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<mods:title id="02631DE4A50281A21CF9E6A3CDE6C74A">An overview of the Onthophagini from the Arabian Peninsula (Coleoptera: Scarabaeoidea: Scarabaeidae)</mods:title>
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<mods:namePart id="68FD670C2C72935279FB22A85BD438EE">Ziani, Stefano</mods:namePart>
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<mods:namePart id="6A63298C3B8E0617E5F55141D1011D1E">Abdel-Dayem, Mahmoud Saleh</mods:namePart>
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<mods:namePart id="B708996C1F103BA3E16E7E01FC4D3B34">Aldhafer, Hathal Mohammed</mods:namePart>
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<mods:namePart id="B4FD283453833012455EDE41FA64DABE">Barbero, Enrico</mods:namePart>
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<mods:title id="B6CABBB888B02A13CDAC6319C5C213C0">Zootaxa</mods:title>
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<mods:number id="A74CB5F1A205240768A62C4A93C467F8">2019-08-21</mods:number>
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<treatment id="8861532B8854182AFF760E0EFA9EC6BC" ID-DOI="http://doi.org/10.5281/zenodo.5667065" ID-GBIF-Taxon="159217233" ID-Zenodo-Dep="5667065" LSID="urn:lsid:plazi:treatment:8861532B8854182AFF760E0EFA9EC6BC" httpUri="http://treatment.plazi.org/id/8861532B8854182AFF760E0EFA9EC6BC" lastPageId="8" lastPageNumber="8" pageId="7" pageNumber="8">
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<subSubSection id="48D2B1B688541825FF760E0EFD07C038" box="[151,676,1881,1908]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="0077E23D88541825FF760E0EFD07C038" blockId="7.[151,676,1881,1908]" box="[151,676,1881,1908]" pageId="7" pageNumber="8">
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<heading id="5B3F555188541825FF760E0EFD07C038" bold="true" box="[151,676,1881,1908]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<emphasis id="32BC3E2F88541825FF760E0EFD07C038" bold="true" box="[151,676,1881,1908]" pageId="7" pageNumber="8">
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Subgenus
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<taxonomicName id="C7C899BE88541825FEF60E0EFD07C038" ID-CoL="649SW" authority="Zunino, 1979" authorityName="Zunino" authorityYear="1979" box="[279,676,1881,1908]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Eremonthophagus">
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<emphasis id="32BC3E2F88541825FEF60E0EFE5BC03F" bold="true" box="[279,504,1881,1908]" italics="true" pageId="7" pageNumber="8">Eremonthophagus</emphasis>
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<bibRefCitation id="64599FCC88541825FE1E0E0EFD07C038" author="Zunino, M." box="[511,676,1881,1907]" pageId="7" pageNumber="8" pagination="289 - 292" refId="ref40222" refString="Zunino, M. (1979) Insects of Saudi Arabia Coleoptera: Fam. Scarabaeidae Tribus Onthophagini. Fauna of Saudi Arabia, 1, 289 - 292." type="journal article" year="1979">Zunino, 1979</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="48D2B1B68854182AFF760EF6FA9EC6BC" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="0077E23D8854182AFF760EF6FA9EC6BC" blockId="7.[151,1436,1953,2015]" lastBlockId="8.[151,1437,153,503]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
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<emphasis id="32BC3E2F88541825FF760EF6FEA7C0F0" bold="true" box="[151,260,1953,1979]" pageId="7" pageNumber="8">Remarks</emphasis>
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. According to
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<bibRefCitation id="64599FCC88541825FE4B0EF6FDC3C0F7" author="Kabakov, O. N." box="[426,608,1953,1980]" pageId="7" pageNumber="8" refId="ref38133" refString="Kabakov, O. N. (2006) Plastinchatousye zhuki podsemeystva Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) fauny Rossii i sopredel'nykh stran. [The lamellicorn beetle subfamily Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) in the fauna of Russia and adjacent countries]. Tovarishchestvo Nauchnykh Izdanii KMK, Moskva, 374 pp." type="book" year="2006">Kabakov (2006)</bibRefCitation>
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the subgenus
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<taxonomicName id="C7C899BE88541825FD1E0EF6FBCBC0F0" authority="Zunino, 1979" authorityName="Zunino" authorityYear="1979" box="[767,1128,1953,1979]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Eremonthophagus">
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<emphasis id="32BC3E2F88541825FD1E0EF6FC68C0F0" box="[767,971,1953,1979]" italics="true" pageId="7" pageNumber="8">Eremonthophagus</emphasis>
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<bibRefCitation id="64599FCC88541825FC330EF6FBCBC0F0" author="Zunino, M." box="[978,1128,1953,1979]" pageId="7" pageNumber="8" pagination="289 - 292" refId="ref40222" refString="Zunino, M. (1979) Insects of Saudi Arabia Coleoptera: Fam. Scarabaeidae Tribus Onthophagini. Fauna of Saudi Arabia, 1, 289 - 292." type="journal article" year="1979">Zunino, 1979</bibRefCitation>
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</taxonomicName>
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is to be considered a junior synonym of the subgenus
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<taxonomicName id="C7C899BE88541825FE560E92FCE0C094" authority="Balthasar, 1963" authorityName="Balthasar" authorityYear="1963" box="[439,835,1989,2015]" class="Insecta" family="Scarabaeidae" genus="Colobonthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="32BC3E2F88541825FE560E92FD2FC094" box="[439,652,1989,2015]" italics="true" pageId="7" pageNumber="8">Colobonthophagus</emphasis>
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<bibRefCitation id="64599FCC88541825FD730E92FCE0C094" author="Balthasar, V." box="[658,835,1989,2015]" pageId="7" pageNumber="8" refId="ref36564" refString="Balthasar, V. (1963) Monographie der Scarabaeidae und Aphodiidae der palaearktischen und orientalischen Region. Coleoptera: Lamellicornia. Band 2. Coprinae. Tschechoslowakischen Akademie der Wissenschaften, Prag, 628 pp." type="book" year="1963">Balthasar, 1963</bibRefCitation>
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</taxonomicName>
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, and all the species of the former must be placed in the latter, even if they do not have the bulbous elevation at base of justasutural interstria, that is, one of the distinctive characters of
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<taxonomicName id="C7C899BE885B182AFED009EAFDA5C79C" authorityName="Balthasar" authorityYear="1963" box="[305,518,189,215]" class="Insecta" family="Scarabaeidae" genus="Colobonthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="32BC3E2F885B182AFED009EAFDA5C79C" box="[305,518,189,215]" italics="true" pageId="8" pageNumber="9">Colobonthophagus</emphasis>
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</taxonomicName>
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: “...а таКЖе виды подрода
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<taxonomicName id="C7C899BE885B182AFCA909EAFBD2C79C" authority="Zunino" authorityName="Zunino" authorityYear="1979" box="[840,1137,189,215]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Eremonthophagus">
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<emphasis id="32BC3E2F885B182AFCA909EAFBB7C79C" box="[840,1044,189,215]" italics="true" pageId="8" pageNumber="9">Eremonthophagus</emphasis>
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Zunino
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</taxonomicName>
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, ЯвЛЯЮЩиеСЯ по нашему мнениЮ аридным дериватом типицных
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<emphasis id="32BC3E2F885B182AFD8D09B6FCF2C7B0" box="[620,849,225,251]" italics="true" pageId="8" pageNumber="9">
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<taxonomicName id="C7C899BE885B182AFD8D09B6FCE4C7B0" authorityName="Balthasar" authorityYear="1963" box="[620,839,225,251]" class="Insecta" family="Scarabaeidae" genus="Colobonthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">Colobonthophagus</taxonomicName>
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”
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</emphasis>
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[...as well as species of the subgenus
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<taxonomicName id="C7C899BE885B182AFB0F09B6FF1DC654" authorityName="Zunino" authorityYear="1979" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subGenus" subGenus="Eremonthophagus">
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<emphasis id="32BC3E2F885B182AFB0F09B6FF1DC654" italics="true" pageId="8" pageNumber="9">Eremonthophagus</emphasis>
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</taxonomicName>
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, which in our opinion are arid derivatives of typical
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<taxonomicName id="C7C899BE885B182AFCF10852FC46C654" authorityName="Balthasar" authorityYear="1963" box="[784,997,261,287]" class="Insecta" family="Scarabaeidae" genus="Colobonthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="32BC3E2F885B182AFCF10852FC46C654" box="[784,997,261,287]" italics="true" pageId="8" pageNumber="9">Colobonthophagus</emphasis>
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</taxonomicName>
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]. That “arid”, meant as living in desert areas, is referred to the xerothermic habitat of some
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<taxonomicName id="C7C899BE885B182AFD36087EFC00C608" authorityName="Zunino" authorityYear="1979" box="[727,931,297,323]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="32" phylum="Arthropoda" rank="subGenus" subGenus="Eremonthophagus">
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<emphasis id="32BC3E2F885B182AFD36087EFC00C608" box="[727,931,297,323]" italics="true" pageId="8" pageNumber="9">Eremonthophagus</emphasis>
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</taxonomicName>
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species. According to
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<bibRefCitation id="64599FCC885B182AFB7F087EFAF4C60F" author="Kabakov, O. N." box="[1182,1367,297,324]" pageId="8" pageNumber="9" refId="ref38133" refString="Kabakov, O. N. (2006) Plastinchatousye zhuki podsemeystva Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) fauny Rossii i sopredel'nykh stran. [The lamellicorn beetle subfamily Scarabaeinae (Insecta: Coleoptera: Scarabaeidae) in the fauna of Russia and adjacent countries]. Tovarishchestvo Nauchnykh Izdanii KMK, Moskva, 374 pp." type="book" year="2006">Kabakov (2006)</bibRefCitation>
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, some other external morphological features justify the synonymy, and allow
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<taxonomicName id="C7C899BE885B182AFC5C081AFAE5C62C" authority="sensu Kabakov" authorityName="Kabakov" box="[957,1350,333,359]" class="Insecta" family="Scarabaeidae" genus="Colobonthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="32BC3E2F885B182AFC5C081AFB7AC62C" box="[957,1241,333,359]" italics="true" pageId="8" pageNumber="9">Colobonthophagus sensu</emphasis>
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Kabakov
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</taxonomicName>
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to be a homogenous group. However, the morphology of male genitalia, particularly the lamella copulatrix, and the external character of the base of sutural interstria of elytra, that can be considered a synapomorphic, and do not allow us to agree with the synonymy of Kabakov. As already stated by
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<bibRefCitation id="64599FCC885B182AFCB008EEFBFAC698" author="Tagliaferri, F. & Moretto, P. & Tarasov, S." box="[849,1113,441,467]" pageId="8" pageNumber="9" pagination="1 - 31" refId="ref39627" refString="Tagliaferri, F., Moretto, P. & Tarasov, S. (2012) Essai sur la systematique et la phylogenie des Onthophagus Latreille, 1802, d'Afrique tropicale appartenant au septieme groupe de d'Orbigny. Description d'un sous-genre nouveau et de trois especes nouvelles (Coleoptera, Scarabaeoidea, Onthophagini). Catharsius La Revue, 6, 1 - 31." type="journal article" year="2012">
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Tagliaferri
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<emphasis id="32BC3E2F885B182AFC2F08EDFBA3C698" box="[974,1024,441,467]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
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. (2012)
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</bibRefCitation>
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and
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<bibRefCitation id="64599FCC885B182AFB7108EEFA3BC69F" author="Ziani, S. & Bezdek, A." box="[1168,1432,441,468]" pageId="8" pageNumber="9" pagination="180 - 204" refId="ref39999" refString="Ziani, S. & Bezdek, A. (2016) Scarabaeidae, subfamily Scarabaeinae, tribe Onthophagini. In: Lobl, I. & Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea - Scirtoidea - Dascilloidea - Buprestoidea - Byrrhoidea. Revised and updated edition. Brill, Leiden-Boston, pp. 180 - 204." type="book chapter" year="2016">Ziani & Bezděk (2016)</bibRefCitation>
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, the subgenus
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<taxonomicName id="C7C899BE885B182AFED1088AFE5FC6BC" authorityName="Zunino" authorityYear="1979" box="[304,508,477,503]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subGenus" subGenus="Eremonthophagus">
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<emphasis id="32BC3E2F885B182AFED1088AFE5FC6BC" box="[304,508,477,503]" italics="true" pageId="8" pageNumber="9">Eremonthophagus</emphasis>
|
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</taxonomicName>
|
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is herein considered a valid subgenus, separated from
|
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<taxonomicName id="C7C899BE885B182AFB80088AFA95C6BC" box="[1121,1334,477,503]" class="Insecta" family="Scarabaeidae" genus="Colobonthophagus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
|
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<emphasis id="32BC3E2F885B182AFB80088AFA95C6BC" box="[1121,1334,477,503]" italics="true" pageId="8" pageNumber="9">Colobonthophagus</emphasis>
|
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|
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</taxonomicName>
|
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|
|
.
|
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|
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</paragraph>
|
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</subSubSection>
|
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</treatment>
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</document>
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