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<document ID-DOI="https://doi.org/10.11646/zootaxa.4982.1.1" ID-GBIF-Dataset="f68b381d-d0dd-44e2-b9fe-6cf3c1aab1fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921842" ID-ZooBank="58388FB8-128A-4381-83D1-3C508D0D3873" checkinTime="1623285106829" checkinUser="plazi" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2021" docId="D446F043FFEEFFC68C92FF39FA1406E4" docLanguage="en" docName="zootaxa.4982.1.1.pdf" docOrigin="Zootaxa 4982 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.9:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="9" docTitle="Bellardithala undetermined" docType="treatment" docVersion="7" lastPageNumber="28" masterDocId="287F883BFFF5FFDD8C05FFAEFF890000" masterDocTitle="An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea" masterLastPageNumber="70" masterPageNumber="1" pageNumber="28" updateTime="1632097679370" updateUser="ExternalLinkService">
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<mods:title>An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Harzhauser, Mathias</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Landau, Bernard</mods:namePart>
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<mods:number>2021-06-09</mods:number>
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<paragraph blockId="27.[151,354,151,213]" box="[151,354,151,178]" pageId="27" pageNumber="28">
<heading bold="true" box="[151,354,151,178]" fontSize="11" level="2" pageId="27" pageNumber="28" reason="4">
<taxonomicName box="[151,354,151,178]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="species">
<emphasis bold="true" box="[151,354,151,178]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[151,311,151,178]" italics="true" pageId="27" pageNumber="28">Bellardithala</emphasis>
sp.
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,241,187,213]" pageId="27" pageNumber="28" type="description">
<paragraph blockId="27.[151,354,151,213]" box="[151,241,187,213]" pageId="27" pageNumber="28">
<figureCitation box="[151,241,187,213]" captionStart="FIGURE 8" captionStartId="22.[151,250,1800,1824]" captionTargetBox="[151,1436,669,1777]" captionTargetId="figure-225@22.[151,1436,669,1777]" captionTargetPageId="22" captionText="FIGURE 8 A1A2. Bellardithala dacica nov. sp., holotype, Lăpugiu de Sus (Romania), NHMW 1854/0035/0105. B1B2. Bellardithala dacica nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0061/0001. C1C2. Bellardithala dacica nov. sp., paratype, Lăpugiu de Sus (Romania), NHMW 2020/0061/0002. D1D2. Bellardithala kostejana (Boettger, 1906), lectotype, Coşteiu de Sus (Romania), SMF 360340. E1E2. Bellardithala kostejana (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 2020/0064/0001. F. Bellardithala kostejana (Boettger, 1906), Lăpugiu de Sus (Romania), NHMW 2020/0064/0002. G. Bellardithala sp., Devínska Nová Ves (Slovakia), Z23408-P2003/27." figureDoi="http://doi.org/10.5281/zenodo.4921858" httpUri="https://zenodo.org/record/4921858/files/figure.png" pageId="27" pageNumber="28">Figs 8G</figureCitation>
</paragraph>
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<subSubSection box="[151,827,256,280]" pageId="27" pageNumber="28" type="reference_group">
<paragraph blockId="27.[151,827,256,280]" box="[151,827,256,280]" pageId="27" pageNumber="28">
<treatmentCitationGroup box="[151,827,256,280]" pageId="27" pageNumber="28">
<taxonomicName authority="" box="[151,356,256,280]" class="Gastropoda" family="Costellariidae" genus="Vexillum" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="subGenus" subGenus="Uromitra">
<emphasis box="[151,238,256,280]" italics="true" pageId="27" pageNumber="28">Vexillum</emphasis>
(
<emphasis box="[253,348,256,279]" italics="true" pageId="27" pageNumber="28">Uromitra</emphasis>
)
</taxonomicName>
sp.—
<treatmentCitation author="Tomasovych, A." box="[413,656,256,280]" page="378" pageId="27" pageNumber="28" year="1998">
<bibRefCitation author="Tomasovych, A." box="[413,656,256,280]" pageId="27" pageNumber="28" pagination="357 - 386" refId="ref62902" refString="Tomasovych, A., (1998) Mollusca from the Devinska Nova Ves - brickyard locality (Bratislava, Slovakia), Badenian. Mineralia Slovaca, 30, 357 - 386. [https: // www. geology. sk / mineralia /]" type="journal article" year="1998">Tomašových 1998: 378</bibRefCitation>
</treatmentCitation>
, pl. 10, figs 34.
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</paragraph>
</subSubSection>
<subSubSection box="[151,1280,325,352]" pageId="27" pageNumber="28" type="materials_examined">
<paragraph blockId="27.[151,1437,325,857]" box="[151,1280,325,352]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[151,262,325,351]" pageId="27" pageNumber="28">Material.</emphasis>
<materialsCitation accessionNumber="Z23408" box="[269,1276,326,352]" collectorName="Bratislava, SL" location="Natural History Museum in Bratislava" pageId="27" pageNumber="28" specimenCount="1">
Inv.
<accessionNumber box="[319,408,326,352]" httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/Z23408" pageId="27" pageNumber="28">Z23408</accessionNumber>
-P2003/27,
<location LSID="urn:lsid:plazi:treatment:D446F043FFEEFFC68C92FF39FA1406E4:5930178EFFEEFFC68E1CFEE8FC410160" box="[537,968,326,352]" name="Natural History Museum in Bratislava" pageId="27" pageNumber="28">
Natural History Museum in
<collectorName box="[852,968,326,352]" pageId="27" pageNumber="28">Bratislava</collectorName>
</location>
, SL:
<quantity box="[1027,1115,326,352]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="27" pageNumber="28" unit="mm" value="9.4">9.4 mm</quantity>
, MD:
<quantity box="[1187,1276,326,352]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="27" pageNumber="28" unit="mm" value="3.3">3.3 mm</quantity>
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.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="description">
<paragraph blockId="27.[151,1437,325,857]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[199,345,361,387]" pageId="27" pageNumber="28">Description.</emphasis>
Shell medium-sized, moderately slender with slightly gradate spire. Protoconch and first teleoconch whorls unknown. Early teleoconch whorls with broad, strongly opisthocline axial ribs, with interspaces of similar width. Axial ribs becoming sigmoidal and densely beaded on later whorls, with two more spiral rows of slightly more prominent beads along adapical suture. About eight spiral rows of beads on penultimate whorl. Beads replaced by prominent, convex spiral cords on strongly constricted base. Aperture is largely destroyed with three prominent columellar folds and much weaker abapical fourth fold. Siphonal canal long.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="discussion">
<paragraph blockId="27.[151,1437,325,857]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[199,332,577,603]" pageId="27" pageNumber="28">Discussion.</emphasis>
<bibRefCitation author="Tomasovych, A." box="[338,562,578,604]" pageId="27" pageNumber="28" pagination="357 - 386" refId="ref62902" refString="Tomasovych, A., (1998) Mollusca from the Devinska Nova Ves - brickyard locality (Bratislava, Slovakia), Badenian. Mineralia Slovaca, 30, 357 - 386. [https: // www. geology. sk / mineralia /]" type="journal article" year="1998">Tomašových (1998)</bibRefCitation>
described a single fragmentary specimen of a
<taxonomicName authorityName="Harzhauser &amp; Landau" authorityYear="2021" box="[1078,1226,578,604]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[1078,1226,578,604]" italics="true" pageId="27" pageNumber="28">Bellardithala</emphasis>
</taxonomicName>
from basinal clays at Devínska Nová Ves. Due to its peculiar sculpture of strongly opisthocline to sigmoidal, beaded axial ribs, this specimen differs from all other Paratethyan
<taxonomicName box="[656,895,650,676]" class="Gastropoda" family="Costellariidae" genus="Bellardithala" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="species">
<emphasis box="[656,804,650,676]" italics="true" pageId="27" pageNumber="28">Bellardithala</emphasis>
species
</taxonomicName>
and might represent a yet undescribed species. Due to its fragmentary preservation, we refrain from naming it formally.
</paragraph>
<paragraph blockId="27.[151,1437,325,857]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[199,438,721,747]" pageId="27" pageNumber="28">Palaeoenvironment.</emphasis>
The mollusc assemblage of Devínska Nová Ves indicates deeper neritic environments (
<bibRefCitation author="Tomasovych, A." box="[159,370,758,784]" pageId="27" pageNumber="28" pagination="357 - 386" refId="ref62902" refString="Tomasovych, A., (1998) Mollusca from the Devinska Nova Ves - brickyard locality (Bratislava, Slovakia), Badenian. Mineralia Slovaca, 30, 357 - 386. [https: // www. geology. sk / mineralia /]" type="journal article" year="1998">Tomašových 1998</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="distribution">
<paragraph blockId="27.[151,1437,325,857]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[199,620,793,820]" pageId="27" pageNumber="28">Distribution in Central Paratethys.</emphasis>
Badenian (middle Miocene):
<emphasis box="[959,1109,795,820]" italics="true" pageId="27" pageNumber="28">Vienna Basin</emphasis>
: Devínska Nová Ves (
<collectingCountry name="Slovakia" pageId="27" pageNumber="28">Slovakia</collectingCountry>
).
</paragraph>
<paragraph blockId="27.[151,622,937,964]" box="[151,622,937,964]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[151,622,937,964]" pageId="27" pageNumber="28">
Genus
<taxonomicName authority="Monterosato, 1917" authorityName="Monterosato" authorityYear="1917" box="[238,622,937,964]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis bold="true" box="[238,381,937,964]" italics="true" pageId="27" pageNumber="28">Ebenomitra</emphasis>
<bibRefCitation author="Monterosato T. A." box="[389,622,938,964]" pageId="27" pageNumber="28" pagination="1 - 28" refId="ref59881" refString="Monterosato T. A. (1917). Molluschi viventi e quaternari raccolti lungo le coste della Tripolitania dall'ing. Camillo Crema. Bollettino della Societa Zoologica Italiana, Series 3, 4, 1 - 28. [https: // www. biodiversitylibrary. org / item / 268776 page / 7 / mode / 1 up]" type="journal article" year="1917">Monterosato, 1917</bibRefCitation>
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="type_taxon">
<paragraph blockId="27.[151,1437,1009,1504]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[151,308,1009,1036]" pageId="27" pageNumber="28">
<typeStatus box="[151,210,1010,1036]" pageId="27" pageNumber="28">Type</typeStatus>
species.
</emphasis>
<taxonomicName authority="Lamarck, 1811" authorityName="Lamarck" authorityYear="1811" box="[318,650,1010,1036]" class="Gastropoda" family="Mitridae" genus="Mitra" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="species">
<emphasis box="[318,469,1010,1036]" italics="true" pageId="27" pageNumber="28">Mitra ebenus</emphasis>
<bibRefCitation author="Lamarck, M." box="[477,650,1010,1036]" pageId="27" pageNumber="28" pagination="195 - 222" refId="ref58636" refString="Lamarck, M. (1811) Sur la determination des especes parmi les animaux sans vertebres, et particulicrement parmi les mollusques testaces. Annales du Museum d'histoire naturelle, Paris, 17, 195 - 222. [https: // www. biodiversitylibrary. org / item / 93161 page / 198 / mode / 1 up] https: // doi. org / 10.5962 / bhl. title. 11732" type="journal article" year="1811">Lamarck, 1811</bibRefCitation>
</taxonomicName>
, subsequent designation by
<bibRefCitation author="Coan, E." box="[973,1179,1010,1036]" pageId="27" pageNumber="28" pagination="127 - 137" refId="ref54988" refString="Coan, E. (1966) Nomenclatural units in the gastropod family Mitridae. Veliger, 9, 127 - 137. [https: // www. biodiversitylibrary. org / item / 134958 page / 161 / mode / 1 up]" type="journal article" year="1966">Coan (1966: 130)</bibRefCitation>
; Present-day, Mediterranean Sea.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="description">
<paragraph blockId="27.[151,1437,1009,1504]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[199,345,1081,1107]" pageId="27" pageNumber="28">Description.</emphasis>
<emphasis italics="true" pageId="27" pageNumber="28">Shell small, turriform or fusiform to widely fusiform. Protoconch bulbous, paucispiral, with about two glossy whorls. Teleoconch of up to 7.5 moderately or strongly convex whorls. Early spire whorls with strong ribs or very strong and wide folds; later whorls with strong wide folds vanishing below periphery of last adult whorl, or smooth; spiral sculpture not pronounced. Siphonal canal short to moderately long, sculptured with between two and four strong oblique cords. Aperture ovate to elongate, sometimes constricted towards siphonal canal; outer aperture lip lirate within. Inner apertural lip with three strong oblique columellar folds, adapicalmost fold strongest</emphasis>
” (
<bibRefCitation author="Fedosov, A. &amp; Puillandre, N. &amp; Herrmann, M. &amp; Dgebuadze, P. &amp; Bouchet, P." box="[330,621,1298,1325]" pageId="27" pageNumber="28" pagination="541 - 626" refId="ref56065" refString="Fedosov, A. E, Puillandre, N., Herrmann, M., Dgebuadze, P. &amp; Bouchet, P. (2017) Phylogeny, systematics, and evolution of the family Costellariidae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 179, 541 - 626. https: // doi. org / 10.1111 / zoj. 12431" type="journal article" year="2017">
Fedosov
<emphasis box="[434,483,1298,1324]" italics="true" pageId="27" pageNumber="28">et al</emphasis>
., 2017: 601
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="discussion">
<paragraph blockId="27.[151,1437,1009,1504]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[199,332,1333,1359]" pageId="27" pageNumber="28">Discussion.</emphasis>
<bibRefCitation author="Fedosov, A. &amp; Puillandre, N. &amp; Herrmann, M. &amp; Dgebuadze, P. &amp; Bouchet, P." box="[340,564,1334,1360]" pageId="27" pageNumber="28" pagination="541 - 626" refId="ref56065" refString="Fedosov, A. E, Puillandre, N., Herrmann, M., Dgebuadze, P. &amp; Bouchet, P. (2017) Phylogeny, systematics, and evolution of the family Costellariidae (Gastropoda: Neogastropoda). Zoological Journal of the Linnean Society, 179, 541 - 626. https: // doi. org / 10.1111 / zoj. 12431" type="journal article" year="2017">
Fedosov
<emphasis box="[443,500,1334,1360]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2017
</bibRefCitation>
considered
<taxonomicName authorityName="Monterosato" authorityYear="1917" box="[701,833,1334,1360]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[701,833,1334,1360]" italics="true" pageId="27" pageNumber="28">Ebenomitra</emphasis>
</taxonomicName>
as a subgenus of
<taxonomicName authorityName="Swainson" authorityYear="1840" box="[1036,1100,1335,1360]" class="Gastropoda" family="Costellariidae" genus="Pusia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[1036,1100,1335,1360]" italics="true" pageId="27" pageNumber="28">Pusia</emphasis>
</taxonomicName>
. The pros and cons of the usage of subgenera was controversially discussed in a plethora of papers (e.g.
<bibRefCitation author="Laurin, M." box="[1025,1169,1370,1396]" pageId="27" pageNumber="28" pagination="131 - 146" refId="ref58888" refString="Laurin, M. (2010) The subjective nature of Linnaean categories and its impact in evolutionary biology and biodiversity studies. Contributions in Zoology, 79, 131 - 146. https: // doi. org / 10.1163 / 18759866 - 07904001" type="journal article" year="2010">Laurin 2010</bibRefCitation>
;
<bibRefCitation author="Puillandre, N. &amp; Duda, T. F. &amp; Meyer, C. &amp; Olivera, B. M. &amp; Bouchet, P." box="[1181,1432,1370,1396]" pageId="27" pageNumber="28" pagination="1 - 23" refId="ref60824" refString="Puillandre, N., Duda, T. F., Meyer, C., Olivera, B. M. &amp; Bouchet, P. (2014) One, four or 100 genera? A new classification of the cone snails. Journal of Molluscan Studies, 8, 1 - 23. https: // doi. org / 10.1093 / mollus / eyu 055" type="journal article" year="2014">
Puillandre
<emphasis box="[1306,1364,1370,1396]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2014
</bibRefCitation>
;
<bibRefCitation author="Teta, P." box="[151,263,1406,1432]" pageId="27" pageNumber="28" pagination="209 - 211" refId="ref62753" refString="Teta, P. (2018) The usage of subgenera in mammalian taxonomy. Mammalia, 83, 209 - 211. https: // doi. org / 10.1515 / mammalia- 2018 - 0059" type="journal article" year="2018">Teta 2018</bibRefCitation>
and references therein). As stated in our previous contributions (Harzhauser &amp; Landau 2016, 2019) we consider supraspecific classification levels as arbitrary and try to avoid the usage of subgenera. Therefore, we prefer to raise
<taxonomicName authorityName="Monterosato" authorityYear="1917" box="[240,372,1478,1504]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="genus">
<emphasis box="[240,372,1478,1504]" italics="true" pageId="27" pageNumber="28">Ebenomitra</emphasis>
</taxonomicName>
to full genus rank.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="28" type="key">
<paragraph blockId="27.[151,650,1585,1612]" box="[151,650,1585,1612]" pageId="27" pageNumber="28">
<emphasis bold="true" box="[151,650,1585,1612]" pageId="27" pageNumber="28">
Key to Paratethyan
<taxonomicName box="[403,645,1585,1612]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="undetermined">
<emphasis bold="true" box="[403,546,1585,1612]" italics="true" pageId="27" pageNumber="28">Ebenomitra</emphasis>
species
</taxonomicName>
:
</emphasis>
</paragraph>
<key pageId="27" pageNumber="28">
<keyStep pageId="27" pageNumber="28">
<paragraph blockId="27.[151,1437,1655,1765]" box="[151,1436,1655,1677]" pageId="27" pageNumber="28">
<keyLead box="[151,1436,1655,1677]" pageId="27" pageNumber="28">
1. Shell very broadly fusiform....................................................................
<taxonomicName authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Andrzejowski" baseAuthorityYear="1830" box="[1312,1436,1655,1676]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="leucozona">
<emphasis box="[1312,1436,1655,1676]" italics="true" pageId="27" pageNumber="28">E. leucozona</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="27.[151,1437,1655,1765]" box="[151,1436,1684,1706]" pageId="27" pageNumber="28">
<keyLead box="[151,1436,1684,1706]" pageId="27" pageNumber="28">
<emphasis box="[151,159,1685,1706]" italics="true" pageId="27" pageNumber="28">-</emphasis>
Shell narrow fusiform.................................................................................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep pageId="27" pageNumber="28">
<paragraph blockId="27.[151,1437,1655,1765]" box="[151,1436,1713,1735]" pageId="27" pageNumber="28">
<keyLead box="[151,1436,1713,1735]" pageId="27" pageNumber="28">
2. Axial sculpture restricted to early teleoconch whorls........................................
<taxonomicName authorityName="Harzhauser &amp; Landau" authorityYear="2021" baseAuthorityName="Boettger" baseAuthorityYear="1906" box="[1226,1436,1713,1735]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="pseudopyramidella">
<emphasis box="[1226,1436,1713,1735]" italics="true" pageId="27" pageNumber="28">E. pseudopyramidella</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph blockId="27.[151,1437,1655,1765]" box="[151,1437,1743,1765]" pageId="27" pageNumber="28">
<keyLead box="[151,1437,1743,1765]" pageId="27" pageNumber="28">
- Axial sculpture almost absent, suggestion of ribs on last whorl....................................
<taxonomicName authorityName="Friedberg" authorityYear="1928" box="[1271,1437,1743,1764]" class="Gastropoda" family="Costellariidae" genus="Ebenomitra" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="januszkiewiczi">
<emphasis box="[1271,1437,1743,1764]" italics="true" pageId="27" pageNumber="28">E. januszkiewiczi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>