabia" pageId="5" pageNumber="506">Saudi Arabia</collectingCountry>
. Habitat of the new species is steep coral-reef sloping down to about
<quantity id="4C989B745F79FFB5FBE8FA95FBEBFA39" box="[1031,1090,1404,1431]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="506" unit="m" value="30.0">30 m</quantity>
depth, upper half of the slope characterized by short steps covered with hard and soft corals, the specimen was collected at the entrance of small cave.
</paragraph>
</subSubSection>
<subSubSection id="C37A651A5F79FFB7FF28FA02FC51FC41" lastPageId="7" lastPageNumber="508" pageId="5" pageNumber="506" type="discussion">
<paragraph id="8BDF36915F79FFB6FF28FA02FC69FF55" blockId="5.[151,1437,152,2019]" lastBlockId="6.[151,1436,153,251]" lastPageId="6" lastPageNumber="507" pageId="5" pageNumber="506">
<emphasis id="B914EA835F79FFB5FF28FA02FE9DF9AB" bold="true" box="[199,308,1515,1541]" pageId="5" pageNumber="506">Remarks</emphasis>
. The new species is unique among the species of the genus
<taxonomicName id="4C604D125F79FFB5FC08FA02FBD3F9AB" authorityName="Bleeker" authorityYear="1874" box="[999,1146,1515,1541]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="506" phylum="Chordata" rank="genus">
<emphasis id="B914EA835F79FFB5FC08FA02FBD3F9AB" box="[999,1146,1515,1541]" italics="true" pageId="5" pageNumber="506">Hetereleotris</emphasis>
</taxonomicName>
, except for
<taxonomicName id="4C604D125F79FFB5FAEBFA02FAC2F9AA" baseAuthorityName="Hoese &amp; Winterbottom" baseAuthorityYear="1979" box="[1284,1387,1515,1540]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="apora">
<emphasis id="B914EA835F79FFB5FAEBFA02FAC2F9AA" box="[1284,1387,1515,1540]" italics="true" pageId="5" pageNumber="506">H. apora</emphasis>
</taxonomicName>
and
<taxonomicName id="4C604D125F79FFB5FF78F9F9FF5DF987" authorityName="Shibukawa" authorityYear="2010" box="[151,244,1551,1577]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="exilis">
<emphasis id="B914EA835F79FFB5FF78F9F9FF5DF987" box="[151,244,1551,1577]" italics="true" pageId="5" pageNumber="506">H. exilis</emphasis>
</taxonomicName>
, in the combination of absence of head canals and squamation reduced to caudal peduncle. From
<taxonomicName id="4C604D125F79FFB5FAD8F9F9FA35F987" baseAuthorityName="Hoese &amp; Winterbottom" baseAuthorityYear="1979" box="[1335,1436,1552,1577]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="apora">
<emphasis id="B914EA835F79FFB5FAD8F9F9FA35F987" box="[1335,1436,1552,1577]" italics="true" pageId="5" pageNumber="506">H. apora</emphasis>
</taxonomicName>
it differs by series of characters among which the most useful are: body with squamation reduced to a few scales on caudal peduncle at caudal-fin base (vs. 4–5 rows of scales restricted to caudal peduncle); anterior nostril with a long nasal tube without projection from rim, posterior nostril just pore with erected rim, height of rim less than 1/10 of anterior nasal tube length (vs. anterior and posterior nostril tubes, anterior nostril slightly longer than posterior nostril); operculum without spines (vs. operculum with 2 spines dorsally); dorsal-fin rays VI + I,10 and anal-fin rays I,9 (vs. dorsal-fin rays VI + I,11 and anal-fin rays I,10); pectoral-fin rays 14 (vs. pectoral-fin rays 15–16), pelvic fins I,5 + 5,I (vs. pelvic fins I,4 + 4,I); body yellowish orange, with five vertical brown bars (vs. body dusky grey with diffuse dark bars on upper half). From
<taxonomicName id="4C604D125F79FFB5FD6AF8DEFD4BF8FE" authorityName="Shibukawa" authorityYear="2010" box="[645,738,1846,1872]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="506" phylum="Chordata" rank="species" species="exilis">
<emphasis id="B914EA835F79FFB5FD6AF8DEFD4BF8FE" box="[645,738,1846,1872]" italics="true" pageId="5" pageNumber="506">H. exilis</emphasis>
</taxonomicName>
it differs by series of characters among which the most useful are: body with squamation reduced to a few ctenoid scales on caudal peduncle at caudal-fin base (vs. 14–15 rows of cycloid scales on caudal peduncle); posterior angle of jaws extending posteriorly to below posterior edge of pupil (vs. posterior angle of jaws extending posteriorly to below middle or anterior edge of pupil); anterior nostril with a long nasal tube without projection from rim, posterior nostril just pore with erected rim, height of rim less than 1/10 of anterior nasal tube length (vs. anterior and posterior nostril with nasal tubes); dorsal-fin rays VI + I,10 and anal-fin rays I,9 (vs. dorsal-fin rays VI + I,12 and anal-fin rays I,11); pectoral-fin rays 14 (vs. pectoral-fin rays 16); pelvic fins ending at anus (vs. pelvic fins not reaching posteriorly anus).
</paragraph>
<caption id="DF1F66195F7AFFB6FF78F883FD53F86D" httpUri="https://zenodo.org/record/3066725/files/figure.png" pageId="6" pageNumber="507" startId="6.[151,250,1898,1923]" targetBox="[151,1436,251,1865]" targetPageId="6">
<paragraph id="8BDF36915F7AFFB6FF78F883FD53F86D" blockId="6.[151,1437,1898,1987]" pageId="6" pageNumber="507">
<emphasis id="B914EA835F7AFFB6FF78F883FEBAF82D" bold="true" box="[151,275,1898,1923]" pageId="6" pageNumber="507">FIGURE 1.</emphasis>
<taxonomicName id="4C604D125F7AFFB6FEF5F883FDBDF82C" authority="Kovačić &amp; Bogorodsky &amp; Mal, 2019" authorityName="Kovačić &amp; Bogorodsky &amp; Mal" authorityYear="2019" box="[282,532,1898,1922]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="507" phylum="Chordata" rank="species" species="aurantiaca" status="sp. nov.">
<emphasis id="B914EA835F7AFFB6FEF5F883FDBDF82C" box="[282,532,1898,1922]" italics="true" pageId="6" pageNumber="507">Hetereleotris aurantiaca</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22757F85F7AFFB6FDF4F882FDC5F82D" box="[539,620,1899,1923]" pageId="6" pageNumber="507" rank="species">
<emphasis id="B914EA835F7AFFB6FDF4F882FDC5F82D" bold="true" box="[539,620,1899,1923]" pageId="6" pageNumber="507">sp. nov.</emphasis>
</taxonomicNameLabel>
SMF 35966, holotype, female, 14.51+3.40 mm, Red Sea, Saudi Arabia, Jeddah, Obhur creek (Sharm Obhur).
<emphasis id="B914EA835F7AFFB6FE27F863FE73F80D" bold="true" box="[456,474,1930,1955]" pageId="6" pageNumber="507">A</emphasis>
: freshly collected specimen;
<emphasis id="B914EA835F7AFFB6FCE5F862FCB2F80D" bold="true" box="[778,795,1931,1955]" pageId="6" pageNumber="507">B</emphasis>
: preserved specimen;
<emphasis id="B914EA835F7AFFB6FBECF863FBBCF80D" bold="true" box="[1027,1045,1930,1955]" pageId="6" pageNumber="507">C</emphasis>
: three peduncular scales marked with arrows. Photos by S.V. Bogorodsky (A), M. Kovačić (B, C).
</paragraph>
</caption>
<paragraph id="8BDF36915F7BFFB7FF28FF70FC41FD61" blockId="7.[151,1437,153,1007]" pageId="7" pageNumber="508">
The new species could be easily appeared as scaleless. The three species, among 19 currently known species of the genus, have the combination of absence of head canals and scales:
<taxonomicName id="4C604D125F7BFFB7FC40FF57FBEEFF79" baseAuthorityName="Ruppell" baseAuthorityYear="1830" box="[943,1095,189,215]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="diademata">
<emphasis id="B914EA835F7BFFB7FC40FF57FBEEFF79" box="[943,1095,189,215]" italics="true" pageId="7" pageNumber="508">H. diademata</emphasis>
</taxonomicName>
,
<taxonomicName id="4C604D125F7BFFB7FBBAFF57FB56FF79" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" box="[1109,1279,189,215]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="dorsovittata">
<emphasis id="B914EA835F7BFFB7FBBAFF57FB56FF79" box="[1109,1279,189,215]" italics="true" pageId="7" pageNumber="508">H. dorsovittata</emphasis>
</taxonomicName>
and
<emphasis id="B914EA835F7BFFB7FADAFF57FDBEFF55" italics="true" pageId="7" pageNumber="508">
<taxonomicName id="4C604D125F7BFFB7FADAFF57FF5FFF55" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="psammophila">H. psammophila</taxonomicName>
.
<taxonomicName id="4C604D125F7BFFB7FEEEFF08FDBEFF55" authorityName="Kovačić &amp; Bogorodsky &amp; Mal" authorityYear="2019" box="[257,535,225,251]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="aurantiaca">Hetereleotris aurantiaca</taxonomicName>
</emphasis>
differs from these three species, in addition to presence of peduncular scales, in having anterior nostril with a long nasal tube without projection from rim, posterior nostril just pore with erected rim, height of rim less than 1/10 of anterior nasal tube length vs. anterior and posterior nasal tubes, anterior nasal tube slightly longer than posterior nasal tube in
<taxonomicName id="4C604D125F7BFFB7FD47FEA7FC96FEC9" baseAuthorityName="Ruppell" baseAuthorityYear="1830" box="[680,831,333,359]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="diademata">
<emphasis id="B914EA835F7BFFB7FD47FEA7FC96FEC9" box="[680,831,333,359]" italics="true" pageId="7" pageNumber="508">H. diademata</emphasis>
</taxonomicName>
, anterior and posterior nasal tubes, posterior tube onehalf of anterior tube in
<taxonomicName id="4C604D125F7BFFB7FE70FE9BFDE0FE25" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" box="[415,585,369,395]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="dorsovittata">
<emphasis id="B914EA835F7BFFB7FE70FE9BFDE0FE25" box="[415,585,369,395]" italics="true" pageId="7" pageNumber="508">H. dorsovittata</emphasis>
</taxonomicName>
, anterior and posterior nasal tubes, posterior tube 1/3 of anterior tube in
<taxonomicName id="4C604D125F7BFFB7FA6EFE9BFE87FE01" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="psammophila">
<emphasis id="B914EA835F7BFFB7FA6EFE9BFE87FE01" italics="true" pageId="7" pageNumber="508">H. psammophila</emphasis>
</taxonomicName>
; larger mouth with posterior angle of jaws extending posteriorly to below posterior edge of pupil vs. posterior angle of jaws extending posteriorly to below anterior edge of orbit in
<taxonomicName id="4C604D125F7BFFB7FC13FE53FB3AFE7D" baseAuthorityName="Ruppell" baseAuthorityYear="1830" box="[1020,1171,441,467]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="diademata">
<emphasis id="B914EA835F7BFFB7FC13FE53FB3AFE7D" box="[1020,1171,441,467]" italics="true" pageId="7" pageNumber="508">H. diademata</emphasis>
</taxonomicName>
, posterior angle of jaws extending posteriorly to below anterior edge of pupil in
<taxonomicName id="4C604D125F7BFFB7FCE3FE37FC1CFE59" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" box="[780,949,477,503]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="dorsovittata">
<emphasis id="B914EA835F7BFFB7FCE3FE37FC1CFE59" box="[780,949,477,503]" italics="true" pageId="7" pageNumber="508">H. dorsovittata</emphasis>
</taxonomicName>
and
<taxonomicName id="4C604D125F7BFFB7FC03FE37FB0CFE59" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" box="[1004,1189,477,503]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="psammophila">
<emphasis id="B914EA835F7BFFB7FC03FE37FB0CFE59" box="[1004,1189,477,503]" italics="true" pageId="7" pageNumber="508">H. psammophila</emphasis>
</taxonomicName>
; alive coloration with head orange, body yellowish orange, with five vertical brown bands on body side vs. head and body whitish, mottled sandy brown and with transverse dark band below first dorsal fin, dark diagonal bar extending from eye, diffused line from eye to lip, and 7–9 more or less defined vertical brown bars starting at bases of dorsal fins in
<taxonomicName id="4C604D125F7BFFB7FB11FDA3FA3CFDCD" baseAuthorityName="Ruppell" baseAuthorityYear="1830" box="[1278,1429,585,611]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="diademata">
<emphasis id="B914EA835F7BFFB7FB11FDA3FA3CFDCD" box="[1278,1429,585,611]" italics="true" pageId="7" pageNumber="508">H. diademata</emphasis>
</taxonomicName>
, head and body yellowish white with irregular brown mottling above midlateral and dorsal fins with longitudinal dark submarginal band in
<taxonomicName id="4C604D125F7BFFB7FE59FD7BFDF4FD05" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" box="[438,605,657,683]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="dorsovittata">
<emphasis id="B914EA835F7BFFB7FE59FD7BFDF4FD05" box="[438,605,657,683]" italics="true" pageId="7" pageNumber="508">H. dorsovittata</emphasis>
</taxonomicName>
, head and body transparent whitish with rare dark brown markings on head and along back and with blackish bar at caudal-fin base in
<taxonomicName id="4C604D125F7BFFB7FCC7FD5FFC48FD61" authorityName="Kovacic &amp; Bogorodsky" authorityYear="2014" box="[808,993,693,719]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="species" species="psammophila">
<emphasis id="B914EA835F7BFFB7FCC7FD5FFC48FD61" box="[808,993,693,719]" italics="true" pageId="7" pageNumber="508">H. psammophila</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDF36915F7BFFB7FF28FD30FC51FC41" blockId="7.[151,1437,153,1007]" pageId="7" pageNumber="508">
The presence of three scales observed on the
<typeStatus id="54DB88335F7BFFB7FD53FD30FCB7FD5D" box="[700,798,729,755]" pageId="7" pageNumber="508" type="holotype">holotype</typeStatus>
was included in the description. However, we have taken precaution using the phrase of a few scales in the key and in the diagnosis since the species count, when more material would be collected, could extend the present
<typeStatus id="54DB88335F7BFFB7FD2FFCC8FC8BFC95" box="[704,802,801,827]" pageId="7" pageNumber="508" type="holotype">holotype</typeStatus>
count. The fin meristic characters in diagnosis has to be taken also with cautions since they are based on a single specimen. However, for the most
<taxonomicName id="4C604D125F7BFFB7FB94FCACFAA7FCF1" authorityName="Bleeker" authorityYear="1874" box="[1147,1294,837,863]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="genus">
<emphasis id="B914EA835F7BFFB7FB94FCACFAA7FCF1" box="[1147,1294,837,863]" italics="true" pageId="7" pageNumber="508">Hetereleotris</emphasis>
</taxonomicName>
species with the data known from numerous specimens the second dorsal-fin count and the anal-fin count remain restricted to a single value (
<bibRefCitation id="EFF14B605F7BFFB7FEC2FC64FE1EFC06" author="Hoese, D. F." box="[301,439,909,936]" pageId="7" pageNumber="508" pagination="1 - 25" refId="ref13090" refString="Hoese, D. F. (1986) Descriptions of two new species of Hetereleotris (Pisces: Gobiidae) from the western Indian Ocean, with discussion of related species. J. L. B. Smith Institute of Ichthyology, Special Publication, 41, 1 - 25." type="journal article" year="1986">Hoese 1986</bibRefCitation>
). The pectoral-fin count is more variable within the species (
<bibRefCitation id="EFF14B605F7BFFB7FBB0FC64FB43FC06" author="Hoese, D. F." box="[1119,1258,909,936]" pageId="7" pageNumber="508" pagination="1 - 25" refId="ref13090" refString="Hoese, D. F. (1986) Descriptions of two new species of Hetereleotris (Pisces: Gobiidae) from the western Indian Ocean, with discussion of related species. J. L. B. Smith Institute of Ichthyology, Special Publication, 41, 1 - 25." type="journal article" year="1986">Hoese 1986</bibRefCitation>
). Anyway, even with the overlapping of fin meristics, the new species diagnosis would remain firm based on the large set of other differential characters in comparison with other known
<taxonomicName id="4C604D125F7BFFB7FCEAFC3CFC31FC41" authorityName="Bleeker" authorityYear="1874" box="[773,920,981,1007]" class="Actinopterygii" family="Gobiidae" genus="Hetereleotris" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="508" phylum="Chordata" rank="genus">
<emphasis id="B914EA835F7BFFB7FCEAFC3CFC31FC41" box="[773,920,981,1007]" italics="true" pageId="7" pageNumber="508">Hetereleotris</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
</treatment>
</document>
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