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<document id="2218F71C498AB5AD486E11E9FCE25564" ID-CLB-Dataset="25961" ID-DOI="10.11646/zootaxa.4658.1.1" ID-GBIF-Dataset="31d6e081-95c3-4086-a45a-c11555dda3aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="3373040" ID-ZooBank="ADCBCCA2-63C1-42CD-A3C9-C14F954F4A21" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1566364910789" checkinUser="plazi" docAuthor="Ziani, Stefano, Abdel-Dayem, Mahmoud Saleh, Aldhafer, Hathal Mohammed &amp; Barbero, Enrico" docDate="2019" docId="8861532B88711801FF760C0AFDB8C416" docLanguage="en" docName="zootaxa.4658.1.1.pdf" docOrigin="Zootaxa 4658 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus brittoni Paulian 1948" docType="treatment" docVersion="13" lastPageNumber="36" masterDocId="74582B5388531822FFE10957FFA3C74B" masterDocTitle="An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="35" updateTime="1698751869822" updateUser="ExternalLinkService">
<mods:mods id="4EFA53761044B02AA0E8A22FCAF0E7F7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="0E63C07E683C4BB06EBFF61B0DBCD1C3">An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="81618E64B94EDB856393B447B82E673B">Ziani, Stefano</mods:namePart>
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<mods:namePart id="0F4BD5C0ACD52E1738A83AC847A2ED6D">Abdel-Dayem, Mahmoud Saleh</mods:namePart>
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<mods:namePart id="0719B0202482E9278DDC0E9844BCE44D">Aldhafer, Hathal Mohammed</mods:namePart>
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<mods:namePart id="F5C138A9E66B9B1F396B3D789F4B3366">Barbero, Enrico</mods:namePart>
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<mods:title id="1752102204BA9A7370A5D122A68B5D5E">Zootaxa</mods:title>
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<mods:part id="0DCA03C832D96410C236D21EF04CF3FA">
<mods:date id="8AFAC3F28D7EF91434AA0C5774CA6417">2019</mods:date>
<mods:detail id="3CA03214325ABB1DE8860730727B033C" type="pubDate">
<mods:number id="5D0E520E62513E06D5ACC37DEE739C75">2019-08-21</mods:number>
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<mods:detail id="E4F2411C5D41843D8CB3D6A4ECD1BA8A" type="volume">
<mods:number id="CF9AFB7279D3C2AF38E1383344636DF9">4658</mods:number>
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<mods:identifier id="2DA02EE23A18E24532A48C28E1329AD9" type="DOI">10.11646/zootaxa.4658.1.1</mods:identifier>
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<treatment id="8861532B88711801FF760C0AFDB8C416" ID-DOI="http://doi.org/10.5281/zenodo.5667112" ID-GBIF-Taxon="159217222" ID-Zenodo-Dep="5667112" LSID="urn:lsid:plazi:treatment:8861532B88711801FF760C0AFDB8C416" httpUri="http://treatment.plazi.org/id/8861532B88711801FF760C0AFDB8C416" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
<subSubSection id="48D2B1B688711800FF760C0AFD99C233" box="[151,570,1373,1400]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="0077E23D88711800FF760C0AFD99C233" blockId="34.[151,570,1373,1436]" box="[151,570,1373,1400]" pageId="34" pageNumber="35">
<heading id="5B3F555188711800FF760C0AFD99C233" bold="true" box="[151,570,1373,1400]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<taxonomicName id="C7C899BE88711800FF760C0AFD99C233" ID-CoL="7XMJ6" authority="Paulian, 1948" authorityName="Paulian" authorityYear="1948" box="[151,570,1373,1400]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="brittoni">
<emphasis id="32BC3E2F88711800FF760C0AFD99C233" bold="true" box="[151,570,1373,1400]" pageId="34" pageNumber="35">
<emphasis id="32BC3E2F88711800FF760C0AFE32C233" bold="true" box="[151,401,1373,1400]" italics="true" pageId="34" pageNumber="35">Onthophagus brittoni</emphasis>
Paulian, 1948
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="48D2B1B688711800FF760CD5FE8DC2D7" box="[151,302,1409,1436]" pageId="34" pageNumber="35" type="description">
<paragraph id="0077E23D88711800FF760CD5FE8DC2D7" blockId="34.[151,570,1373,1436]" box="[151,302,1409,1436]" pageId="34" pageNumber="35">
[
<figureCitation id="98F3FEB888711800FF7F0CD5FE84C2D7" box="[158,295,1409,1436]" captionStart="FIGURES 19–26" captionStartId="25.[151,264,1834,1859]" captionTargetBox="[216,1343,177,1780]" captionTargetId="figure@25.[211,1377,166,1814]" captionTargetPageId="25" captionText="FIGURES 19–26. Onthophagini of Arabian Peninsula. 19, Onthophagus bicolor Raffray, 1877: OM (Dhofar) Wadi Ashawq; 20, Onthophagus latipennis d’Orbigny, 1898: OM (Dhofar) Al Mughsayl; 21, Onthophagus trapezicornis d’Orbigny, 1902: SA (Bahah) 5 km N Al Makhwah; 22, Onthophagus brittoni Paulian, 1948, ♂: SA (Asir) Abha, Raydah; 23, Onthophagus brittoni Paulian, 1948, ♀: SA (Asir) Abha, Raydah; 24, Onthophagus brittoni Paulian, 1948, paramera, lateral view, 25, Onthophagus brittoni Paulian, 1948, parameres, dorsal view; 26, Onthophagus brittoni Paulian, 1948, internal sac: accessory lamellae (above) and lamina copulatrix (below), ventral view. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373053/files/figure.png" pageId="34" pageNumber="35">
Figs. 22
<emphasis id="32BC3E2F88711800FF1A0CD4FEAAC2D7" box="[251,265,1411,1436]" italics="true" pageId="34" pageNumber="35">–</emphasis>
26
</figureCitation>
]
</paragraph>
</subSubSection>
<subSubSection id="48D2B1B688711800FF760C91FE02C2B6" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="0077E23D88711800FF760C91FE02C2B6" blockId="34.[151,1436,1477,1533]" pageId="34" pageNumber="35">
<taxonomicName id="C7C899BE88711800FF760C91FD9BC295" ID-CoL="7XMJ6" authority="Paulian, 1948: 148" authorityName="Paulian" authorityPageNumber="148" authorityYear="1948" box="[151,568,1477,1502]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="brittoni">
<emphasis id="32BC3E2F88711800FF760C91FED1C295" box="[151,370,1478,1502]" italics="true" pageId="34" pageNumber="35">Onthophagus brittoni</emphasis>
Paulian, 1948: 148
</taxonomicName>
; Paulian 1980: 153; Zunino 1980: 413; Löbl
<emphasis id="32BC3E2F88711800FC1F0C90FB89C295" box="[1022,1066,1478,1502]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2006:173; Krajcik 2012: 176; Ziani &amp; Bezdĕk 2016: 201.
</paragraph>
</subSubSection>
<subSubSection id="48D2B1B688711801FF760F7AFDB8C416" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="0077E23D88711800FF760F7AFCA1C10F" blockId="34.[151,1437,1580,2037]" box="[151,770,1580,1605]" pageId="34" pageNumber="35">
<materialsCitation id="B0A0E86088711800FF760F7AFD5DC10F" ID-GBIF-Occurrence="2350486361" box="[151,766,1580,1605]" country="Yemen" location="Western" pageId="34" pageNumber="35" specimenCount="1" stateProvince="Ad Dali'" typeStatus="holotype">
<emphasis id="32BC3E2F88711800FF760F7AFE81C10E" bold="true" box="[151,290,1580,1605]" pageId="34" pageNumber="35">
<typeStatus id="DF735C9F88711800FF760F7AFF6FC10E" box="[151,204,1581,1605]" pageId="34" pageNumber="35">Type</typeStatus>
locality
</emphasis>
. “
<location id="0517B4E688711800FED60F7AFE2FC10E" LSID="urn:lsid:plazi:treatment:8861532B88711801FF760C0AFDB8C416:0517B4E688711800FED60F7AFE2FC10E" box="[311,396,1581,1605]" country="Yemen" name="Western" pageId="34" pageNumber="35" stateProvince="Ad Dali'">Western</location>
Aden Protectorate,
<collectingRegion id="C20C2CDF88711800FDB40F7BFD37C10E" box="[597,660,1580,1605]" country="Yemen" name="Ad Dali'" pageId="34" pageNumber="35">Dhala</collectingRegion>
” [
<collectingCountry id="78DFA2AD88711800FD480F7AFD5BC10F" box="[681,760,1581,1604]" name="Yemen" pageId="34" pageNumber="35">Yemen</collectingCountry>
]
</materialsCitation>
.
</paragraph>
<paragraph id="0077E23D88711800FF260F06FB87C122" blockId="34.[151,1437,1580,2037]" box="[199,1060,1616,1641]" pageId="34" pageNumber="35">
<emphasis id="32BC3E2F88711800FF260F06FE6FC122" bold="true" box="[199,460,1616,1641]" pageId="34" pageNumber="35">Type material examined</emphasis>
.
<materialsCitation id="B0A0E86088711800FE380F07FB87C122" ID-GBIF-Occurrence="2350486248" box="[473,1060,1616,1641]" collectionCode="NHMUK" pageId="34" pageNumber="35" specimenCount="1" typeStatus="holotype">
<typeStatus id="DF735C9F88711800FE380F07FD99C122" box="[473,570,1616,1641]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
, male, fixed by original designation (
<collectionCode id="66D97AF888711800FC520F06FBBAC122" box="[947,1049,1617,1641]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="34" pageNumber="35" type="Museum">NHMUK</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="0077E23D88711800FF260F22FAE2C056" blockId="34.[151,1437,1580,2037]" pageId="34" pageNumber="35">
<emphasis id="32BC3E2F88711800FF260F22FEFCC1C6" bold="true" box="[199,351,1652,1677]" pageId="34" pageNumber="35">
<typeStatus id="DF735C9F88711800FF260F22FF5FC1C6" box="[199,252,1653,1677]" pageId="34" pageNumber="35">Type</typeStatus>
labelling
</emphasis>
. 1
<superScript id="F7BD4F7588711800FE990F24FE21C1CA" attach="none" box="[376,386,1651,1665]" fontSize="6" pageId="34" pageNumber="35">st</superScript>
, white [where specimen and edeagus are attached], handwritten in black: “
<taxonomicName id="C7C899BE88711800FB950F23FB5EC1C6" box="[1140,1277,1652,1677]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
/ brittoni / Paulian /
<typeStatus id="DF735C9F88711800FF300FCEFEB3C1FB" box="[209,272,1689,1712]" pageId="34" pageNumber="35">TYPE</typeStatus>
.”; 2
<superScript id="F7BD4F7588711800FEDC0FC0FEEFC1EE" attach="none" box="[317,332,1687,1701]" fontSize="6" pageId="34" pageNumber="35">nd</superScript>
, white, printed in black: “TAKEN AT LIGHT.”; 3
<superScript id="F7BD4F7588711800FCAD0FC0FCFBC1EE" attach="none" box="[844,856,1687,1701]" fontSize="6" pageId="34" pageNumber="35">rd</superScript>
, circular, white and red, printed in black: “
<typeStatus id="DF735C9F88711800FAEF0FCEFAE0C1FA" box="[1294,1347,1689,1713]" pageId="34" pageNumber="35">Type</typeStatus>
/ H. T.”; 4
<superScript id="F7BD4F7588711800FF450FECFF0CC182" attach="none" box="[164,175,1723,1737]" fontSize="6" pageId="34" pageNumber="35">th</superScript>
, white, printed in black: “W.
<collectingRegion id="C20C2CDF88711800FE3A0FEAFD82C19E" box="[475,545,1725,1749]" country="Yemen" name="'Adan" pageId="34" pageNumber="35">ADEN</collectingRegion>
PROT. /
<collectingRegion id="C20C2CDF88711800FD600FEBFD62C19E" box="[641,705,1724,1749]" country="Yemen" name="Ad Dali'" pageId="34" pageNumber="35">Dhala</collectingRegion>
. /
<quantity id="C7304FD888711800FD390FEBFC8BC19E" box="[728,808,1724,1749]" metricMagnitude="3" metricUnit="m" metricValue="1.46304" pageId="34" pageNumber="35" unit="ft" value="4800.0">4,800ft.</quantity>
/
<date id="7476C4FD88711800FCDA0FEBFC0DC19E" box="[827,942,1724,1749]" pageId="34" pageNumber="35" value="1937-09-14">14.ix.1937</date>
.”; 5
<superScript id="F7BD4F7588711800FC320FECFC7DC182" attach="none" box="[979,990,1723,1737]" fontSize="6" pageId="34" pageNumber="35">th</superScript>
, white, printed in black: “B. M. Exp. to / S. W.
<taxonomicName id="C7C899BE88711800FF590FB7FEA3C1B2" baseAuthorityName="Klug" baseAuthorityYear="1845" box="[184,256,1760,1785]" class="Insecta" family="Cerambycidae" genus="Arabia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Arabia</taxonomicName>
. / H. Scott &amp; / E. B. Britton.”; 6
<superScript id="F7BD4F7588711800FDA30F88FDEEC1A6" attach="none" box="[578,589,1759,1773]" fontSize="6" pageId="34" pageNumber="35">th</superScript>
, white, handwritten and printed in black: “
<taxonomicName id="C7C899BE88711800FC1C0FB7FB25C1B2" box="[1021,1158,1760,1785]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Onthophagus</taxonomicName>
/ brittoni
<taxonomicNameLabel id="298F835488711800FB0B0FB6FA83C1B3" box="[1258,1312,1761,1784]" pageId="34" pageNumber="35" rank="species">n. sp.</taxonomicNameLabel>
/ R. Paulian det.”; 7
<superScript id="F7BD4F7588711800FF030E54FF4EC05A" attach="none" box="[226,237,1795,1809]" fontSize="6" pageId="34" pageNumber="35">th</superScript>
, white, handwritten and printed in black: “
<taxonomicName id="C7C899BE88711800FD7E0E53FC38C056" authority="(Phanaeo.)" baseAuthorityName="Phanaeo." box="[671,923,1796,1821]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Onthophagus (Phanaeo.)</taxonomicName>
/
<taxonomicName id="C7C899BE88711800FC4E0E53FB97C056" authority="Paul." authorityName="Paulian" authorityYear="1948" box="[943,1076,1796,1821]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="brittoni">brittoni Paul.</taxonomicName>
/ J. SCHEUERN det. 98”.
</paragraph>
<paragraph id="0077E23D88711800FF260E7FFB4CC02E" blockId="34.[151,1437,1580,2037]" pageId="34" pageNumber="35">
<emphasis id="32BC3E2F88711800FF260E7FFE35C00A" bold="true" box="[199,406,1832,1857]" pageId="34" pageNumber="35">Diagnostic features</emphasis>
. Length 6.0 to
<quantity id="C7304FD888711800FDD10E7FFD22C00B" box="[560,641,1832,1857]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="34" pageNumber="35" unit="mm" value="8.5">8.5 mm</quantity>
. Colour blackish-brown to black, moderately glossy, with distinct isodiametric microreticulation; antennal scape, pedicel and funicle dark yellow, antennal club yellow; setation pale yellow.
</paragraph>
<paragraph id="0077E23D88711800FF260E26FAC8C0BE" blockId="34.[151,1437,1580,2037]" pageId="34" pageNumber="35">
<emphasis id="32BC3E2F88711800FF260E26FF5DC0C2" bold="true" box="[199,254,1905,1929]" pageId="34" pageNumber="35">Male</emphasis>
(
<figureCitation id="98F3FEB888711800FEEE0E27FEFDC0C2" box="[271,350,1904,1929]" captionStart="FIGURES 19–26" captionStartId="25.[151,264,1834,1859]" captionTargetBox="[216,1343,177,1780]" captionTargetId="figure@25.[211,1377,166,1814]" captionTargetPageId="25" captionText="FIGURES 19–26. Onthophagini of Arabian Peninsula. 19, Onthophagus bicolor Raffray, 1877: OM (Dhofar) Wadi Ashawq; 20, Onthophagus latipennis d’Orbigny, 1898: OM (Dhofar) Al Mughsayl; 21, Onthophagus trapezicornis d’Orbigny, 1902: SA (Bahah) 5 km N Al Makhwah; 22, Onthophagus brittoni Paulian, 1948, ♂: SA (Asir) Abha, Raydah; 23, Onthophagus brittoni Paulian, 1948, ♀: SA (Asir) Abha, Raydah; 24, Onthophagus brittoni Paulian, 1948, paramera, lateral view, 25, Onthophagus brittoni Paulian, 1948, parameres, dorsal view; 26, Onthophagus brittoni Paulian, 1948, internal sac: accessory lamellae (above) and lamina copulatrix (below), ventral view. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373053/files/figure.png" pageId="34" pageNumber="35">Fig. 22</figureCitation>
). Head with clypeal margin medially fairly strongly prolonged into a round point, sinuated at sides and strongly reflexed upward; rest of head margin evenly round to genal angles, which are round; frons flat, without any carina or depression; occipital margin raised at middle into a single semi-circular slightly prominent tubercle, flanked by a small concave hollow on each side, close to the eye; all the surface densely, coarsely and regularly punctate, except the hollows, smooth.
</paragraph>
<paragraph id="0077E23D88701801FF2609CEFB23C656" blockId="35.[151,1437,152,861]" pageId="35" pageNumber="36">Pronotum with two ridges converging anteriorly toward the middle and forming the upper anterior edge of the highly elevated disc, shaped like an isosceles triangle, with an anterolateral depression on either side; pronotal sides distinctly margined, not sinuate behind anterior angles; dorsal surface sub-regularly and densely punctate on disc, punctures separated by half to one their diameters, granulate at sides; each puncture bears a yellow seta, short on disc, longer at sides.</paragraph>
<paragraph id="0077E23D88701801FF26087FFE0DC62F" blockId="35.[151,1437,152,861]" pageId="35" pageNumber="36">Elytral striae wide, their punctures crenating interstriae; the latter flat on disc; interstriae with irregular granules, each of them bearing a yellow seta.</paragraph>
<paragraph id="0077E23D88701801FF260827FDB1C6E6" blockId="35.[151,1437,152,861]" pageId="35" pageNumber="36">Pygidium margined basally, setigerous punctate, punctures large, coarse, sub-regularly impressed, surface between punctures shining, clearly microreticulate.</paragraph>
<paragraph id="0077E23D88701801FF2608EFFBA9C552" blockId="35.[151,1437,152,861]" pageId="35" pageNumber="36">Middle and hind femora ventrally with sparse setigerous punctures; foretibiae quadridentate externally; middle and hind tibiae short, triangular, broad apically; apical spur clearly longer than first segment of hind tarsi; middle and hind tarsi more or less widened and flat, the second to fourth tarsal joints differing only slightly in length.</paragraph>
<paragraph id="0077E23D88701801FF260B72FE5AC5CE" blockId="35.[151,1437,152,861]" pageId="35" pageNumber="36">
<emphasis id="32BC3E2F88701801FF260B72FEB5C576" bold="true" box="[199,278,549,573]" pageId="35" pageNumber="36">Female</emphasis>
(
<figureCitation id="98F3FEB888701801FEC70B73FED0C576" box="[294,371,548,573]" captionStart="FIGURES 19–26" captionStartId="25.[151,264,1834,1859]" captionTargetBox="[216,1343,177,1780]" captionTargetId="figure@25.[211,1377,166,1814]" captionTargetPageId="25" captionText="FIGURES 19–26. Onthophagini of Arabian Peninsula. 19, Onthophagus bicolor Raffray, 1877: OM (Dhofar) Wadi Ashawq; 20, Onthophagus latipennis d’Orbigny, 1898: OM (Dhofar) Al Mughsayl; 21, Onthophagus trapezicornis d’Orbigny, 1902: SA (Bahah) 5 km N Al Makhwah; 22, Onthophagus brittoni Paulian, 1948, ♂: SA (Asir) Abha, Raydah; 23, Onthophagus brittoni Paulian, 1948, ♀: SA (Asir) Abha, Raydah; 24, Onthophagus brittoni Paulian, 1948, paramera, lateral view, 25, Onthophagus brittoni Paulian, 1948, parameres, dorsal view; 26, Onthophagus brittoni Paulian, 1948, internal sac: accessory lamellae (above) and lamina copulatrix (below), ventral view. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373053/files/figure.png" pageId="35" pageNumber="36">Fig. 23</figureCitation>
). Head with clypeal margin prolonged medially but not forming a round point; pronotal process weak, represented only by two anterolateral faint depressions; pronotal punctation smaller and sparser, punctures separated by one and half to two their diameters on disc.
</paragraph>
<paragraph id="0077E23D88701801FF260BC7FF4CC5BA" blockId="35.[151,1437,152,861]" pageId="35" pageNumber="36">
<emphasis id="32BC3E2F88701801FF260BC7FE98C5E2" bold="true" box="[199,315,656,681]" pageId="35" pageNumber="36">Variability</emphasis>
. As with most
<taxonomicName id="C7C899BE88701801FE360BC7FDC1C5E2" authorityName="Burmeister" authorityYear="1846" box="[471,610,656,681]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="tribe" tribe="Onthophagini">Onthophagini</taxonomicName>
species, intraspecific variability stands in the degree of development of pronotal process: minor males have a like-female pronotum ornamentation. Furthermore, the colour of the body varies from dark brown to black.
</paragraph>
<paragraph id="0077E23D88701801FF260BAAFDB8C416" blockId="35.[151,1437,152,861]" pageId="35" pageNumber="36">
<emphasis id="32BC3E2F88701801FF260BAAFE1AC45E" bold="true" box="[199,441,764,789]" pageId="35" pageNumber="36">Male genital armature</emphasis>
. Parameres very short, sinuate laterally in dorsal view, with strong latero-apical spatulae bent ventrally, not divergent apically (
<figureCitation id="98F3FEB888701801FE270A77FDE1C472" box="[454,578,800,825]" captionStart="FIGURES 19–26" captionStartId="25.[151,264,1834,1859]" captionTargetBox="[216,1343,177,1780]" captionTargetId="figure@25.[211,1377,166,1814]" captionTargetPageId="25" captionText="FIGURES 19–26. Onthophagini of Arabian Peninsula. 19, Onthophagus bicolor Raffray, 1877: OM (Dhofar) Wadi Ashawq; 20, Onthophagus latipennis d’Orbigny, 1898: OM (Dhofar) Al Mughsayl; 21, Onthophagus trapezicornis d’Orbigny, 1902: SA (Bahah) 5 km N Al Makhwah; 22, Onthophagus brittoni Paulian, 1948, ♂: SA (Asir) Abha, Raydah; 23, Onthophagus brittoni Paulian, 1948, ♀: SA (Asir) Abha, Raydah; 24, Onthophagus brittoni Paulian, 1948, paramera, lateral view, 25, Onthophagus brittoni Paulian, 1948, parameres, dorsal view; 26, Onthophagus brittoni Paulian, 1948, internal sac: accessory lamellae (above) and lamina copulatrix (below), ventral view. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373053/files/figure.png" pageId="35" pageNumber="36">
Figs. 24
<emphasis id="32BC3E2F88701801FDF80A76FD85C473" box="[537,550,801,824]" italics="true" pageId="35" pageNumber="36">–</emphasis>
25
</figureCitation>
); internal sac with some accessory lamellae and a comma-shaped lamina copulatrix with two counterposed arms (
<figureCitation id="98F3FEB888701801FE220A13FDB3C416" box="[451,528,836,861]" captionStart="FIGURES 19–26" captionStartId="25.[151,264,1834,1859]" captionTargetBox="[216,1343,177,1780]" captionTargetId="figure@25.[211,1377,166,1814]" captionTargetPageId="25" captionText="FIGURES 19–26. Onthophagini of Arabian Peninsula. 19, Onthophagus bicolor Raffray, 1877: OM (Dhofar) Wadi Ashawq; 20, Onthophagus latipennis d’Orbigny, 1898: OM (Dhofar) Al Mughsayl; 21, Onthophagus trapezicornis d’Orbigny, 1902: SA (Bahah) 5 km N Al Makhwah; 22, Onthophagus brittoni Paulian, 1948, ♂: SA (Asir) Abha, Raydah; 23, Onthophagus brittoni Paulian, 1948, ♀: SA (Asir) Abha, Raydah; 24, Onthophagus brittoni Paulian, 1948, paramera, lateral view, 25, Onthophagus brittoni Paulian, 1948, parameres, dorsal view; 26, Onthophagus brittoni Paulian, 1948, internal sac: accessory lamellae (above) and lamina copulatrix (below), ventral view. Photos by Gabriele Fiumi." httpUri="https://zenodo.org/record/3373053/files/figure.png" pageId="35" pageNumber="36">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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