treatments-xml/data/D4/3E/03/D43E034BFF994160FF17FE0764A1FE78.xml

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<mods:title>Species review of the genus Gnypeta Thomson from Canada, Alaska and Greenland (Coleoptera, Staphylinidae, Aleocharinae): systematics, bionomics and distribution</mods:title>
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<emphasis box="[140,216,448,474]" inLineHeading="true" pageId="4" pageNumber="15" reason="1">Genus</emphasis>
<taxonomicName authority="Thomson" authorityName="Thomson" box="[223,444,448,474]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[223,324,448,474]" italics="true" pageId="4" pageNumber="15">Gnypeta</emphasis>
Thomson
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<emphasis box="[140,230,519,545]" italics="true" pageId="4" pageNumber="15">Gnypeta</emphasis>
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<bibRefCitation author="Blackwelder" firstAuthor="Blackwelder" page="173" pageId="4" pageNumber="15" pagination="1 - 483" refId="ref22950" refString="Blackwelder RE (1952) The generic names of the beetle family Staphylinidae with an essay on genotypy. Bulletin of the United States National Museum 200: I-IV, 1 - 483." type="journal article" year="1952">Blackwelder 1952: 173</bibRefCitation>
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species:
<taxonomicName authority="Erichson 1839" authorityName="Erichson" authorityYear="1839" class="Insecta" family="Staphylinidae" genus="Homalota" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="labilis">
<emphasis italics="true" pageId="4" pageNumber="15">Homalota labilis</emphasis>
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(=
<taxonomicName authority="Mannerheim 1830" authorityName="Mannerheim" authorityYear="1830" box="[455,929,589,615]" class="Insecta" family="Staphylinidae" genus="Bolitochara" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis box="[455,703,589,615]" italics="true" pageId="4" pageNumber="15">Bolitochara carbonaria</emphasis>
<bibRefCitation author="Mannerheim" box="[711,929,589,615]" firstAuthor="Mannerheim" pageId="4" pageNumber="15" pagination="415 - 501" refId="ref23744" refString="Mannerheim CG (1830) Precis d'un nouvel arrangement de la famille des brachelytres de l'ordre des insectes coleopteres. Memoires de l'Academie Imperiale des Sciences de St. - Petersburg 1: 415 - 501. Sep. St. Petersburg, 87 pp." type="book chapter" year="1830">Mannerheim 1830</bibRefCitation>
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<emphasis box="[140,221,624,650]" italics="true" pageId="4" pageNumber="15">Euliusa</emphasis>
<bibRefCitation author="Casey" box="[228,420,624,651]" firstAuthor="Casey" page="215" pageId="4" pageNumber="15" pagination="125 - 434" refId="ref23153" refString="Casey TL (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis 16: 125 - 434." type="journal article" year="1906">Casey 1906: 215</bibRefCitation>
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<treatmentCitation author="Moore I &amp; Legner EF" box="[431,766,624,651]" page="421" pageId="4" pageNumber="15" year="1975">
<bibRefCitation author="Moore and Legner" box="[431,766,624,651]" firstAuthor="Moore" page="421" pageId="4" pageNumber="15" pagination="1 - 514" refId="ref23790" refString="Moore I., Legner EF (1975) A catalogue of the Staphylinidae of America north of Mexico (Coleoptera). University of California Division of Agricultural Sciences. Special Publication No. 3015: 1 - 514." type="book chapter" year="1975">Moore and Legner 1975: 421</bibRefCitation>
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<treatmentCitation author="Seevers CH" box="[777,969,624,650]" page="83" pageId="4" pageNumber="15" year="1978">
<bibRefCitation author="Seevers" box="[777,969,624,650]" firstAuthor="Seevers" page="83" pageId="4" pageNumber="15" pagination="1 - 289" refId="ref24091" refString="Seevers CH (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoology 71: i-vi, 1 - 289." type="journal article" year="1978">Seevers 1978: 83</bibRefCitation>
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. Synonymized by
<treatmentCitation author="Blackwelder RE" box="[682,948,659,686]" page="173" pageId="4" pageNumber="15" year="1952">
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</treatmentCitation>
.
<typeStatus box="[960,1015,660,686]" pageId="4" pageNumber="15">Type</typeStatus>
species:
<taxonomicName authority="Bernhauer 1905" authorityName="Bernhauer" authorityYear="1905" box="[185,541,694,721]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="lucens">
<emphasis box="[185,347,694,721]" italics="true" pageId="4" pageNumber="15">Gnypeta lucens</emphasis>
<bibRefCitation author="Bernhauer" box="[354,541,694,721]" firstAuthor="Bernhauer" pageId="4" pageNumber="15" pagination="580 - 595" refId="ref22896" refString="Bernhauer M (1905) Folge neuer Staphyliniden der palaarktischen Fauna, nebst Bemerkungen. Verhandlungen der Kaiserlich-Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 55: 580 - 595." type="journal article" year="1905">Bernhauer 1905</bibRefCitation>
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.
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<treatmentCitation author="Casey TL" box="[140,470,729,756]" page="196" pageId="4" pageNumber="15" year="1906">
<taxonomicName authority="Casey 1906: 196" authorityName="Casey" authorityPageNumber="196" authorityYear="1906" box="[140,470,729,756]" class="Insecta" family="Staphylinidae" genus="Gnypetoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis box="[140,263,730,756]" italics="true" pageId="4" pageNumber="15">Gnypetoma</emphasis>
<bibRefCitation author="Casey" box="[272,470,729,756]" firstAuthor="Casey" page="196" pageId="4" pageNumber="15" pagination="125 - 434" refId="ref23153" refString="Casey TL (1906) Observations on the staphylinid groups Aleocharinae and Xantholinini, chiefly of America. Transactions of the Academy of Sciences of St. Louis 16: 125 - 434." type="journal article" year="1906">Casey 1906: 196</bibRefCitation>
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<treatmentCitation author="Ashe JS" page="363" pageId="4" pageNumber="15" year="2001">
<bibRefCitation author="Ashe" firstAuthor="Ashe" page="363" pageId="4" pageNumber="15" pagination="299 - 374" refId="ref22843" refString="Ashe JS (2001) Keys to the tribes and genera of Nearctic Aleocharinae. In: Arnett RH, Thomas MC (Eds) American Beetles. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press LLC, Boca Raton, Florida, 299 - 374." type="book chapter" year="2001">Ashe 2001: 363</bibRefCitation>
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<treatmentCitation author="Smetana A" box="[323,554,765,791]" page="489" pageId="4" pageNumber="15" year="2004">
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. Synonymized by
<treatmentCitation author="Blackwelder RE" box="[768,1039,765,792]" page="173" pageId="4" pageNumber="15" year="1952">
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</treatmentCitation>
.
<typeStatus box="[1052,1107,765,791]" pageId="4" pageNumber="15">Type</typeStatus>
species:
<taxonomicName authority="LeConte 1863" authorityName="LeConte" authorityYear="1863" box="[276,647,800,826]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="15" phylum="Arthropoda" rank="species" species="baltifera">
<emphasis box="[276,471,800,826]" italics="true" pageId="4" pageNumber="15">Tachyusa baltifera</emphasis>
<bibRefCitation author="LeConte" box="[479,647,800,826]" firstAuthor="LeConte" pageId="4" pageNumber="15" pagination="1 - 56" refId="ref23584" refString="LeConte JL (1863) New species of North American Coleoptera. Smithsonian Miscellaneous Collections 6: 1 - 56." type="journal article" year="1863">LeConte 1863</bibRefCitation>
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.
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</paragraph>
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<paragraph blockId="4.[140,1108,870,1672]" box="[140,259,870,896]" pageId="4" pageNumber="15">
<emphasis bold="true" box="[140,259,870,896]" inLineHeading="true" pageId="4" pageNumber="15" reason="1">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="4.[140,1108,870,1672]" pageId="4" pageNumber="15">
Members of this genus may be distinguished by the following combination of characters: tarsi 4, 5, 5-articulated (
<figureCitation box="[504,576,941,967]" captionStart="Fig" captionStartId="5.[140,174,1654,1676]" captionTargetBox="[243,974,176,1625]" captionTargetId="figure@5.[136,1110,161,1637]" captionTargetPageId="5" captionText="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774051" httpUri="https://zenodo.org/record/3774051/files/figure.png" pageId="4" pageNumber="15">Figs 1</figureCitation>
,
<figureCitation box="[594,611,941,967]" captionStart="Fig" captionStartId="6.[140,174,1654,1676]" captionTargetBox="[197,1035,184,1604]" captionTargetId="figure@6.[136,1110,161,1637]" captionTargetPageId="6" captionText="Fig. 2. Gnypeta nigrella (LeConte), in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770346" httpUri="https://zenodo.org/record/3770346/files/figure.png" pageId="4" pageNumber="15">2</figureCitation>
); body medium sized, length 2.3-3.6 mm, moderately to strongly glossy, integument finely punctate and pubescent, pubescence moderately long and often silky in appearance; head and pronotum distinctly narrower than elytra (
<figureCitation box="[281,394,1046,1073]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Figs 3-8" captionStart-3="Figs 9-14" captionStart-4="Figs 15-20" captionStartId-0="5.[140,174,1654,1676]" captionStartId-1="6.[140,174,1654,1676]" captionStartId-2="7.[140,184,1622,1644]" captionStartId-3="8.[140,184,1622,1644]" captionStartId-4="9.[140,184,1622,1644]" captionTargetBox-0="[243,974,176,1625]" captionTargetBox-1="[197,1035,184,1604]" captionTargetBox-2="[181,994,433,1595]" captionTargetBox-3="[237,1043,441,1595]" captionTargetBox-4="[214,1020,442,1588]" captionTargetId-0="figure@5.[136,1110,161,1637]" captionTargetId-1="figure@6.[136,1110,161,1637]" captionTargetId-2="figure@7.[173,1072,410,1609]" captionTargetId-3="figure@8.[176,1074,410,1609]" captionTargetId-4="figure@9.[176,1072,358,1560]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." captionText-1="Fig. 2. Gnypeta nigrella (LeConte), in ventral view. Scale = 1 mm." captionText-2="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" captionText-3="Figs 9-14. Gnypeta species: 9, G. brevicornis; 10, G. caerulea; 11, G. lohsei; 12, G. crebrepunctata; 13, G. manitobae [apical part of abdomen removed]; 14, G. uteana. Scale = 1 mm" captionText-4="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.3774051" figureDoi-1="http://doi.org/10.5281/zenodo.3770346" figureDoi-2="http://doi.org/10.5281/zenodo.3774053" figureDoi-3="http://doi.org/10.5281/zenodo.3774055" figureDoi-4="http://doi.org/10.5281/zenodo.3774057" httpUri-0="https://zenodo.org/record/3774051/files/figure.png" httpUri-1="https://zenodo.org/record/3770346/files/figure.png" httpUri-2="https://zenodo.org/record/3774053/files/figure.png" httpUri-3="https://zenodo.org/record/3774055/files/figure.png" httpUri-4="https://zenodo.org/record/3774057/files/figure.png" pageId="4" pageNumber="15">Figs 1-20</figureCitation>
); abdomen as broad as elytra at base or moderately narrower (
<figureCitation box="[147,258,1082,1108]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Figs 3-8" captionStart-3="Figs 9-14" captionStart-4="Figs 15-20" captionStartId-0="5.[140,174,1654,1676]" captionStartId-1="6.[140,174,1654,1676]" captionStartId-2="7.[140,184,1622,1644]" captionStartId-3="8.[140,184,1622,1644]" captionStartId-4="9.[140,184,1622,1644]" captionTargetBox-0="[243,974,176,1625]" captionTargetBox-1="[197,1035,184,1604]" captionTargetBox-2="[181,994,433,1595]" captionTargetBox-3="[237,1043,441,1595]" captionTargetBox-4="[214,1020,442,1588]" captionTargetId-0="figure@5.[136,1110,161,1637]" captionTargetId-1="figure@6.[136,1110,161,1637]" captionTargetId-2="figure@7.[173,1072,410,1609]" captionTargetId-3="figure@8.[176,1074,410,1609]" captionTargetId-4="figure@9.[176,1072,358,1560]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." captionText-1="Fig. 2. Gnypeta nigrella (LeConte), in ventral view. Scale = 1 mm." captionText-2="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" captionText-3="Figs 9-14. Gnypeta species: 9, G. brevicornis; 10, G. caerulea; 11, G. lohsei; 12, G. crebrepunctata; 13, G. manitobae [apical part of abdomen removed]; 14, G. uteana. Scale = 1 mm" captionText-4="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.3774051" figureDoi-1="http://doi.org/10.5281/zenodo.3770346" figureDoi-2="http://doi.org/10.5281/zenodo.3774053" figureDoi-3="http://doi.org/10.5281/zenodo.3774055" figureDoi-4="http://doi.org/10.5281/zenodo.3774057" httpUri-0="https://zenodo.org/record/3774051/files/figure.png" httpUri-1="https://zenodo.org/record/3770346/files/figure.png" httpUri-2="https://zenodo.org/record/3774053/files/figure.png" httpUri-3="https://zenodo.org/record/3774055/files/figure.png" httpUri-4="https://zenodo.org/record/3774057/files/figure.png" pageId="4" pageNumber="15">Figs 1-20</figureCitation>
), subparallel (
<figureCitation box="[421,517,1082,1108]" captionStart="Figs 3-8" captionStartId="7.[140,184,1622,1644]" captionTargetBox="[181,994,433,1595]" captionTargetId="figure@7.[173,1072,410,1609]" captionTargetPageId="7" captionText="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="4" pageNumber="15">Figs 3-7</figureCitation>
), or broadening apically (
<figureCitation box="[815,889,1082,1108]" captionStart="Figs 9-14" captionStartId="8.[140,184,1622,1644]" captionTargetBox="[237,1043,441,1595]" captionTargetId="figure@8.[176,1074,410,1609]" captionTargetPageId="8" captionText="Figs 9-14. Gnypeta species: 9, G. brevicornis; 10, G. caerulea; 11, G. lohsei; 12, G. crebrepunctata; 13, G. manitobae [apical part of abdomen removed]; 14, G. uteana. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774055" httpUri="https://zenodo.org/record/3774055/files/figure.png" pageId="4" pageNumber="15">Figs 9</figureCitation>
,
<figureCitation box="[903,974,1082,1108]" captionStart="Figs 15-20" captionStartId="9.[140,184,1622,1644]" captionTargetBox="[214,1020,442,1588]" captionTargetId="figure@9.[176,1072,358,1560]" captionTargetPageId="9" captionText="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi="http://doi.org/10.5281/zenodo.3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="4" pageNumber="15">18 20</figureCitation>
), first three visible tergites with deep basal impressions bearing coarse punctures but without pronounced longitudinal ridges (
<figureCitation box="[476,541,1152,1178]" captionStart="Fig" captionStartId="5.[140,174,1654,1676]" captionTargetBox="[243,974,176,1625]" captionTargetId="figure@5.[136,1110,161,1637]" captionTargetPageId="5" captionText="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774051" httpUri="https://zenodo.org/record/3774051/files/figure.png" pageId="4" pageNumber="15">Fig. 1</figureCitation>
); postgenal carinae incomplete or absent (
<bibRefCitation author="Seevers" firstAuthor="Seevers" pageId="4" pageNumber="15" pagination="1 - 289" refId="ref24091" refString="Seevers CH (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoology 71: i-vi, 1 - 289." type="journal article" year="1978">Seevers 1978</bibRefCitation>
); pronotum broadest in apical third or in the middle of the disc, and converging apically and basally (
<figureCitation box="[375,441,1222,1249]" captionStart="Fig" captionStartId="5.[140,174,1654,1676]" captionTargetBox="[243,974,176,1625]" captionTargetId="figure@5.[136,1110,161,1637]" captionTargetPageId="5" captionText="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774051" httpUri="https://zenodo.org/record/3774051/files/figure.png" pageId="4" pageNumber="15">Fig. 1</figureCitation>
), pubescence distributed postero-laterad from the midline of the disc (
<figureCitation box="[275,386,1258,1284]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Figs 3-8" captionStart-3="Figs 9-14" captionStart-4="Figs 15-20" captionStartId-0="5.[140,174,1654,1676]" captionStartId-1="6.[140,174,1654,1676]" captionStartId-2="7.[140,184,1622,1644]" captionStartId-3="8.[140,184,1622,1644]" captionStartId-4="9.[140,184,1622,1644]" captionTargetBox-0="[243,974,176,1625]" captionTargetBox-1="[197,1035,184,1604]" captionTargetBox-2="[181,994,433,1595]" captionTargetBox-3="[237,1043,441,1595]" captionTargetBox-4="[214,1020,442,1588]" captionTargetId-0="figure@5.[136,1110,161,1637]" captionTargetId-1="figure@6.[136,1110,161,1637]" captionTargetId-2="figure@7.[173,1072,410,1609]" captionTargetId-3="figure@8.[176,1074,410,1609]" captionTargetId-4="figure@9.[176,1072,358,1560]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." captionText-1="Fig. 2. Gnypeta nigrella (LeConte), in ventral view. Scale = 1 mm." captionText-2="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" captionText-3="Figs 9-14. Gnypeta species: 9, G. brevicornis; 10, G. caerulea; 11, G. lohsei; 12, G. crebrepunctata; 13, G. manitobae [apical part of abdomen removed]; 14, G. uteana. Scale = 1 mm" captionText-4="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.3774051" figureDoi-1="http://doi.org/10.5281/zenodo.3770346" figureDoi-2="http://doi.org/10.5281/zenodo.3774053" figureDoi-3="http://doi.org/10.5281/zenodo.3774055" figureDoi-4="http://doi.org/10.5281/zenodo.3774057" httpUri-0="https://zenodo.org/record/3774051/files/figure.png" httpUri-1="https://zenodo.org/record/3770346/files/figure.png" httpUri-2="https://zenodo.org/record/3774053/files/figure.png" httpUri-3="https://zenodo.org/record/3774055/files/figure.png" httpUri-4="https://zenodo.org/record/3774057/files/figure.png" pageId="4" pageNumber="15">Figs 1-20</figureCitation>
); elytra much broader than both head or pronotum, and often with wavy pubescence pattern on both sides (
<figureCitation box="[666,777,1293,1319]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Figs 3-8" captionStart-3="Figs 9-14" captionStart-4="Figs 15-20" captionStartId-0="5.[140,174,1654,1676]" captionStartId-1="6.[140,174,1654,1676]" captionStartId-2="7.[140,184,1622,1644]" captionStartId-3="8.[140,184,1622,1644]" captionStartId-4="9.[140,184,1622,1644]" captionTargetBox-0="[243,974,176,1625]" captionTargetBox-1="[197,1035,184,1604]" captionTargetBox-2="[181,994,433,1595]" captionTargetBox-3="[237,1043,441,1595]" captionTargetBox-4="[214,1020,442,1588]" captionTargetId-0="figure@5.[136,1110,161,1637]" captionTargetId-1="figure@6.[136,1110,161,1637]" captionTargetId-2="figure@7.[173,1072,410,1609]" captionTargetId-3="figure@8.[176,1074,410,1609]" captionTargetId-4="figure@9.[176,1072,358,1560]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionText-0="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." captionText-1="Fig. 2. Gnypeta nigrella (LeConte), in ventral view. Scale = 1 mm." captionText-2="Figs 3-8. Gnypeta species: 3, G. ashei; 4, G. brincki; 5, G. atrolucens; 6, G. sellmani: 7, G. dentata; and 8, G. groenlandica. Scale = 1 mm" captionText-3="Figs 9-14. Gnypeta species: 9, G. brevicornis; 10, G. caerulea; 11, G. lohsei; 12, G. crebrepunctata; 13, G. manitobae [apical part of abdomen removed]; 14, G. uteana. Scale = 1 mm" captionText-4="Figs 15-20. Gnypeta species: 15, G. carbonaria; 16, G. helenae; 17, G. canadensis; 18, G. nigrella; 19, G. saccharina; and 20, G. minuta [18-20, apical part of abdomen removed]. Scale = 1 mm" figureDoi-0="http://doi.org/10.5281/zenodo.3774051" figureDoi-1="http://doi.org/10.5281/zenodo.3770346" figureDoi-2="http://doi.org/10.5281/zenodo.3774053" figureDoi-3="http://doi.org/10.5281/zenodo.3774055" figureDoi-4="http://doi.org/10.5281/zenodo.3774057" httpUri-0="https://zenodo.org/record/3774051/files/figure.png" httpUri-1="https://zenodo.org/record/3770346/files/figure.png" httpUri-2="https://zenodo.org/record/3774053/files/figure.png" httpUri-3="https://zenodo.org/record/3774055/files/figure.png" httpUri-4="https://zenodo.org/record/3774057/files/figure.png" pageId="4" pageNumber="15">Figs 1-20</figureCitation>
); mesoventral process moderately broad, attaining middle of mesocoxae, its apex truncate (
<figureCitation box="[880,951,1328,1354]" captionStart="Fig" captionStartId="6.[140,174,1654,1676]" captionTargetBox="[197,1035,184,1604]" captionTargetId="figure@6.[136,1110,161,1637]" captionTargetPageId="6" captionText="Fig. 2. Gnypeta nigrella (LeConte), in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770346" httpUri="https://zenodo.org/record/3770346/files/figure.png" pageId="4" pageNumber="15">Fig. 2</figureCitation>
); metaventral process broadly rounded (
<figureCitation box="[430,497,1363,1389]" captionStart="Fig" captionStartId="6.[140,174,1654,1676]" captionTargetBox="[197,1035,184,1604]" captionTargetId="figure@6.[136,1110,161,1637]" captionTargetPageId="6" captionText="Fig. 2. Gnypeta nigrella (LeConte), in ventral view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3770346" httpUri="https://zenodo.org/record/3770346/files/figure.png" pageId="4" pageNumber="15">Fig. 2</figureCitation>
); first visible three tergites with deep basal impressions bearing coarse punctures; legs long and slender; basal article of metatarsus moderately elongate and usually shorter than the two following articles combined (
<figureCitation box="[937,1002,1434,1460]" captionStart="Fig" captionStartId="5.[140,174,1654,1676]" captionTargetBox="[243,974,176,1625]" captionTargetId="figure@5.[136,1110,161,1637]" captionTargetPageId="5" captionText="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774051" httpUri="https://zenodo.org/record/3774051/files/figure.png" pageId="4" pageNumber="15">Fig. 1</figureCitation>
). GENITAL STRUCTURES: median lobe of aedeagus consists of an enlarged, swollen bulbus and short, triangularly produced tubus (
<figureCitation box="[597,729,1504,1530]" captionStart="Figs 39-47" captionStartId="17.[140,184,1590,1612]" captionTargetBox="[140,1101,234,1493]" captionTargetId="figure@17.[140,1108,227,1517]" captionTargetPageId="17" captionText="Figs 39-47. Genital structures of Gnypeta ashei: 39, median lobe of the aedeagus in lateral view, 40, dorsal view, 41, ventral view; 42, paramere; 43, male tergite 8; 44, male sternite 8; 45, spermatheca; 46, female tergite 8; 47, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774096" httpUri="https://zenodo.org/record/3774096/files/figure.png" pageId="4" pageNumber="15">Figs 40, 41</figureCitation>
,
<figureCitation box="[742,774,1504,1530]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="4" pageNumber="15">49</figureCitation>
,
<figureCitation box="[786,818,1504,1531]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="4" pageNumber="15">58</figureCitation>
,
<figureCitation box="[831,908,1504,1530]" captionStart="Figs 66-74" captionStartId="24.[140,184,1590,1612]" captionTargetBox="[140,1088,235,1499]" captionTargetId="figure@24.[140,1108,233,1523]" captionTargetPageId="24" captionText="Figs 66-74. Genital structures of Gnypeta sellmani: 66, median lobe of the aedeagus in lateral view, 67, dorsal view, 68, apical part of median lobe in ventral view; 69, paramere; 70, male tergite 8; 71, male sternite 8; 72, spermatheca; 73, female tergite 8; 74, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774102" httpUri="https://zenodo.org/record/3774102/files/figure.png" pageId="4" pageNumber="15">67, 68</figureCitation>
,
<figureCitation box="[921,953,1504,1530]" captionStart="Figs 75-82" captionStartId="27.[140,184,1590,1612]" captionTargetBox="[150,1098,227,1521]" captionTargetId="figure@27.[136,1110,216,1540]" captionTargetPageId="27" captionText="Figs 75-82. Genital structures of Gnypeta dentata: 75, median lobe of the aedeagus in lateral view, 76, dorsal view; 77, paramere; 78, male tergite 8; 79, male sternite 8; 80, spermatheca; 81, female tergite 8; 82, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774104" httpUri="https://zenodo.org/record/3774104/files/figure.png" pageId="4" pageNumber="15">76</figureCitation>
,
<figureCitation box="[966,998,1504,1530]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="4" pageNumber="15">84</figureCitation>
,
<figureCitation box="[1011,1043,1504,1530]" captionStart-0="Figs 91-98" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStartId-0="31.[140,184,1590,1612]" captionStartId-1="60.[140,174,1558,1580]" captionStartId-2="60.[140,174,1622,1644]" captionStartId-3="60.[140,174,1654,1676]" captionStartId-4="61.[140,174,883,905]" captionStartId-5="61.[140,174,915,937]" captionStartId-6="61.[140,174,947,969]" captionStartId-7="61.[140,174,979,1001]" captionTargetBox-0="[151,1101,241,1488]" captionTargetBox-1="[140,1106,794,1132]" captionTargetBox-2="[140,1101,1127,1467]" captionTargetBox-3="[140,1101,1127,1467]" captionTargetBox-4="[140,1106,165,502]" captionTargetBox-5="[140,1106,165,502]" captionTargetBox-6="[140,1106,515,855]" captionTargetBox-7="[140,1106,515,855]" captionTargetId-0="figure@31.[130,1115,227,1529]" captionTargetId-1="figure@60.[140,1108,447,1475]" captionTargetPageId-0="31" captionTargetPageId-1="60" captionTargetPageId-2="60" captionTargetPageId-3="60" captionTargetPageId-4="61" captionTargetPageId-5="61" captionTargetPageId-6="61" captionTargetPageId-7="61" captionText-0="Figs 91-98. Genital structures of Gnypeta brevicornis: 91, median lobe of the aedeagus in lateral view, 92, ventral view; 93, paramere; 94, male tergite 8; 95, male sternite 8; 96, spermatheca; 97, female tergite 8; 98, female sternite 8." captionText-1="Fig. 192. Collection localities in Canada of G. atrolucens and G. nigrella." captionText-2="Fig. 194. Collection localities in Canada of G. crebrepunctata, G. dentata, and G. helenae." captionText-3="Fig. 195. Collection localities in Canada and Alaska of G. groenlandica." captionText-4="Fig. 196. Collection localities in Canada of G. brevicornis and G. uteana." captionText-5="Fig. 197. Collection localities in Canada and Alaska of G. caerulea." captionText-6="Fig. 198. Collection localities in Canada and Alaska of G. lohsei and G. canadiensis." captionText-7="Fig. 199. Collection localities in Canada and Alaska of G. carbonaria and G. saccharina." figureDoi-0="http://doi.org/10.5281/zenodo.3774108" figureDoi-1="http://doi.org/10.5281/zenodo.3774138" figureDoi-2="http://doi.org/10.5281/zenodo.3774142" figureDoi-3="http://doi.org/10.5281/zenodo.3774144" figureDoi-4="http://doi.org/10.5281/zenodo.3774146" figureDoi-5="http://doi.org/10.5281/zenodo.3774148" figureDoi-6="http://doi.org/10.5281/zenodo.3774150" figureDoi-7="http://doi.org/10.5281/zenodo.3774152" httpUri-0="https://zenodo.org/record/3774108/files/figure.png" httpUri-1="https://zenodo.org/record/3774138/files/figure.png" httpUri-2="https://zenodo.org/record/3774142/files/figure.png" httpUri-3="https://zenodo.org/record/3774144/files/figure.png" httpUri-4="https://zenodo.org/record/3774146/files/figure.png" httpUri-5="https://zenodo.org/record/3774148/files/figure.png" httpUri-6="https://zenodo.org/record/3774150/files/figure.png" httpUri-7="https://zenodo.org/record/3774152/files/figure.png" pageId="4" pageNumber="15">92</figureCitation>
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<figureCitation box="[1056,1103,1504,1530]" captionStart="Figs 99-107" captionStartId="33.[140,184,1590,1612]" captionTargetBox="[147,1102,243,1529]" captionTargetId="figure@33.[137,1111,233,1559]" captionTargetPageId="33" captionText="Figs 99-107. Genital structures of Gnypeta brincki: 99, median lobe of the aedeagus in lateral view, 100, dorsal view, 101, apical part of median lobe in ventral view; 102, paramere; 103, male tergite 8; 104, male sternite 8; 105, spermatheca; 106, female tergite 8; 107, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774110" httpUri="https://zenodo.org/record/3774110/files/figure.png" pageId="4" pageNumber="15">100</figureCitation>
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<figureCitation box="[140,187,1539,1565]" captionStart="Figs 108-116" captionStartId="36.[140,184,1590,1612]" captionTargetBox="[144,1104,247,1505]" captionTargetId="figure@36.[143,1111,230,1520]" captionTargetPageId="36" captionText="Figs 108-116. Genital structures of Gnypeta lohsei: 108, median lobe of the aedeagus in lateral view, 109, dorsal view, 110, apical part of median lobe in ventral view; 111, paramere; 112, male tergite 8; 113, male sternite 8; 114, spermatheca; 115, female tergite 8; 116, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774112" httpUri="https://zenodo.org/record/3774112/files/figure.png" pageId="4" pageNumber="15">109</figureCitation>
,
<figureCitation box="[198,245,1539,1565]" captionStart="Figs 108-116" captionStartId="36.[140,184,1590,1612]" captionTargetBox="[144,1104,247,1505]" captionTargetId="figure@36.[143,1111,230,1520]" captionTargetPageId="36" captionText="Figs 108-116. Genital structures of Gnypeta lohsei: 108, median lobe of the aedeagus in lateral view, 109, dorsal view, 110, apical part of median lobe in ventral view; 111, paramere; 112, male tergite 8; 113, male sternite 8; 114, spermatheca; 115, female tergite 8; 116, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774112" httpUri="https://zenodo.org/record/3774112/files/figure.png" pageId="4" pageNumber="15">110</figureCitation>
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<figureCitation box="[256,302,1539,1565]" captionStart="Figs 117-124" captionStartId="39.[140,184,1590,1612]" captionTargetBox="[140,1108,219,1480]" captionTargetId="figure@39.[140,1108,219,1510]" captionTargetPageId="39" captionText="Figs 117-124. Genital structures of Gnypeta crebrepunctata: 117, median lobe of the aedeagus in lateral view, 118, dorsal view; 119, paramere; 120, male tergite 8; 121, male sternite 8; 122, spermatheca; 123, female tergite 8; 124, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774114" httpUri="https://zenodo.org/record/3774114/files/figure.png" pageId="4" pageNumber="15">118</figureCitation>
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<figureCitation box="[314,360,1539,1565]" captionStart="Figs 130-138" captionStartId="44.[140,184,1590,1612]" captionTargetBox="[140,1097,230,1503]" captionTargetId="figure@44.[140,1108,228,1520]" captionTargetPageId="44" captionText="Figs 130-138. Genital structures of Gnypeta uteana: 130, median lobe of the aedeagus in lateral view, 131, dorsal view, 132, ventral view; 133, paramere; 134, male tergite 8; 135, male sternite 8; 136, spermatheca; 137, female tergite 8; 138, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774118" httpUri="https://zenodo.org/record/3774118/files/figure.png" pageId="4" pageNumber="15">131</figureCitation>
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<figureCitation box="[372,419,1539,1565]" captionStart="Figs 130-138" captionStartId="44.[140,184,1590,1612]" captionTargetBox="[140,1097,230,1503]" captionTargetId="figure@44.[140,1108,228,1520]" captionTargetPageId="44" captionText="Figs 130-138. Genital structures of Gnypeta uteana: 130, median lobe of the aedeagus in lateral view, 131, dorsal view, 132, ventral view; 133, paramere; 134, male tergite 8; 135, male sternite 8; 136, spermatheca; 137, female tergite 8; 138, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774118" httpUri="https://zenodo.org/record/3774118/files/figure.png" pageId="4" pageNumber="15">132</figureCitation>
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<figureCitation box="[430,477,1539,1565]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="4" pageNumber="15">140</figureCitation>
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<figureCitation box="[489,535,1539,1565]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="4" pageNumber="15">141</figureCitation>
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<figureCitation box="[547,594,1539,1565]" captionStart="Figs 148-155" captionStartId="49.[140,184,1590,1612]" captionTargetBox="[142,1103,241,1513]" captionTargetId="figure@49.[140,1111,225,1529]" captionTargetPageId="49" captionText="Figs 148-155. Genital structures of Gnypeta helenae: 148, median lobe of the aedeagus in lateral view, 149, dorsal view; 150, paramere; 151, male tergite 8; 152, male sternite 8; 153, spermatheca; 154, female tergite 8; 155, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774122" httpUri="https://zenodo.org/record/3774122/files/figure.png" pageId="4" pageNumber="15">149</figureCitation>
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<figureCitation box="[605,651,1539,1566]" captionStart="Figs 156-163" captionStartId="52.[140,184,1590,1612]" captionTargetBox="[148,1107,237,1505]" captionTargetId="figure@52.[140,1108,237,1527]" captionTargetPageId="52" captionText="Figs 156-163. Genital structures of Gnypeta canadensis: 156, median lobe of the aedeagus in lateral view, 157, ventral view; 158, paramere; 159, male tergite 8; 160, male sternite 8; 161, spermatheca; 162, female tergite 8; 163, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774124" httpUri="https://zenodo.org/record/3774124/files/figure.png" pageId="4" pageNumber="15">157</figureCitation>
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<figureCitation box="[663,709,1539,1566]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="4" pageNumber="15">165</figureCitation>
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<figureCitation box="[721,767,1539,1565]" captionStart="Figs 172-180" captionStartId="56.[140,184,1590,1612]" captionTargetBox="[142,1108,243,1519]" captionTargetId="figure@56.[140,1108,242,1532]" captionTargetPageId="56" captionText="Figs 172-180. Genital structures of Gnypeta saccharina: 172, median lobe of the aedeagus in lateral view, 173, dorsal view, 174, apical part of median lobe in ventral view; 175, paramere; 176, male tergite 8; 177, male sternite 8; 178, spermatheca; 179, female tergite 8; 180, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774128" httpUri="https://zenodo.org/record/3774128/files/figure.png" pageId="4" pageNumber="15">173</figureCitation>
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<figureCitation box="[779,826,1539,1565]" captionStart="Figs 181-189" captionStartId="59.[140,184,1590,1612]" captionTargetBox="[152,1108,234,1496]" captionTargetId="figure@59.[140,1109,234,1545]" captionTargetPageId="59" captionText="Figs 181-189. Genital structures of Gnypeta minuta: 181, median lobe of the aedeagus in lateral view, 182, dorsal view; 183, paramere; 184, male tergite 8; 185, male sternite 8; 186, 187, spermatheca; 188, female tergite 8; 189, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774130" httpUri="https://zenodo.org/record/3774130/files/figure.png" pageId="4" pageNumber="15">182</figureCitation>
), crista apicalis of bulbus large and subtriangular in shape laterally (
<figureCitation box="[621,709,1574,1601]" captionStart="Figs 39-47" captionStartId="17.[140,184,1590,1612]" captionTargetBox="[140,1101,234,1493]" captionTargetId="figure@17.[140,1108,227,1517]" captionTargetPageId="17" captionText="Figs 39-47. Genital structures of Gnypeta ashei: 39, median lobe of the aedeagus in lateral view, 40, dorsal view, 41, ventral view; 42, paramere; 43, male tergite 8; 44, male sternite 8; 45, spermatheca; 46, female tergite 8; 47, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774096" httpUri="https://zenodo.org/record/3774096/files/figure.png" pageId="4" pageNumber="15">Figs 39</figureCitation>
,
<figureCitation box="[723,755,1575,1601]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="4" pageNumber="15">48</figureCitation>
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<figureCitation box="[768,800,1574,1601]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="4" pageNumber="15">57</figureCitation>
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<figureCitation box="[814,846,1574,1601]" captionStart-0="Figs 48-56" captionStart-1="Figs 181-189" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStart-6="Fig" captionStart-7="Fig" captionStart-8="Fig" captionStart-9="Fig" captionStartId-0="20.[140,184,1590,1612]" captionStartId-1="59.[140,184,1590,1612]" captionStartId-2="60.[140,174,1494,1516]" captionStartId-3="60.[140,174,1558,1580]" captionStartId-4="60.[140,174,1622,1644]" captionStartId-5="60.[140,174,1654,1676]" captionStartId-6="61.[140,174,883,905]" captionStartId-7="61.[140,174,915,937]" captionStartId-8="61.[140,174,947,969]" captionStartId-9="61.[140,174,979,1001]" captionTargetBox-0="[152,1097,238,1512]" captionTargetBox-1="[152,1108,234,1496]" captionTargetBox-2="[140,1103,451,785]" captionTargetBox-3="[140,1106,794,1132]" captionTargetBox-4="[140,1101,1127,1467]" captionTargetBox-5="[140,1101,1127,1467]" captionTargetBox-6="[140,1106,165,502]" captionTargetBox-7="[140,1106,165,502]" captionTargetBox-8="[140,1106,515,855]" captionTargetBox-9="[140,1106,515,855]" captionTargetId-0="figure@20.[140,1108,238,1528]" captionTargetId-1="figure@59.[140,1109,234,1545]" captionTargetId-3="figure@60.[140,1108,447,1475]" captionTargetPageId-0="20" captionTargetPageId-1="59" captionTargetPageId-2="60" captionTargetPageId-3="60" captionTargetPageId-4="60" captionTargetPageId-5="60" captionTargetPageId-6="61" captionTargetPageId-7="61" captionTargetPageId-8="61" captionTargetPageId-9="61" captionText-0="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." captionText-1="Figs 181-189. Genital structures of Gnypeta minuta: 181, median lobe of the aedeagus in lateral view, 182, dorsal view; 183, paramere; 184, male tergite 8; 185, male sternite 8; 186, 187, spermatheca; 188, female tergite 8; 189, female sternite 8." captionText-2="Fig. 190. Collection localities in Canada and Alaska of G. ashei." captionText-3="Fig. 192. Collection localities in Canada of G. atrolucens and G. nigrella." captionText-4="Fig. 194. Collection localities in Canada of G. crebrepunctata, G. dentata, and G. helenae." captionText-5="Fig. 195. Collection localities in Canada and Alaska of G. groenlandica." captionText-6="Fig. 196. Collection localities in Canada of G. brevicornis and G. uteana." captionText-7="Fig. 197. Collection localities in Canada and Alaska of G. caerulea." captionText-8="Fig. 198. Collection localities in Canada and Alaska of G. lohsei and G. canadiensis." captionText-9="Fig. 199. Collection localities in Canada and Alaska of G. carbonaria and G. saccharina." figureDoi-0="http://doi.org/10.5281/zenodo.3774098" figureDoi-1="http://doi.org/10.5281/zenodo.3774130" figureDoi-2="http://doi.org/10.5281/zenodo.3774132" figureDoi-3="http://doi.org/10.5281/zenodo.3774138" figureDoi-4="http://doi.org/10.5281/zenodo.3774142" figureDoi-5="http://doi.org/10.5281/zenodo.3774144" figureDoi-6="http://doi.org/10.5281/zenodo.3774146" figureDoi-7="http://doi.org/10.5281/zenodo.3774148" figureDoi-8="http://doi.org/10.5281/zenodo.3774150" figureDoi-9="http://doi.org/10.5281/zenodo.3774152" httpUri-0="https://zenodo.org/record/3774098/files/figure.png" httpUri-1="https://zenodo.org/record/3774130/files/figure.png" httpUri-2="https://zenodo.org/record/3774132/files/figure.png" httpUri-3="https://zenodo.org/record/3774138/files/figure.png" httpUri-4="https://zenodo.org/record/3774142/files/figure.png" httpUri-5="https://zenodo.org/record/3774144/files/figure.png" httpUri-6="https://zenodo.org/record/3774146/files/figure.png" httpUri-7="https://zenodo.org/record/3774148/files/figure.png" httpUri-8="https://zenodo.org/record/3774150/files/figure.png" httpUri-9="https://zenodo.org/record/3774152/files/figure.png" pageId="4" pageNumber="15">56</figureCitation>
); paramere broad with short apical lobe bearing three long subapical and one short apical macrosetae (
<figureCitation box="[1018,1103,1609,1636]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="4" pageNumber="15">Figs 60</figureCitation>
,
<figureCitation box="[140,172,1645,1671]" captionStart="Figs 75-82" captionStartId="27.[140,184,1590,1612]" captionTargetBox="[150,1098,227,1521]" captionTargetId="figure@27.[136,1110,216,1540]" captionTargetPageId="27" captionText="Figs 75-82. Genital structures of Gnypeta dentata: 75, median lobe of the aedeagus in lateral view, 76, dorsal view; 77, paramere; 78, male tergite 8; 79, male sternite 8; 80, spermatheca; 81, female tergite 8; 82, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774104" httpUri="https://zenodo.org/record/3774104/files/figure.png" pageId="4" pageNumber="15">77</figureCitation>
,
<figureCitation box="[182,214,1645,1672]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="4" pageNumber="15">85</figureCitation>
,
<figureCitation box="[225,272,1645,1671]" captionStart="Figs 117-124" captionStartId="39.[140,184,1590,1612]" captionTargetBox="[140,1108,219,1480]" captionTargetId="figure@39.[140,1108,219,1510]" captionTargetPageId="39" captionText="Figs 117-124. Genital structures of Gnypeta crebrepunctata: 117, median lobe of the aedeagus in lateral view, 118, dorsal view; 119, paramere; 120, male tergite 8; 121, male sternite 8; 122, spermatheca; 123, female tergite 8; 124, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774114" httpUri="https://zenodo.org/record/3774114/files/figure.png" pageId="4" pageNumber="15">119</figureCitation>
,
<figureCitation box="[282,329,1645,1671]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="4" pageNumber="15">142</figureCitation>
,
<figureCitation box="[339,386,1645,1672]" captionStart="Figs 148-155" captionStartId="49.[140,184,1590,1612]" captionTargetBox="[142,1103,241,1513]" captionTargetId="figure@49.[140,1111,225,1529]" captionTargetPageId="49" captionText="Figs 148-155. Genital structures of Gnypeta helenae: 148, median lobe of the aedeagus in lateral view, 149, dorsal view; 150, paramere; 151, male tergite 8; 152, male sternite 8; 153, spermatheca; 154, female tergite 8; 155, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774122" httpUri="https://zenodo.org/record/3774122/files/figure.png" pageId="4" pageNumber="15">150</figureCitation>
); male tergite 8 truncate, or rarely emarginated apically, its apical
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3774051" ID-Zenodo-Dep="3774051" box="[140,740,1654,1676]" httpUri="https://zenodo.org/record/3774051/files/figure.png" pageId="5" pageNumber="16" startId="5.[140,174,1654,1676]" targetBox="[243,974,176,1625]" targetPageId="5">
<paragraph blockId="5.[140,740,1654,1676]" box="[140,740,1654,1676]" pageId="5" pageNumber="16">
<emphasis bold="true" box="[140,202,1654,1676]" pageId="5" pageNumber="16">Fig. 1.</emphasis>
<taxonomicName authority="(LeConte)" baseAuthorityName="LeConte" box="[209,459,1654,1676]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="16" phylum="Arthropoda" rank="species" species="nigrella">
<emphasis box="[209,354,1654,1676]" italics="true" pageId="5" pageNumber="16">Gnypeta nigrella</emphasis>
(LeConte)
</taxonomicName>
, in dorsal view. Scale =
<quantity box="[677,736,1654,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="16" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3770346" ID-Zenodo-Dep="3770346" box="[140,750,1654,1676]" httpUri="https://zenodo.org/record/3770346/files/figure.png" pageId="6" pageNumber="17" startId="6.[140,174,1654,1676]" targetBox="[197,1035,184,1604]" targetPageId="6">
<paragraph blockId="6.[140,750,1654,1676]" box="[140,750,1654,1676]" pageId="6" pageNumber="17">
<emphasis bold="true" box="[140,205,1654,1676]" pageId="6" pageNumber="17">Fig. 2.</emphasis>
<taxonomicName authority="(LeConte)" baseAuthorityName="LeConte" box="[210,461,1654,1676]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="17" phylum="Arthropoda" rank="species" species="nigrella">
<emphasis box="[210,284,1655,1676]" italics="true" pageId="6" pageNumber="17">Gnypeta</emphasis>
<emphasis box="[290,356,1654,1675]" italics="true" pageId="6" pageNumber="17">nigrella</emphasis>
(LeConte)
</taxonomicName>
, in ventral view. Scale = 1 mm.
</paragraph>
</caption>
<paragraph blockId="7.[140,1108,166,369]" pageId="7" pageNumber="18">
margin smooth or with small 2-4 dents (
<figureCitation box="[588,673,166,193]" captionStart="Figs 39-47" captionStartId="17.[140,184,1590,1612]" captionTargetBox="[140,1101,234,1493]" captionTargetId="figure@17.[140,1108,227,1517]" captionTargetPageId="17" captionText="Figs 39-47. Genital structures of Gnypeta ashei: 39, median lobe of the aedeagus in lateral view, 40, dorsal view, 41, ventral view; 42, paramere; 43, male tergite 8; 44, male sternite 8; 45, spermatheca; 46, female tergite 8; 47, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774096" httpUri="https://zenodo.org/record/3774096/files/figure.png" pageId="7" pageNumber="18">Figs 43</figureCitation>
,
<figureCitation box="[684,716,166,193]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="7" pageNumber="18">52</figureCitation>
,
<figureCitation box="[727,759,166,192]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="7" pageNumber="18">61</figureCitation>
,
<figureCitation box="[769,801,167,193]" captionStart="Figs 75-82" captionStartId="27.[140,184,1590,1612]" captionTargetBox="[150,1098,227,1521]" captionTargetId="figure@27.[136,1110,216,1540]" captionTargetPageId="27" captionText="Figs 75-82. Genital structures of Gnypeta dentata: 75, median lobe of the aedeagus in lateral view, 76, dorsal view; 77, paramere; 78, male tergite 8; 79, male sternite 8; 80, spermatheca; 81, female tergite 8; 82, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774104" httpUri="https://zenodo.org/record/3774104/files/figure.png" pageId="7" pageNumber="18">78</figureCitation>
,
<figureCitation box="[812,858,167,193]" captionStart="Figs 99-107" captionStartId="33.[140,184,1590,1612]" captionTargetBox="[147,1102,243,1529]" captionTargetId="figure@33.[137,1111,233,1559]" captionTargetPageId="33" captionText="Figs 99-107. Genital structures of Gnypeta brincki: 99, median lobe of the aedeagus in lateral view, 100, dorsal view, 101, apical part of median lobe in ventral view; 102, paramere; 103, male tergite 8; 104, male sternite 8; 105, spermatheca; 106, female tergite 8; 107, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774110" httpUri="https://zenodo.org/record/3774110/files/figure.png" pageId="7" pageNumber="18">103</figureCitation>
,
<figureCitation box="[869,916,167,193]" captionStart="Figs 108-116" captionStartId="36.[140,184,1590,1612]" captionTargetBox="[144,1104,247,1505]" captionTargetId="figure@36.[143,1111,230,1520]" captionTargetPageId="36" captionText="Figs 108-116. Genital structures of Gnypeta lohsei: 108, median lobe of the aedeagus in lateral view, 109, dorsal view, 110, apical part of median lobe in ventral view; 111, paramere; 112, male tergite 8; 113, male sternite 8; 114, spermatheca; 115, female tergite 8; 116, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774112" httpUri="https://zenodo.org/record/3774112/files/figure.png" pageId="7" pageNumber="18">112</figureCitation>
,
<figureCitation box="[927,974,166,193]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="7" pageNumber="18">167</figureCitation>
); spermatheca of four
<typeStatus box="[270,332,202,228]" pageId="7" pageNumber="18">types</typeStatus>
: S-shaped (
<figureCitation box="[461,562,201,228]" captionStart="Figs 99-107" captionStartId="33.[140,184,1590,1612]" captionTargetBox="[147,1102,243,1529]" captionTargetId="figure@33.[137,1111,233,1559]" captionTargetPageId="33" captionText="Figs 99-107. Genital structures of Gnypeta brincki: 99, median lobe of the aedeagus in lateral view, 100, dorsal view, 101, apical part of median lobe in ventral view; 102, paramere; 103, male tergite 8; 104, male sternite 8; 105, spermatheca; 106, female tergite 8; 107, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774110" httpUri="https://zenodo.org/record/3774110/files/figure.png" pageId="7" pageNumber="18">Figs 105</figureCitation>
,
<figureCitation box="[576,623,202,228]" captionStart="Figs 108-116" captionStartId="36.[140,184,1590,1612]" captionTargetBox="[144,1104,247,1505]" captionTargetId="figure@36.[143,1111,230,1520]" captionTargetPageId="36" captionText="Figs 108-116. Genital structures of Gnypeta lohsei: 108, median lobe of the aedeagus in lateral view, 109, dorsal view, 110, apical part of median lobe in ventral view; 111, paramere; 112, male tergite 8; 113, male sternite 8; 114, spermatheca; 115, female tergite 8; 116, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774112" httpUri="https://zenodo.org/record/3774112/files/figure.png" pageId="7" pageNumber="18">114</figureCitation>
,
<figureCitation box="[636,682,201,228]" captionStart="Figs 148-155" captionStartId="49.[140,184,1590,1612]" captionTargetBox="[142,1103,241,1513]" captionTargetId="figure@49.[140,1111,225,1529]" captionTargetPageId="49" captionText="Figs 148-155. Genital structures of Gnypeta helenae: 148, median lobe of the aedeagus in lateral view, 149, dorsal view; 150, paramere; 151, male tergite 8; 152, male sternite 8; 153, spermatheca; 154, female tergite 8; 155, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774122" httpUri="https://zenodo.org/record/3774122/files/figure.png" pageId="7" pageNumber="18">153</figureCitation>
,
<figureCitation box="[695,741,201,228]" captionStart="Figs 156-163" captionStartId="52.[140,184,1590,1612]" captionTargetBox="[148,1107,237,1505]" captionTargetId="figure@52.[140,1108,237,1527]" captionTargetPageId="52" captionText="Figs 156-163. Genital structures of Gnypeta canadensis: 156, median lobe of the aedeagus in lateral view, 157, ventral view; 158, paramere; 159, male tergite 8; 160, male sternite 8; 161, spermatheca; 162, female tergite 8; 163, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774124" httpUri="https://zenodo.org/record/3774124/files/figure.png" pageId="7" pageNumber="18">161</figureCitation>
), C-shaped (
<figureCitation box="[888,990,202,228]" captionStart="Figs 117-124" captionStartId="39.[140,184,1590,1612]" captionTargetBox="[140,1108,219,1480]" captionTargetId="figure@39.[140,1108,219,1510]" captionTargetPageId="39" captionText="Figs 117-124. Genital structures of Gnypeta crebrepunctata: 117, median lobe of the aedeagus in lateral view, 118, dorsal view; 119, paramere; 120, male tergite 8; 121, male sternite 8; 122, spermatheca; 123, female tergite 8; 124, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774114" httpUri="https://zenodo.org/record/3774114/files/figure.png" pageId="7" pageNumber="18">Figs 122</figureCitation>
,
<figureCitation box="[1003,1103,202,228]" captionStart="Figs 125-129" captionStartId="41.[140,184,1622,1644]" captionTargetBox="[151,1097,842,1571]" captionTargetId="figure@41.[140,1110,828,1617]" captionTargetPageId="41" captionText="Figs 125-129. Genital structures of Gnypeta manitobae (lectotype): 125, female tergite 8; 126, female sternite 8; 127-129, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3774116" httpUri="https://zenodo.org/record/3774116/files/figure.png" pageId="7" pageNumber="18">127-129</figureCitation>
,
<figureCitation box="[140,187,237,264]" captionStart="Figs 130-138" captionStartId="44.[140,184,1590,1612]" captionTargetBox="[140,1097,230,1503]" captionTargetId="figure@44.[140,1108,228,1520]" captionTargetPageId="44" captionText="Figs 130-138. Genital structures of Gnypeta uteana: 130, median lobe of the aedeagus in lateral view, 131, dorsal view, 132, ventral view; 133, paramere; 134, male tergite 8; 135, male sternite 8; 136, spermatheca; 137, female tergite 8; 138, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774118" httpUri="https://zenodo.org/record/3774118/files/figure.png" pageId="7" pageNumber="18">136</figureCitation>
), hatchet-shaped (
<figureCitation box="[393,479,237,264]" captionStart="Figs 39-47" captionStartId="17.[140,184,1590,1612]" captionTargetBox="[140,1101,234,1493]" captionTargetId="figure@17.[140,1108,227,1517]" captionTargetPageId="17" captionText="Figs 39-47. Genital structures of Gnypeta ashei: 39, median lobe of the aedeagus in lateral view, 40, dorsal view, 41, ventral view; 42, paramere; 43, male tergite 8; 44, male sternite 8; 45, spermatheca; 46, female tergite 8; 47, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774096" httpUri="https://zenodo.org/record/3774096/files/figure.png" pageId="7" pageNumber="18">Figs 45</figureCitation>
,
<figureCitation box="[490,522,237,264]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="7" pageNumber="18">54</figureCitation>
,
<figureCitation box="[534,566,237,263]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="7" pageNumber="18">63</figureCitation>
,
<figureCitation box="[578,610,237,263]" captionStart="Figs 66-74" captionStartId="24.[140,184,1590,1612]" captionTargetBox="[140,1088,235,1499]" captionTargetId="figure@24.[140,1108,233,1523]" captionTargetPageId="24" captionText="Figs 66-74. Genital structures of Gnypeta sellmani: 66, median lobe of the aedeagus in lateral view, 67, dorsal view, 68, apical part of median lobe in ventral view; 69, paramere; 70, male tergite 8; 71, male sternite 8; 72, spermatheca; 73, female tergite 8; 74, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774102" httpUri="https://zenodo.org/record/3774102/files/figure.png" pageId="7" pageNumber="18">72</figureCitation>
,
<figureCitation box="[622,654,237,263]" captionStart="Figs 75-82" captionStartId="27.[140,184,1590,1612]" captionTargetBox="[150,1098,227,1521]" captionTargetId="figure@27.[136,1110,216,1540]" captionTargetPageId="27" captionText="Figs 75-82. Genital structures of Gnypeta dentata: 75, median lobe of the aedeagus in lateral view, 76, dorsal view; 77, paramere; 78, male tergite 8; 79, male sternite 8; 80, spermatheca; 81, female tergite 8; 82, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774104" httpUri="https://zenodo.org/record/3774104/files/figure.png" pageId="7" pageNumber="18">80</figureCitation>
,
<figureCitation box="[665,697,237,263]" captionStart="Figs 83-90" captionStartId="29.[140,184,1590,1612]" captionTargetBox="[148,1105,214,1510]" captionTargetId="figure@29.[137,1110,198,1540]" captionTargetPageId="29" captionText="Figs 83-90. Genital structures of Gnypeta groenlandica: 83, median lobe of the aedeagus in lateral view, 84, dorsal view; 85, paramere; 86, male tergite 8; 87, male sternite 8; 88, spermatheca; 89, female tergite 8; 90, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774106" httpUri="https://zenodo.org/record/3774106/files/figure.png" pageId="7" pageNumber="18">88</figureCitation>
,
<figureCitation box="[709,741,237,264]" captionStart-0="Figs 91-98" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStartId-0="31.[140,184,1590,1612]" captionStartId-1="61.[140,174,883,905]" captionStartId-2="61.[140,174,915,937]" captionStartId-3="61.[140,174,947,969]" captionStartId-4="61.[140,174,979,1001]" captionTargetBox-0="[151,1101,241,1488]" captionTargetBox-1="[140,1106,165,502]" captionTargetBox-2="[140,1106,165,502]" captionTargetBox-3="[140,1106,515,855]" captionTargetBox-4="[140,1106,515,855]" captionTargetId-0="figure@31.[130,1115,227,1529]" captionTargetPageId-0="31" captionTargetPageId-1="61" captionTargetPageId-2="61" captionTargetPageId-3="61" captionTargetPageId-4="61" captionText-0="Figs 91-98. Genital structures of Gnypeta brevicornis: 91, median lobe of the aedeagus in lateral view, 92, ventral view; 93, paramere; 94, male tergite 8; 95, male sternite 8; 96, spermatheca; 97, female tergite 8; 98, female sternite 8." captionText-1="Fig. 196. Collection localities in Canada of G. brevicornis and G. uteana." captionText-2="Fig. 197. Collection localities in Canada and Alaska of G. caerulea." captionText-3="Fig. 198. Collection localities in Canada and Alaska of G. lohsei and G. canadiensis." captionText-4="Fig. 199. Collection localities in Canada and Alaska of G. carbonaria and G. saccharina." figureDoi-0="http://doi.org/10.5281/zenodo.3774108" figureDoi-1="http://doi.org/10.5281/zenodo.3774146" figureDoi-2="http://doi.org/10.5281/zenodo.3774148" figureDoi-3="http://doi.org/10.5281/zenodo.3774150" figureDoi-4="http://doi.org/10.5281/zenodo.3774152" httpUri-0="https://zenodo.org/record/3774108/files/figure.png" httpUri-1="https://zenodo.org/record/3774146/files/figure.png" httpUri-2="https://zenodo.org/record/3774148/files/figure.png" httpUri-3="https://zenodo.org/record/3774150/files/figure.png" httpUri-4="https://zenodo.org/record/3774152/files/figure.png" pageId="7" pageNumber="18">96</figureCitation>
), or club-shaped (
<figureCitation box="[944,1045,237,263]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="7" pageNumber="18">Figs 169</figureCitation>
,
<figureCitation box="[1056,1102,237,263]" captionStart="Figs 172-180" captionStartId="56.[140,184,1590,1612]" captionTargetBox="[142,1108,243,1519]" captionTargetId="figure@56.[140,1108,242,1532]" captionTargetPageId="56" captionText="Figs 172-180. Genital structures of Gnypeta saccharina: 172, median lobe of the aedeagus in lateral view, 173, dorsal view, 174, apical part of median lobe in ventral view; 175, paramere; 176, male tergite 8; 177, male sternite 8; 178, spermatheca; 179, female tergite 8; 180, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774128" httpUri="https://zenodo.org/record/3774128/files/figure.png" pageId="7" pageNumber="18">178</figureCitation>
,
<figureCitation box="[140,248,272,298]" captionStart="Figs 181-189" captionStartId="59.[140,184,1590,1612]" captionTargetBox="[152,1108,234,1496]" captionTargetId="figure@59.[140,1109,234,1545]" captionTargetPageId="59" captionText="Figs 181-189. Genital structures of Gnypeta minuta: 181, median lobe of the aedeagus in lateral view, 182, dorsal view; 183, paramere; 184, male tergite 8; 185, male sternite 8; 186, 187, spermatheca; 188, female tergite 8; 189, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774130" httpUri="https://zenodo.org/record/3774130/files/figure.png" pageId="7" pageNumber="18">186, 187</figureCitation>
); capsule tubular with apical part approximately spherical (
<figureCitation box="[937,1041,272,299]" captionStart="Figs 139-147" captionStartId="46.[140,184,1590,1612]" captionTargetBox="[152,1103,246,1521]" captionTargetId="figure@46.[140,1108,249,1540]" captionTargetPageId="46" captionText="Figs 139-147. Genital structures of Gnypeta brincki: 139, median lobe of the aedeagus in lateral view, 140, dorsal view, 141, apical part of median lobe in ventral view; 142, paramere; 143, male tergite 8; 144, male sternite 8; 145, spermatheca; 146, female tergite 8; 147, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774120" httpUri="https://zenodo.org/record/3774120/files/figure.png" pageId="7" pageNumber="18">Figs 145</figureCitation>
,
<figureCitation box="[1056,1103,272,298]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="7" pageNumber="18">169</figureCitation>
,
<figureCitation box="[140,186,307,333]" captionStart="Figs 172-180" captionStartId="56.[140,184,1590,1612]" captionTargetBox="[142,1108,243,1519]" captionTargetId="figure@56.[140,1108,242,1532]" captionTargetPageId="56" captionText="Figs 172-180. Genital structures of Gnypeta saccharina: 172, median lobe of the aedeagus in lateral view, 173, dorsal view, 174, apical part of median lobe in ventral view; 175, paramere; 176, male tergite 8; 177, male sternite 8; 178, spermatheca; 179, female tergite 8; 180, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774128" httpUri="https://zenodo.org/record/3774128/files/figure.png" pageId="7" pageNumber="18">178</figureCitation>
,
<figureCitation box="[203,313,307,333]" captionStart="Figs 181-189" captionStartId="59.[140,184,1590,1612]" captionTargetBox="[152,1108,234,1496]" captionTargetId="figure@59.[140,1109,234,1545]" captionTargetPageId="59" captionText="Figs 181-189. Genital structures of Gnypeta minuta: 181, median lobe of the aedeagus in lateral view, 182, dorsal view; 183, paramere; 184, male tergite 8; 185, male sternite 8; 186, 187, spermatheca; 188, female tergite 8; 189, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774130" httpUri="https://zenodo.org/record/3774130/files/figure.png" pageId="7" pageNumber="18">186, 187</figureCitation>
), tubular (
<figureCitation box="[440,546,307,333]" captionStart="Figs 117-124" captionStartId="39.[140,184,1590,1612]" captionTargetBox="[140,1108,219,1480]" captionTargetId="figure@39.[140,1108,219,1510]" captionTargetPageId="39" captionText="Figs 117-124. Genital structures of Gnypeta crebrepunctata: 117, median lobe of the aedeagus in lateral view, 118, dorsal view; 119, paramere; 120, male tergite 8; 121, male sternite 8; 122, spermatheca; 123, female tergite 8; 124, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774114" httpUri="https://zenodo.org/record/3774114/files/figure.png" pageId="7" pageNumber="18">Figs 122</figureCitation>
,
<figureCitation box="[562,609,307,334]" captionStart="Figs 125-129" captionStartId="41.[140,184,1622,1644]" captionTargetBox="[151,1097,842,1571]" captionTargetId="figure@41.[140,1110,828,1617]" captionTargetPageId="41" captionText="Figs 125-129. Genital structures of Gnypeta manitobae (lectotype): 125, female tergite 8; 126, female sternite 8; 127-129, spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3774116" httpUri="https://zenodo.org/record/3774116/files/figure.png" pageId="7" pageNumber="18">127</figureCitation>
,
<figureCitation box="[613,659,307,334]" captionStart="Figs 148-155" captionStartId="49.[140,184,1590,1612]" captionTargetBox="[142,1103,241,1513]" captionTargetId="figure@49.[140,1111,225,1529]" captionTargetPageId="49" captionText="Figs 148-155. Genital structures of Gnypeta helenae: 148, median lobe of the aedeagus in lateral view, 149, dorsal view; 150, paramere; 151, male tergite 8; 152, male sternite 8; 153, spermatheca; 154, female tergite 8; 155, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774122" httpUri="https://zenodo.org/record/3774122/files/figure.png" pageId="7" pageNumber="18">153</figureCitation>
,
<figureCitation box="[676,722,307,334]" captionStart="Figs 156-163" captionStartId="52.[140,184,1590,1612]" captionTargetBox="[148,1107,237,1505]" captionTargetId="figure@52.[140,1108,237,1527]" captionTargetPageId="52" captionText="Figs 156-163. Genital structures of Gnypeta canadensis: 156, median lobe of the aedeagus in lateral view, 157, ventral view; 158, paramere; 159, male tergite 8; 160, male sternite 8; 161, spermatheca; 162, female tergite 8; 163, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774124" httpUri="https://zenodo.org/record/3774124/files/figure.png" pageId="7" pageNumber="18">161</figureCitation>
), or funnel-shaped (
<figureCitation box="[964,1054,307,334]" captionStart="Figs 39-47" captionStartId="17.[140,184,1590,1612]" captionTargetBox="[140,1101,234,1493]" captionTargetId="figure@17.[140,1108,227,1517]" captionTargetPageId="17" captionText="Figs 39-47. Genital structures of Gnypeta ashei: 39, median lobe of the aedeagus in lateral view, 40, dorsal view, 41, ventral view; 42, paramere; 43, male tergite 8; 44, male sternite 8; 45, spermatheca; 46, female tergite 8; 47, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774096" httpUri="https://zenodo.org/record/3774096/files/figure.png" pageId="7" pageNumber="18">Figs 45</figureCitation>
,
<figureCitation box="[1071,1103,307,334]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="7" pageNumber="18">54</figureCitation>
,
<figureCitation box="[140,172,342,368]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="7" pageNumber="18">63</figureCitation>
,
<figureCitation box="[186,219,343,369]" captionStart="Figs 66-74" captionStartId="24.[140,184,1590,1612]" captionTargetBox="[140,1088,235,1499]" captionTargetId="figure@24.[140,1108,233,1523]" captionTargetPageId="24" captionText="Figs 66-74. Genital structures of Gnypeta sellmani: 66, median lobe of the aedeagus in lateral view, 67, dorsal view, 68, apical part of median lobe in ventral view; 69, paramere; 70, male tergite 8; 71, male sternite 8; 72, spermatheca; 73, female tergite 8; 74, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774102" httpUri="https://zenodo.org/record/3774102/files/figure.png" pageId="7" pageNumber="18">72</figureCitation>
); stem tubular, elongate, more or less sinuate and moderately swollen basally.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3774053" ID-Zenodo-Dep="3774053" httpUri="https://zenodo.org/record/3774053/files/figure.png" pageId="7" pageNumber="18" startId="7.[140,184,1622,1644]" targetBox="[181,994,433,1595]" targetPageId="7">
<paragraph blockId="7.[140,1108,1622,1676]" pageId="7" pageNumber="18">
<emphasis bold="true" box="[140,231,1622,1644]" pageId="7" pageNumber="18">Figs 3-8.</emphasis>
<taxonomicName box="[238,312,1623,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis box="[238,312,1623,1644]" italics="true" pageId="7" pageNumber="18">Gnypeta</emphasis>
</taxonomicName>
species: 3,
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[415,487,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="ashei">
<emphasis box="[415,487,1622,1644]" italics="true" pageId="7" pageNumber="18">G. ashei</emphasis>
</taxonomicName>
; 4,
<taxonomicName authorityName="Palm" authorityYear="1966" box="[523,616,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="brincki">
<emphasis box="[523,616,1622,1644]" italics="true" pageId="7" pageNumber="18">G. brincki</emphasis>
</taxonomicName>
; 5,
<taxonomicName authorityName="Casey" authorityYear="1893" box="[652,768,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="atrolucens">
<emphasis box="[652,768,1622,1644]" italics="true" pageId="7" pageNumber="18">G. atrolucens</emphasis>
</taxonomicName>
; 6,
<taxonomicName box="[804,906,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="sellmani">
<emphasis box="[804,906,1622,1644]" italics="true" pageId="7" pageNumber="18">G. sellmani</emphasis>
</taxonomicName>
: 7,
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[942,1037,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="dentata">
<emphasis box="[942,1037,1622,1644]" italics="true" pageId="7" pageNumber="18">G. dentata</emphasis>
</taxonomicName>
; and 8,
<taxonomicName authorityName="Lohse" authorityYear="1989" box="[140,282,1654,1676]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="18" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis box="[140,282,1654,1676]" italics="true" pageId="7" pageNumber="18">G. groenlandica</emphasis>
</taxonomicName>
. Scale =
<quantity box="[364,420,1654,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="18" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
</caption>
<subSection pageId="8" pageNumber="19" type="multiple">
<paragraph blockId="8.[140,1108,166,369]" pageId="8" pageNumber="19">
Sternite 8 of male and female with broad space between basal margin and antecostal suture (
<figureCitation box="[224,353,202,228]" captionStart="Figs 39-47" captionStartId="17.[140,184,1590,1612]" captionTargetBox="[140,1101,234,1493]" captionTargetId="figure@17.[140,1108,227,1517]" captionTargetPageId="17" captionText="Figs 39-47. Genital structures of Gnypeta ashei: 39, median lobe of the aedeagus in lateral view, 40, dorsal view, 41, ventral view; 42, paramere; 43, male tergite 8; 44, male sternite 8; 45, spermatheca; 46, female tergite 8; 47, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774096" httpUri="https://zenodo.org/record/3774096/files/figure.png" pageId="8" pageNumber="19">Figs 44, 47</figureCitation>
,
<figureCitation box="[365,440,201,228]" captionStart="Figs 48-56" captionStartId="20.[140,184,1590,1612]" captionTargetBox="[152,1097,238,1512]" captionTargetId="figure@20.[140,1108,238,1528]" captionTargetPageId="20" captionText="Figs 48-56. Genital structures of Gnypeta brincki: 48, median lobe of the aedeagus in lateral view, 49, dorsal view, 50, apical part of median lobe in ventral view; 51, paramere; 52, male tergite 8; 53, male sternite 8; 54, spermatheca; 55, female tergite 8; 56, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774098" httpUri="https://zenodo.org/record/3774098/files/figure.png" pageId="8" pageNumber="19">53, 56</figureCitation>
,
<figureCitation box="[452,528,201,228]" captionStart="Figs 57-65" captionStartId="22.[140,184,1590,1612]" captionTargetBox="[152,1098,247,1512]" captionTargetId="figure@22.[140,1108,237,1527]" captionTargetPageId="22" captionText="Figs 57-65. Genital structures of Gnypeta atrolucens: 57, median lobe of the aedeagus in lateral view, 58, dorsal view, 59, apical part of median lobe in ventral view; 60, paramere; 61, male tergite 8; 62, male sternite 8; 63, spermatheca; 64, female tergite 8; 65, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774100" httpUri="https://zenodo.org/record/3774100/files/figure.png" pageId="8" pageNumber="19">62, 65</figureCitation>
).
<taxonomicName box="[548,638,202,228]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis box="[548,638,202,228]" italics="true" pageId="8" pageNumber="19">Gnypeta</emphasis>
</taxonomicName>
is readily distinguished from most genera of
<taxonomicName authorityName="Thomson" authorityYear="1859" box="[171,294,237,263]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="tribe" tribe="Oxypodini">Oxypodini</taxonomicName>
by having 4, 5, 5-articulated tarsi and from closely related
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[974,1072,237,263]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis box="[974,1072,237,263]" italics="true" pageId="8" pageNumber="19">Tachyusa</emphasis>
</taxonomicName>
by robust body (slender in
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[413,511,272,298]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis box="[413,511,272,298]" italics="true" pageId="8" pageNumber="19">Tachyusa</emphasis>
</taxonomicName>
), abdomen at base as broad as elytra or only slightly narrower (much narrower in
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[474,572,307,333]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis box="[474,572,307,333]" italics="true" pageId="8" pageNumber="19">Tachyusa</emphasis>
</taxonomicName>
), lack of distinctive ridges in abdominal tergal impressions, and by less elongate basal article of metatarsus (usually shorter than the
</paragraph>
</subSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3774055" ID-Zenodo-Dep="3774055" httpUri="https://zenodo.org/record/3774055/files/figure.png" pageId="8" pageNumber="19" startId="8.[140,184,1622,1644]" targetBox="[237,1043,441,1595]" targetPageId="8">
<paragraph blockId="8.[140,1108,1622,1676]" pageId="8" pageNumber="19">
<emphasis bold="true" box="[140,245,1622,1644]" pageId="8" pageNumber="19">Figs 9-14.</emphasis>
<taxonomicName box="[251,325,1623,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis box="[251,325,1623,1644]" italics="true" pageId="8" pageNumber="19">Gnypeta</emphasis>
</taxonomicName>
species: 9,
<taxonomicName authorityName="Casey" authorityYear="1906" box="[430,554,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis box="[430,554,1622,1644]" italics="true" pageId="8" pageNumber="19">G. brevicornis</emphasis>
</taxonomicName>
; 10,
<taxonomicName baseAuthorityName="C. R. Sahlberg" baseAuthorityYear="1831" box="[603,704,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[603,704,1622,1644]" italics="true" pageId="8" pageNumber="19">G. caerulea</emphasis>
</taxonomicName>
; 11,
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[752,829,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="lohsei">
<emphasis box="[752,829,1622,1644]" italics="true" pageId="8" pageNumber="19">G. lohsei</emphasis>
</taxonomicName>
; 12,
<taxonomicName baseAuthorityName="Casey" baseAuthorityYear="1886" box="[878,1036,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="crebrepunctata">
<emphasis box="[878,1036,1622,1644]" italics="true" pageId="8" pageNumber="19">G. crebrepunctata</emphasis>
</taxonomicName>
; 13,
<taxonomicName authorityName="Casey" authorityYear="1906" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="manitobae">
<emphasis italics="true" pageId="8" pageNumber="19">G. manitobae</emphasis>
</taxonomicName>
[apical part of abdomen removed]; 14,
<taxonomicName box="[602,691,1655,1676]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="19" phylum="Arthropoda" rank="species" species="uteana">
<emphasis box="[602,691,1655,1676]" italics="true" pageId="8" pageNumber="19">G. uteana</emphasis>
</taxonomicName>
. Scale =
<quantity box="[774,830,1654,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="19" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
</caption>
<subSection pageId="9" pageNumber="20" type="multiple">
<paragraph blockId="9.[140,1108,166,369]" pageId="9" pageNumber="20">
two following articles combined).
<taxonomicName authority="Stephens" authorityName="Stephens, Pasnik" authorityYear="2006" box="[522,749,166,192]" class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis box="[522,642,166,192]" italics="true" pageId="9" pageNumber="20">Ischnopoda</emphasis>
Stephens
</taxonomicName>
is another related genus to
<taxonomicName class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="9" pageNumber="20">Gnypeta</emphasis>
</taxonomicName>
, which differs externally by parallel-sided body, and extremely elongate metatarsus, exceeding 4/5 length of metatibia, and with basal article at least as long as the two following articles combined (
<figureCitation box="[468,534,272,298]" captionStart="Fig" captionStartId="5.[140,174,1654,1676]" captionTargetBox="[243,974,176,1625]" captionTargetId="figure@5.[136,1110,161,1637]" captionTargetPageId="5" captionText="Fig. 1. Gnypeta nigrella (LeConte), in dorsal view. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3774051" httpUri="https://zenodo.org/record/3774051/files/figure.png" pageId="9" pageNumber="20">Fig. 1</figureCitation>
). According to
<bibRefCitation author="Pasnik" box="[712,870,272,298]" firstAuthor="Pasnik" pageId="9" pageNumber="20" pagination="1 - 54" refId="ref24022" refString="Pasnik G (2007) Revision and phylogenetic analysis of the African genus Ischnopoderona (Scheerpeltz, 1974) new status (Coleoptera, Staphylinidae, Aleocharinae). Zootaxa 1412: 1 - 54." type="journal article" year="2007">Paśnik (2007)</bibRefCitation>
the genus
<taxonomicName box="[993,1083,272,298]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis box="[993,1083,272,298]" italics="true" pageId="9" pageNumber="20">Gnypeta</emphasis>
</taxonomicName>
is closely related to
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[341,439,307,333]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis box="[341,439,307,333]" italics="true" pageId="9" pageNumber="20">Tachyusa</emphasis>
</taxonomicName>
,
<taxonomicName authority="(Scheerpeltz)" baseAuthorityName="Scheerpeltz" box="[456,777,307,334]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis box="[456,622,307,333]" italics="true" pageId="9" pageNumber="20">Ischnopoderona</emphasis>
(Scheerpeltz)
</taxonomicName>
and
<taxonomicName authorityName="Stephens, Pasnik" authorityYear="2006" box="[839,959,307,333]" class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis box="[839,959,307,333]" italics="true" pageId="9" pageNumber="20">Ischnopoda</emphasis>
</taxonomicName>
. The key for these four genera is provided by
<bibRefCitation author="Pasnik" box="[506,668,342,369]" firstAuthor="Pasnik" pageId="9" pageNumber="20" pagination="1 - 54" refId="ref24022" refString="Pasnik G (2007) Revision and phylogenetic analysis of the African genus Ischnopoderona (Scheerpeltz, 1974) new status (Coleoptera, Staphylinidae, Aleocharinae). Zootaxa 1412: 1 - 54." type="journal article" year="2007">Paśnik (2007)</bibRefCitation>
.
</paragraph>
</subSection>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3774057" ID-Zenodo-Dep="3774057" httpUri="https://zenodo.org/record/3774057/files/figure.png" pageId="9" pageNumber="20" startId="9.[140,184,1622,1644]" targetBox="[214,1020,442,1588]" targetPageId="9">
<paragraph blockId="9.[140,1108,1622,1676]" pageId="9" pageNumber="20">
<emphasis bold="true" box="[140,259,1622,1644]" pageId="9" pageNumber="20">Figs 15-20.</emphasis>
<taxonomicName box="[265,339,1623,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis box="[265,339,1623,1644]" italics="true" pageId="9" pageNumber="20">Gnypeta</emphasis>
</taxonomicName>
species: 15,
<taxonomicName authorityName="Mannerheim" authorityYear="1830" box="[455,580,1622,1644]" class="Insecta" family="Staphylinidae" genus="Bolitochara" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis box="[455,580,1622,1644]" italics="true" pageId="9" pageNumber="20">G. carbonaria</emphasis>
</taxonomicName>
; 16,
<taxonomicName authorityName=": Moore and Legner" authorityYear="1975" box="[629,723,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="helenae">
<emphasis box="[629,723,1622,1644]" italics="true" pageId="9" pageNumber="20">G. helenae</emphasis>
</taxonomicName>
; 17,
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[771,892,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="canadensis">
<emphasis box="[771,892,1622,1644]" italics="true" pageId="9" pageNumber="20">G. canadensis</emphasis>
</taxonomicName>
; 18,
<taxonomicName box="[941,1036,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="nigrella">
<emphasis box="[941,1036,1622,1644]" italics="true" pageId="9" pageNumber="20">G. nigrella</emphasis>
</taxonomicName>
; 19,
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="saccharina">
<emphasis italics="true" pageId="9" pageNumber="20">G. saccharina</emphasis>
</taxonomicName>
; and 20,
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[321,416,1655,1676]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="20" phylum="Arthropoda" rank="species" species="minuta">
<emphasis box="[321,416,1655,1676]" italics="true" pageId="9" pageNumber="20">G. minuta</emphasis>
</taxonomicName>
[18-20, apical part of abdomen removed]. Scale =
<quantity box="[886,942,1654,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="20" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.3774059" ID-Zenodo-Dep="3774059" httpUri="https://zenodo.org/record/3774059/files/figure.png" pageId="10" pageNumber="21" startId="10.[140,184,1462,1484]" targetBox="[230,1007,164,1440]" targetPageId="10">
<paragraph blockId="10.[140,1108,1462,1676]" pageId="10" pageNumber="21">
<emphasis bold="true" box="[140,260,1462,1484]" pageId="10" pageNumber="21">Figs 21-38.</emphasis>
Antennae of
<taxonomicName box="[388,462,1463,1484]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis box="[388,462,1463,1484]" italics="true" pageId="10" pageNumber="21">Gnypeta</emphasis>
</taxonomicName>
species:
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[544,617,1462,1484]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="ashei">
<emphasis box="[544,617,1462,1484]" italics="true" pageId="10" pageNumber="21">G. ashei</emphasis>
</taxonomicName>
: 21a. male, 21b. female;
<taxonomicName authorityName="Palm" authorityYear="1966" box="[856,950,1462,1484]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="brincki">
<emphasis box="[856,950,1462,1484]" italics="true" pageId="10" pageNumber="21">G. brincki</emphasis>
</taxonomicName>
: 22a. male, 22b. female;
<taxonomicName authorityName="Casey" authorityYear="1893" box="[213,330,1494,1516]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="atrolucens">
<emphasis box="[213,330,1494,1516]" italics="true" pageId="10" pageNumber="21">G. atrolucens</emphasis>
</taxonomicName>
: 23a. male, 23b. female;
<taxonomicName box="[565,667,1494,1516]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="sellmani">
<emphasis box="[565,667,1494,1516]" italics="true" pageId="10" pageNumber="21">G. sellmani</emphasis>
</taxonomicName>
: 24a. male, 24b. female;
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[903,999,1494,1516]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="dentata">
<emphasis box="[903,999,1494,1516]" italics="true" pageId="10" pageNumber="21">G. dentata</emphasis>
</taxonomicName>
: 25a. male, 25b. female;
<taxonomicName authorityName="Lohse" authorityYear="1989" box="[261,403,1526,1548]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="groenlandica">
<emphasis box="[261,403,1526,1548]" italics="true" pageId="10" pageNumber="21">G. groenlandica</emphasis>
</taxonomicName>
: 26a. male, 26b. female:
<taxonomicName authorityName="Casey" authorityYear="1906" box="[639,763,1526,1548]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="brevicornis">
<emphasis box="[639,763,1526,1548]" italics="true" pageId="10" pageNumber="21">G. brevicornis</emphasis>
</taxonomicName>
: 27a. male, 27b. female;
<taxonomicName baseAuthorityName="C. R. Sahlberg" baseAuthorityYear="1831" box="[1000,1101,1526,1548]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="caerulea">
<emphasis box="[1000,1101,1526,1548]" italics="true" pageId="10" pageNumber="21">G. caerulea</emphasis>
</taxonomicName>
: 28a. male, 28b. female;
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[367,445,1558,1580]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="lohsei">
<emphasis box="[367,445,1558,1580]" italics="true" pageId="10" pageNumber="21">G. lohsei</emphasis>
</taxonomicName>
: 29a. male, 29b. female;
<taxonomicName baseAuthorityName="Casey" baseAuthorityYear="1886" box="[685,844,1558,1580]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="crebrepunctata">
<emphasis box="[685,844,1558,1580]" italics="true" pageId="10" pageNumber="21">G. crebrepunctata</emphasis>
</taxonomicName>
: 30a. male, 30b. female;
<taxonomicName authorityName="Casey" authorityYear="1906" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="manitobae">
<emphasis italics="true" pageId="10" pageNumber="21">G. manitobae</emphasis>
</taxonomicName>
: 31b. female;
<taxonomicName box="[373,464,1591,1612]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="uteana">
<emphasis box="[373,464,1591,1612]" italics="true" pageId="10" pageNumber="21">G. uteana</emphasis>
</taxonomicName>
: 32a. male, 32b. female;
<taxonomicName authorityName="Mannerheim" authorityYear="1830" box="[711,838,1590,1612]" class="Insecta" family="Staphylinidae" genus="Bolitochara" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="carbonaria">
<emphasis box="[711,838,1590,1612]" italics="true" pageId="10" pageNumber="21">G. carbonaria</emphasis>
</taxonomicName>
: 33a. male, 33b. female;
<taxonomicName authorityName=": Moore and Legner" authorityYear="1975" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="helenae">
<emphasis italics="true" pageId="10" pageNumber="21">G. helenae</emphasis>
</taxonomicName>
: 34a. male, 34b. female;
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[436,556,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="canadensis">
<emphasis box="[436,556,1622,1644]" italics="true" pageId="10" pageNumber="21">G. canadensis</emphasis>
</taxonomicName>
: 35a. male, 35b. female;
<taxonomicName box="[787,882,1622,1644]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="nigrella">
<emphasis box="[787,882,1622,1644]" italics="true" pageId="10" pageNumber="21">G. nigrella</emphasis>
</taxonomicName>
: 36a. male, 36b. female;
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[140,262,1654,1676]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="saccharina">
<emphasis box="[140,262,1654,1676]" italics="true" pageId="10" pageNumber="21">G. saccharina</emphasis>
</taxonomicName>
: 37a. male, 37b. female;
<taxonomicName authorityName="Klimaszewski &amp; Savard &amp; Pelletier &amp; Webster" authorityYear="2008" box="[495,590,1655,1676]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="21" phylum="Arthropoda" rank="species" species="minuta">
<emphasis box="[495,590,1655,1676]" italics="true" pageId="10" pageNumber="21">G. minuta</emphasis>
</taxonomicName>
: 38a. male, 38b. female. Scale =
<quantity box="[894,950,1654,1676]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="21" unit="mm" value="1.0">1 mm</quantity>
</paragraph>
</caption>
<subSubSection pageId="11" pageNumber="22" type="biology_ecology">
<paragraph blockId="11.[140,1108,166,368]" box="[140,467,166,193]" pageId="11" pageNumber="22">
<emphasis bold="true" box="[140,467,166,193]" pageId="11" pageNumber="22">
<emphasis box="[140,265,166,192]" inLineHeading="true" pageId="11" pageNumber="22" reason="1">Collection</emphasis>
and habitat data
</emphasis>
</paragraph>
<paragraph blockId="11.[140,1108,166,368]" pageId="11" pageNumber="22">
Adults are associated with riparian habitats and debris along the margins of marshes, ponds, lakes, and streams (
<bibRefCitation author="Ashe" box="[437,563,237,263]" firstAuthor="Ashe" pageId="11" pageNumber="22" pagination="299 - 374" refId="ref22843" refString="Ashe JS (2001) Keys to the tribes and genera of Nearctic Aleocharinae. In: Arnett RH, Thomas MC (Eds) American Beetles. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press LLC, Boca Raton, Florida, 299 - 374." type="book chapter" year="2001">Ashe 2001</bibRefCitation>
). Some Canadian species were found in cold wet moss alongside streams. They may also occur in vegetation and litter along edges of streams, river, and lakes, in grass tussocks on mud flats, in gravel, wooded bogs, beaver lodges, and decaying fungi.
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="23" pageId="11" pageNumber="22" type="discussion">
<paragraph blockId="11.[140,1114,413,1671]" box="[140,429,413,439]" pageId="11" pageNumber="22">
<emphasis bold="true" box="[140,429,413,439]" pageId="11" pageNumber="22">
<emphasis box="[140,295,413,439]" inLineHeading="true" pageId="11" pageNumber="22" reason="1">Phylogenetic</emphasis>
affiliations
</emphasis>
</paragraph>
<paragraph blockId="11.[140,1114,413,1671]" lastBlockId="12.[140,1108,166,404]" lastPageId="12" lastPageNumber="23" pageId="11" pageNumber="22">
<bibRefCitation author="Seevers" box="[140,306,448,474]" firstAuthor="Seevers" pageId="11" pageNumber="22" pagination="1 - 289" refId="ref24091" refString="Seevers CH (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoology 71: i-vi, 1 - 289." type="journal article" year="1978">Seevers (1978)</bibRefCitation>
classified the genus
<taxonomicName box="[545,635,448,474]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[545,635,448,474]" italics="true" pageId="11" pageNumber="22">Gnypeta</emphasis>
</taxonomicName>
in the tribe
<taxonomicName authorityName="Thomson" authorityYear="1859" box="[786,909,448,474]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Oxypodini">Oxypodini</taxonomicName>
and the subtribe
<taxonomicName box="[140,255,483,509]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="subTribe" subTribe="Tachyusae">Tachyusae</taxonomicName>
(=Tachyusina Thomson), together with the genera
<taxonomicName authority="Erichson" authorityName="Erichson" authorityYear="1837" box="[840,1048,483,509]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[840,937,483,509]" italics="true" pageId="11" pageNumber="22">Tachyusa</emphasis>
Erichson
</taxonomicName>
,
<taxonomicName authority="Casey" authorityName="Casey" authorityYear="1906" class="Insecta" family="Staphylinidae" genus="Trachyota" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="11" pageNumber="22">Trachyota</emphasis>
Casey
</taxonomicName>
,
<taxonomicName authority="Casey" authorityName="Casey" authorityYear="1906" box="[294,444,518,545]" class="Insecta" family="Staphylinidae" genus="Teliusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[294,369,518,544]" italics="true" pageId="11" pageNumber="22">Teliusa</emphasis>
Casey
</taxonomicName>
,
<taxonomicName authority="Casey" authorityName="Casey" authorityYear="1906" box="[456,645,518,545]" class="Insecta" family="Staphylinidae" genus="Gnypetella" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[456,570,518,544]" italics="true" pageId="11" pageNumber="22">Gnypetella</emphasis>
Casey
</taxonomicName>
,
<taxonomicName authority="Sharp" authorityName="Sharp" authorityYear="1887" box="[656,836,518,544]" class="Insecta" family="Staphylinidae" genus="Meronera" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[656,759,519,544]" italics="true" pageId="11" pageNumber="22">Meronera</emphasis>
Sharp
</taxonomicName>
, and
<taxonomicName authority="Mulsant and Rey. He" authorityName="Mulsant and Rey. He" class="Insecta" family="Staphylinidae" genus="Brachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[898,1006,518,544]" italics="true" pageId="11" pageNumber="22">Brachyusa</emphasis>
Mulsant and Rey. He
</taxonomicName>
was tempted to consider
<taxonomicName box="[575,690,554,580]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="subTribe" subTribe="Tachyusae">Tachyusae</taxonomicName>
as a distinct tribe on the grounds of 4, 5, 5-articulated tarsi and the genital features but he was afraid that this arrangement would obscure the relationship of
<taxonomicName box="[538,628,624,650]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[538,628,624,650]" italics="true" pageId="11" pageNumber="22">Gnypeta</emphasis>
</taxonomicName>
to some related oxypodine genera.
<bibRefCitation author="Bernhauer and Scheerpeltz" firstAuthor="Bernhauer" pageId="11" pageNumber="22" pagination="499 - 988" refId="ref22924" refString="Bernhauer M, Scheerpeltz O (1926) Coleopterorum Catalogus, 82, Staphylinidae 6. W. Junk, Berlin, 499 - 988." type="book chapter" year="1926">Bernhauer and Scheerpeltz (1926)</bibRefCitation>
did not separate genera of Tachyusina from Falagriini and grouped them together in the subtribe
<taxonomicName box="[568,666,694,720]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="subTribe" subTribe="Falagriae">Falagriae</taxonomicName>
of the tribe
<taxonomicName authority="Ganglbauer. Lohse (1974)" authorityName="Ganglbauer. Lohse" authorityYear="1974" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Myrmedoniini">
Myrmedoniini Ganglbauer.
<bibRefCitation author="Lohse" box="[140,290,730,756]" firstAuthor="Lohse" pageId="11" pageNumber="22" refId="ref23605" refString="Lohse GA (1974) Die Kafer Mitteleuropas. In: Freude H, Harde KW, Lohse GA (Eds). Band 5: Staphylinidae II (Hypocyphtinae und Aleocharinae) Pselaphidae. Goecke &amp; Evers, Krefeld." type="book" year="1974">Lohse (1974)</bibRefCitation>
</taxonomicName>
placed
<taxonomicName box="[378,468,730,756]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[378,468,730,756]" italics="true" pageId="11" pageNumber="22">Gnypeta</emphasis>
</taxonomicName>
together with
<taxonomicName authority="Leach" authorityName="Leach" authorityYear="1819" box="[633,799,730,756]" class="Insecta" family="Staphylinidae" genus="Falagria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[633,723,730,756]" italics="true" pageId="11" pageNumber="22">Falagria</emphasis>
Leach
</taxonomicName>
,
<taxonomicName authority="Jacobs" authorityName="Jacobs" authorityYear="1925" box="[811,988,730,756]" class="Insecta" family="Staphylinidae" genus="Cordalia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[811,906,730,756]" italics="true" pageId="11" pageNumber="22">Cordalia</emphasis>
Jacobs
</taxonomicName>
,
<taxonomicName authority="Saulcy" authorityName="Saulcy" authorityYear="1864" class="Insecta" family="Staphylinidae" genus="Myrmecopora" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="11" pageNumber="22">Myrmecopora</emphasis>
Saulcy
</taxonomicName>
and allied genera of the Falagriini. We agree with
<bibRefCitation author="Seevers" box="[813,975,765,791]" firstAuthor="Seevers" pageId="11" pageNumber="22" pagination="1 - 289" refId="ref24091" refString="Seevers CH (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoology 71: i-vi, 1 - 289." type="journal article" year="1978">Seevers (1978)</bibRefCitation>
that grouping
<taxonomicName box="[181,296,800,826]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="subTribe" subTribe="Tachyusae">Tachyusae</taxonomicName>
and
<taxonomicName box="[352,450,800,826]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="subTribe" subTribe="Falagriae">Falagriae</taxonomicName>
together, mainly on the grounds of 4, 5, 5-articulated tarsi and some superficial external similarities, was artificial and unwarranted. According to
<bibRefCitation author="Seevers" box="[140,304,871,897]" firstAuthor="Seevers" pageId="11" pageNumber="22" pagination="1 - 289" refId="ref24091" refString="Seevers CH (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoology 71: i-vi, 1 - 289." type="journal article" year="1978">Seevers (1978)</bibRefCitation>
and confirmed here by us, the
<taxonomicName box="[657,772,870,896]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="subTribe" subTribe="Tachyusae">Tachyusae</taxonomicName>
lack the following specialized features of the Falagriini: paramere with codylite velum separated from the paramerite velum; pronotum much narrower at base than apex and with distinct (at least one) median sulcus (sometimes two lateral sulci present); peritremes enlarged, and contiguous with or fused to elongate prosternum; procoxal cavities closed by peritremes, prosternum and inflexed hypomera. In addition the Falagriini have a basal abdominal impression bearing a median ridge, a very distinct shape of the median lobe of the aedeagus (ovoid bulbus and tubular broad tubus), small and narrowly elongate crista apicalis of bulbus, extremely long and coiled flagellum of the internal sac of the median lobe, and the spermatheca is of a different
<typeStatus box="[552,600,1188,1214]" pageId="11" pageNumber="22">type</typeStatus>
with a small spherical capsule connected to a thin stem (
<figureCitation box="[263,480,1222,1249]" captionStart="Figs 164-171" captionStartId="54.[140,184,1590,1612]" captionTargetBox="[149,1086,244,1506]" captionTargetId="figure@54.[140,1108,232,1523]" captionTargetPageId="54" captionText="Figs 164-171. Genital structures of Gnypeta nigrella: 164, median lobe of the aedeagus in lateral view, 165, dorsal view; 166, paramere; 167, male tergite 8; 168, male sternite 8; 169, spermatheca; 170, female tergite 8; 171, female sternite 8." figureDoi="http://doi.org/10.5281/zenodo.3774126" httpUri="https://zenodo.org/record/3774126/files/figure.png" pageId="11" pageNumber="22">Figs 165, 166, 169</figureCitation>
in
<bibRefCitation author="Klimaszewski and Winchester" box="[519,927,1222,1249]" firstAuthor="Klimaszewski" pageId="11" pageNumber="22" pagination="3 - 126" refId="ref23493" refString="Klimaszewski J, Winchester NN (2002) Aleocharine rove beetles (Coleoptera Staphylinidae) of the ancient Sitka spruce forest on Vancouver Island, British Columbia, Canada. Memoires de la Societe royale belge d'Entomologie 40: 3 - 126." type="journal article" year="2002">Klimaszewski and Winchester 2002</bibRefCitation>
). For diagnostic features of Tachyusina, see
<bibRefCitation author="Seevers" box="[451,621,1258,1284]" firstAuthor="Seevers" pageId="11" pageNumber="22" pagination="1 - 289" refId="ref24091" refString="Seevers CH (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana Zoology 71: i-vi, 1 - 289." type="journal article" year="1978">Seevers (1978)</bibRefCitation>
. In his worldwide treatment of
<taxonomicName authority="Stephens, Pasnik (2006 b)" authorityName="Stephens, Pasnik" authorityYear="2006" class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[989,1108,1258,1284]" italics="true" pageId="11" pageNumber="22">Ischnopoda</emphasis>
Stephens,
<bibRefCitation author="Pasnik" box="[256,430,1293,1320]" firstAuthor="Pasnik" pageId="11" pageNumber="22" pagination="1 - 96" refId="ref23987" refString="Pasnik G (2006 b) Taxonomy and phylogeny of the World species of the genus Ischnopoda Stephens, 1837 (Coleoptera, Staphylinidae: Aleocharinae). Zootaxa 1179: 1 - 96." type="journal article" year="2006" yearSuffix="b">Paśnik (2006b)</bibRefCitation>
</taxonomicName>
considered this genus to be closely related to
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[959,1056,1293,1319]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[959,1056,1293,1319]" italics="true" pageId="11" pageNumber="22">Tachyusa</emphasis>
</taxonomicName>
and
<taxonomicName box="[140,230,1328,1354]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[140,230,1328,1354]" italics="true" pageId="11" pageNumber="22">Gnypeta</emphasis>
</taxonomicName>
. Based on a comparative study of mouthparts and the body chaetotaxy,
<bibRefCitation author="Yosii and Sawada" firstAuthor="Yosii" pageId="11" pageNumber="22" pagination="11 - 140" refId="ref24203" refString="Yosii R, Sawada K (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II. Diagnostic characters of genera and subgenera with description of representative species. Contributions of the Biological Laboratory, Kyoto University 25: 11 - 140." type="journal article" year="1976">Yosii and Sawada (1976)</bibRefCitation>
suggested a restricted concept of
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[722,819,1363,1389]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[722,819,1363,1389]" italics="true" pageId="11" pageNumber="22">Tachyusa</emphasis>
</taxonomicName>
-related genera and placed
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[140,237,1398,1424]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[140,237,1398,1424]" italics="true" pageId="11" pageNumber="22">Tachyusa</emphasis>
</taxonomicName>
together with
<emphasis box="[407,504,1399,1425]" italics="true" pageId="11" pageNumber="22">
<taxonomicName box="[407,500,1399,1425]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">Gnypeta</taxonomicName>
,
</emphasis>
and
<taxonomicName authority="Mulsant and Rey" authorityName="Mulsant and Rey" box="[564,852,1398,1425]" class="Insecta" family="Staphylinidae" genus="Dacrila" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[564,646,1398,1424]" italics="true" pageId="11" pageNumber="22">Dacrila</emphasis>
Mulsant and Rey
</taxonomicName>
in the
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[936,1039,1398,1424]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">Tachyusa</taxonomicName>
series of the Athetae. Their tribal affiliation of the
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[649,746,1434,1460]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[649,746,1434,1460]" italics="true" pageId="11" pageNumber="22">Tachyusa</emphasis>
</taxonomicName>
-related genera with athetines is strongly questionable because of the differences in the genital features of the two groups e.g., lack of the “athetine bridge” in the median lobe of the aedeagus in genera of
<taxonomicName authorityName="C.G. Thomson" authorityYear="1859" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Tachyusini">Tachyusini</taxonomicName>
.
<bibRefCitation author="Lohse" box="[208,357,1539,1565]" firstAuthor="Lohse" pageId="11" pageNumber="22" refId="ref23646" refString="Lohse GA (1989 a) Staphylinidae II (Hypocyphtinae und Aleocharinae). In: Lohse GA, Lucht WH (Ed). Die Kafer Mitteleuropas. Band 12. Supplementband mit Katalogteil. Goecke &amp; Evers, Krefeld." type="book" year="1989">Lohse (1989)</bibRefCitation>
excluded
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[468,565,1539,1565]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[468,565,1539,1565]" italics="true" pageId="11" pageNumber="22">Tachyusa</emphasis>
</taxonomicName>
,
<taxonomicName box="[578,668,1540,1566]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[578,668,1540,1566]" italics="true" pageId="11" pageNumber="22">Gnypeta</emphasis>
</taxonomicName>
, and
<taxonomicName authority="Lohse" authorityName="Lohse" authorityYear="1974" box="[729,935,1539,1565]" class="Insecta" family="Staphylinidae" genus="Dasygnypeta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[729,862,1540,1565]" italics="true" pageId="11" pageNumber="22">Dasygnypeta</emphasis>
Lohse
</taxonomicName>
from Falagriini and transferred them together with
<taxonomicName box="[544,652,1574,1600]" class="Insecta" family="Staphylinidae" genus="Brachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[544,652,1574,1600]" italics="true" pageId="11" pageNumber="22">Brachyusa</emphasis>
</taxonomicName>
and
<taxonomicName box="[710,792,1574,1600]" class="Insecta" family="Staphylinidae" genus="Dacrila" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[710,792,1574,1600]" italics="true" pageId="11" pageNumber="22">Dacrila</emphasis>
</taxonomicName>
to the separate tribe
<taxonomicName authorityName="C.G. Thomson" authorityYear="1859" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="tribe" tribe="Tachyusini">Tachyusini</taxonomicName>
.
<bibRefCitation author="Pasnik" box="[209,384,1609,1636]" firstAuthor="Pasnik" pageId="11" pageNumber="22" pagination="1 - 96" refId="ref23987" refString="Pasnik G (2006 b) Taxonomy and phylogeny of the World species of the genus Ischnopoda Stephens, 1837 (Coleoptera, Staphylinidae: Aleocharinae). Zootaxa 1179: 1 - 96." type="journal article" year="2006" yearSuffix="b">Paśnik (2006b)</bibRefCitation>
, in his revision of the world species of
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[820,917,1610,1636]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[820,917,1610,1636]" italics="true" pageId="11" pageNumber="22">Tachyusa</emphasis>
</taxonomicName>
, did not provide his view on the higher classification of
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[583,680,1645,1671]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="22" phylum="Arthropoda" rank="genus">
<emphasis box="[583,680,1645,1671]" italics="true" pageId="11" pageNumber="22">Tachyusa</emphasis>
</taxonomicName>
-related genera but did not contradict the classification proposed by
<bibRefCitation author="Lohse" box="[478,633,166,193]" firstAuthor="Lohse" pageId="12" pageNumber="23" refId="ref23646" refString="Lohse GA (1989 a) Staphylinidae II (Hypocyphtinae und Aleocharinae). In: Lohse GA, Lucht WH (Ed). Die Kafer Mitteleuropas. Band 12. Supplementband mit Katalogteil. Goecke &amp; Evers, Krefeld." type="book" year="1989">Lohse (1989)</bibRefCitation>
.
<bibRefCitation author="Pasnik" box="[644,801,166,193]" firstAuthor="Pasnik" pageId="12" pageNumber="23" pagination="1 - 54" refId="ref24022" refString="Pasnik G (2007) Revision and phylogenetic analysis of the African genus Ischnopoderona (Scheerpeltz, 1974) new status (Coleoptera, Staphylinidae, Aleocharinae). Zootaxa 1412: 1 - 54." type="journal article" year="2007">Paśnik (2007)</bibRefCitation>
published a revision of the African genus
<taxonomicName authority="(Scheerpeltz)" baseAuthorityName="Scheerpeltz" box="[307,631,202,229]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[307,472,202,228]" italics="true" pageId="12" pageNumber="23">Ischnopoderona</emphasis>
(Scheerpeltz)
</taxonomicName>
, with cladistic analysis of the species. He included there the outgroup taxa of the genera
<taxonomicName box="[670,760,237,263]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[670,760,237,263]" italics="true" pageId="12" pageNumber="23">Gnypeta</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Stephens, Pasnik" authorityYear="2006" box="[774,893,237,263]" class="Insecta" family="Staphylinidae" genus="Ischnopoda" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[774,893,237,263]" italics="true" pageId="12" pageNumber="23">Ischnopoda</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[957,1054,237,263]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[957,1054,237,263]" italics="true" pageId="12" pageNumber="23">Tachyusa</emphasis>
</taxonomicName>
. On his single most parsimonious cladogram the
<taxonomicName box="[658,823,272,298]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[658,823,272,298]" italics="true" pageId="12" pageNumber="23">Ischnopoderona</emphasis>
</taxonomicName>
branched off as a sisiter taxon of
<taxonomicName authorityName="Erichson" authorityYear="1837" box="[243,340,307,333]" class="Insecta" family="Staphylinidae" genus="Tachyusa" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[243,340,307,333]" italics="true" pageId="12" pageNumber="23">Tachyusa</emphasis>
</taxonomicName>
and both of them combined formed a sister taxon of
<taxonomicName box="[959,1049,308,334]" class="Insecta" family="Staphylinidae" genus="Gnypeta" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[959,1049,308,334]" italics="true" pageId="12" pageNumber="23">Gnypeta</emphasis>
</taxonomicName>
. The
<taxonomicName box="[140,305,342,368]" class="Insecta" family="Staphylinidae" genus="Ischnopoderona" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="23" phylum="Arthropoda" rank="genus">
<emphasis box="[140,305,342,368]" italics="true" pageId="12" pageNumber="23">Ischnopoderona</emphasis>
</taxonomicName>
was the most basal branch of this tree.We believe that more research is needed before tribal classification of this group becomes clear and stable.
</paragraph>
</subSubSection>
</treatment>
</document>