treatments-xml/data/03/C0/87/03C08782FFDFFFE7FC003E79FDC6FBA2.xml

<document id="286D33C9738ECE68B1B3DBAE6D8E87B7" ID-CLB-Dataset="21692" ID-DOI="10.5252/geodiversitas2020v42a13" ID-GBIF-Dataset="d64d995c-06c4-4541-aae0-737c86d030a9" ID-ISSN="1638-9395" ID-Zenodo-Dep="3894810" ID-ZooBank="urn:lsid:zoobank.org:pub:52D33BB7-2713-4F9D-AACF-3FAA90FDB31F" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1592234070810" checkinUser="valdenar" docAuthor="Solé, Floréal, Marandat, Bernard &amp; Lihoreau, Fabrice" docDate="2020" docId="03C08782FFDFFFE7FC003E79FDC6FBA2" docLanguage="en" docName="GEODIVERSITAS.42.13.185-214.pdf.imf" docOrigin="Geodiversitas 42 (13)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Matthodon peignei Solé &amp; Marandat &amp; Lihoreau 2020, n. sp." docType="treatment" docUuid="F79F2003-3B67-466B-AA9E-03F94442912F" docUuidSource="ZooBank" docVersion="10" lastPageNumber="190" masterDocId="FFF9FFFAFFDBFFE0FFE73A2CFF8FFFB3" masterDocTitle="The hyaenodonts (Mammalia) from the French locality of Aumelas (Hérault), with possible new representatives from the late Ypresian" masterLastPageNumber="214" masterPageNumber="185" pageNumber="187" updateTime="1732705675866" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="8697E051EF7591D3486663D3889E4B16">The hyaenodonts (Mammalia) from the French locality of Aumelas (Hérault), with possible new representatives from the late Ypresian</mods:title>
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<mods:namePart id="ABB821CB2EB56E3A79DE17B05D483C69">Solé, Floréal</mods:namePart>
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<mods:namePart id="963380CBD9FA629607AF3B8DB52C019C">Marandat, Bernard</mods:namePart>
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<mods:namePart id="1D4E99CEBDFEA4BBF711F61482205576">Lihoreau, Fabrice</mods:namePart>
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<mods:title id="41A65440AE9030149A8BDFAFDD18922F">Geodiversitas</mods:title>
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<subSubSection id="C373651FFFDFFFE4FC003E79FB7AFBC2" box="[999,1269,1109,1137]" pageId="4" pageNumber="187" type="nomenclature">
<paragraph id="8BD63694FFDFFFE4FC003E79FB7AFBC2" blockId="4.[999,1269,1109,1169]" box="[999,1269,1109,1137]" pageId="4" pageNumber="187">
<heading id="D09E81F8FFDFFFE4FC003E79FB7AFBC2" box="[999,1269,1109,1137]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="187" reason="2">
<taxonomicName id="4C694D17FFDFFFE4FC003E79FB3CFBC3" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[999,1203,1109,1136]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="4" pageNumber="187" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="B91DEA86FFDFFFE4FC003E79FB3CFBC3" bold="true" box="[999,1203,1109,1136]" italics="true" pageId="4" pageNumber="187">Matthodon peignei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDFFFE4FB5D3E7BFB7AFBC2" box="[1210,1269,1111,1137]" pageId="4" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C373651FFFDFFFE4FBA13E5AFA3FFB74" pageId="4" pageNumber="187" type="description">
<paragraph id="8BD63694FFDFFFE4FBA13E5AFB19FB22" blockId="4.[999,1269,1109,1169]" box="[1094,1174,1142,1169]" pageId="4" pageNumber="187">
(
<figureCitation id="13522A11FFDFFFE4FBA83E5AFB02FB23" box="[1103,1165,1142,1168]" captionStart="FIG" captionStartId="6.[132,143,1962,1979]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure@6.[132,1455,215,1878]" captionTargetPageId="6" captionText="FIG. 1. — Matthodon peignei n. sp. from Aumelas (MP10-MP11?): A-C, UM-AUM694 (Holotype), left mandible bearing the p2-m3 and the alveoli of the canine and p1; A, occlusal view; B, lingual view; C, labial view; D-F, UM-AUM268, left mandible bearing the m2 and m3; D, occlusal view; E, lingual view; F, labial view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3894812" httpUri="https://zenodo.org/record/3894812/files/figure.png" pageId="4" pageNumber="187">Fig. 1</figureCitation>
)
</paragraph>
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urn:lsid:zoobank.org:act: 
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<subSubSection id="C373651FFFDFFFE4FCCA3ED0FBC6FA9C" pageId="4" pageNumber="187" type="materials_examined">
<paragraph id="8BD63694FFDFFFE4FCCA3ED0FBC6FA9C" blockId="4.[813,1455,1276,1327]" pageId="4" pageNumber="187">
<materialsCitation id="3B013CC9FFDFFFE4FCCA3ED0FBC6FA9C" ID-GBIF-Occurrence="2635845304" collectionCode="UM-AUM" pageId="4" pageNumber="187" specimenCode="UM-AUM694" specimenCount="1" typeStatus="holotype">
<typeStatus id="54D28836FFDFFFE4FCCA3ED0FC18FAA1" box="[813,919,1276,1299]" pageId="4" pageNumber="187" type="holotype">HOLOTYPE</typeStatus>
. — 
<specimenCode id="DBCF9EEFFFDFFFE4FC243ED0FBD7FAA7" box="[963,1112,1276,1300]" collectionCode="UM-AUM" pageId="4" pageNumber="187">UM-AUM694</specimenCode>
, left mandible with p2-m3 and the alveoli for the canine and p1.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDFFFE4FCCA3F65FB60FA24" pageId="4" pageNumber="187" type="etymology">
<paragraph id="8BD63694FFDFFFE4FCCA3F65FB60FA24" blockId="4.[813,1456,1353,1431]" pageId="4" pageNumber="187">
ETYMOLOGY. — Dedicated to the memory of Dr. S. Peigné (
<date id="FFD71054FFDFFFE4FA933F66FCECFACF" pageId="4" pageNumber="187" value="1972" valueMax="2017">1972- 2017</date>
), who described numerous carnivorous mammals from the Paleogene and Neogene of Eurasia and Africa.
</paragraph>
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<subSubSection id="C373651FFFDFFFE4FCCA3F9DFCF0FA57" pageId="4" pageNumber="187" type="materials_examined">
<paragraph id="8BD63694FFDFFFE4FCCA3F9DFCF0FA57" blockId="4.[813,1455,1457,1508]" pageId="4" pageNumber="187">
<materialsCitation id="3B013CC9FFDFFFE4FCCA3F9DFCF0FA57" ID-GBIF-Occurrence="2635845306" pageId="4" pageNumber="187">
REFERRED SPECIMENS. 
<emphasis id="B91DEA86FFDFFFE4FBEA3F9DFBA9FA7A" bold="true" box="[1037,1062,1457,1481]" pageId="4" pageNumber="187">—</emphasis>
UM-AUM268, left mandible with m2 and m3.
</materialsCitation>
</paragraph>
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<subSubSection id="C373651FFFDFFFE4FCCA3FD2FBF7F9FF" pageId="4" pageNumber="187" type="description">
<paragraph id="8BD63694FFDFFFE4FCCA3FD2FBF7F9FF" blockId="4.[813,1455,1534,1612]" pageId="4" pageNumber="187">HORIZON AND AGE. — Lacustrine limestone of Montpellier, proposed as Lutetian in age in previous study but here considered possibly late Ypresian/early Lutetian in age</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDFFFE4FCCA3C49FB63F9CD" box="[813,1260,1637,1662]" pageId="4" pageNumber="187" type="distribution">
<paragraph id="8BD63694FFDFFFE4FCCA3C49FB63F9CD" blockId="4.[813,1260,1637,1662]" box="[813,1260,1637,1662]" pageId="4" pageNumber="187">
TYPE LOCALITY. — Aumelas, Hérault, 
<collectingCountry id="F37E7604FFDFFFE4FB443C4BFB68F9CD" box="[1187,1255,1639,1662]" name="France" pageId="4" pageNumber="187">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDFFFE4FCCA3CB4FAF8F80A" pageId="4" pageNumber="187" type="diagnosis">
<paragraph id="8BD63694FFDFFFE4FCCA3CB4FAF8F80A" blockId="4.[813,1456,1688,1978]" pageId="4" pageNumber="187">
DIAGNOSIS. — 
<taxonomicName id="4C694D17FFDFFFE4FC263CB5FBE0F902" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[961,1135,1689,1713]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="4" pageNumber="187" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="B91DEA86FFDFFFE4FC263CB5FBE0F902" box="[961,1135,1689,1713]" italics="true" pageId="4" pageNumber="187">Matthodon peignei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDFFFE4FB933CB5FB27F903" box="[1140,1192,1689,1712]" pageId="4" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
is intermediate both metrically and morphologically between 
<emphasis id="B91DEA86FFDFFFE4FB903C9FFA93F978" box="[1143,1308,1715,1739]" italics="true" pageId="4" pageNumber="187">
<taxonomicName id="4C694D17FFDFFFE4FB903C9FFB51F978" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[1143,1246,1715,1739]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="4" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
menui
</emphasis>
and 
<emphasis id="B91DEA86FFDFFFE4FAAF3C9FFCE7F956" italics="true" pageId="4" pageNumber="187">
<taxonomicName id="4C694D17FFDFFFE4FAAF3C9FFA20F978" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[1352,1455,1715,1739]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="4" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
tritens
</emphasis>
: 
<taxonomicName id="4C694D17FFDFFFE4FC913CE3FC53F956" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[886,988,1742,1766]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="4" pageNumber="187" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="B91DEA86FFDFFFE4FC913CE3FC53F956" box="[886,988,1742,1766]" italics="true" pageId="4" pageNumber="187">M. peignei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDFFFE4FC033CE2FB94F956" box="[996,1051,1742,1765]" pageId="4" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
differs from 
<emphasis id="B91DEA86FFDFFFE4FB783CE3FA8FF956" box="[1183,1280,1742,1766]" italics="true" pageId="4" pageNumber="187">M. menui</emphasis>
by having longer premolars and molars and from 
<emphasis id="B91DEA86FFDFFFE4FB803CC5FB4BF8B3" box="[1127,1220,1769,1792]" italics="true" pageId="4" pageNumber="187">M. tritens</emphasis>
by having mesiodistally shorter premolars and molars. 
<taxonomicName id="4C694D17FFDFFFE4FBB63D2FFA8FF8A9" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[1105,1280,1795,1819]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="4" pageNumber="187" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="B91DEA86FFDFFFE4FBB63D2FFA8FF8A9" box="[1105,1280,1795,1819]" italics="true" pageId="4" pageNumber="187">Matthodon peignei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDFFFE4FAE13D2FFAB5F8A9" box="[1286,1338,1795,1818]" pageId="4" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
differs from 
<emphasis id="B91DEA86FFDFFFE4FCCA3D32FC09F886" box="[813,902,1822,1845]" italics="true" pageId="4" pageNumber="187">M. tritens</emphasis>
by having less developed paraconids on p2, p3 and p4; the larger and more individualized metaconid on the molars; the longer talonid on the m1 and m2; and by shorter molars (4% for the m1, 8% for the m2 and m3), and longer p2 and p4 (respectively 9% and 5%). 
<taxonomicName id="4C694D17FFDFFFE4FC853DABFB9DF82C" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[866,1042,1927,1951]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="4" pageNumber="187" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="B91DEA86FFDFFFE4FC853DABFB9DF82C" box="[866,1042,1927,1951]" italics="true" pageId="4" pageNumber="187">Matthodon peignei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDFFFE4FBF03DA4FBC3F82C" box="[1047,1100,1928,1951]" pageId="4" pageNumber="187" rank="species">n. sp.</taxonomicNameLabel>
differs from 
<emphasis id="B91DEA86FFDFFFE4FB203DA4FAAAF82C" box="[1223,1317,1928,1951]" italics="true" pageId="4" pageNumber="187">M. menui</emphasis>
and 
<emphasis id="B91DEA86FFDFFFE4FAB43DA4FA20F82C" box="[1363,1455,1928,1951]" italics="true" pageId="4" pageNumber="187">M. tritens</emphasis>
by the relative larger size of the p2, and narrower premolars.
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDFFFE5FCCA3DFFFA88FD61" lastPageId="5" lastPageNumber="188" pageId="4" pageNumber="187" type="description">
<paragraph id="8BD63694FFDFFFE4FCCA3DFFFBCBF85F" blockId="4.[813,1092,2003,2028]" box="[813,1092,2003,2028]" pageId="4" pageNumber="187">
MEASUREMENTS. — 
<tableCitation id="C6EB032FFFDFFFE4FC113DF8FBCBF85F" box="[1014,1092,2004,2028]" captionStart="TABLE" captionStartId="5.[132,143,219,236]" captionText="TABLE 1. — Measurements (in mm) of Matthodon peignei n. sp. from Aumelas (MP10-MP11?). Body mass (c. 14.95 kg) estimated after Morlo (1999)." httpUri="http://table.plazi.org/id/DF16661CFFDEFFE5FF633AF7FD44FEB0" pageId="4" pageNumber="187" tableUuid="DF16661CFFDEFFE5FF633AF7FD44FEB0">Table 1.</tableCitation>
</paragraph>
<caption id="DF16661CFFDEFFE5FF633AF7FD44FEB0" ID-Table-UUID="DF16661CFFDEFFE5FF633AF7FD44FEB0" httpUri="http://table.plazi.org/id/DF16661CFFDEFFE5FF633AF7FD44FEB0" pageId="5" pageNumber="188" targetBox="[134,700,317,849]" targetIsTable="true" targetPageId="5">
<paragraph id="8BD63694FFDEFFE5FF633AF7FD44FEB0" blockId="5.[132,775,219,259]" pageId="5" pageNumber="188">
<subScript id="17ED34D1FFDEFFE5FF633AF7FF3DFF5F" attach="left" box="[132,178,219,236]" fontSize="5" pageId="5" pageNumber="188">TABLE</subScript>
1. — Measurements (in mm) of 
<taxonomicName id="4C694D17FFDEFFE5FE5A3AF7FDD5FF5F" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[445,602,219,236]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="188" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="B91DEA86FFDEFFE5FE5A3AF7FDD5FF5F" box="[445,602,219,236]" italics="true" pageId="5" pageNumber="188">Matthodon peignei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDEFFE5FDB83AF7FD00FF5F" box="[607,655,219,236]" pageId="5" pageNumber="188" rank="species">n. sp.</taxonomicNameLabel>
from Aumelas (MP10-MP11?). Body mass (
<emphasis id="B91DEA86FFDEFFE5FE933ADEFE0BFEB0" box="[372,388,242,259]" italics="true" pageId="5" pageNumber="188">c.</emphasis>
14.95 kg) estimated after 
<bibRefCitation id="EFF84B65FFDEFFE5FDBA3ADEFD47FEB0" author="MORLO M." box="[605,712,242,259]" journalOrPublisher="Geobios" pageId="5" pageNumber="188" pagination="297 - 305" part="32" refId="ref15217" refString="MORLO M. 1999. - Niche structure and evolution in creodont (Mammalia) faunas of the European and North American Eocene. Geobios 32: 297 - 305. https: // doi. org / 10.1016 / S 0016 - 6995 (99) 80043 - 6" title="Niche structure and evolution in creodont (Mammalia) faunas of the European and North American Eocene" type="journal article" year="1999">Morlo (1999)</bibRefCitation>
.
</paragraph>
</caption>
<caption id="DF16661CFFDEFFE5FCCA3AF7FC5BFEB0" pageId="5" pageNumber="188">
<paragraph id="8BD63694FFDEFFE5FCCA3AF7FC5BFEB0" blockId="5.[813,1455,219,259]" pageId="5" pageNumber="188">
<subScript id="17ED34D1FFDEFFE5FCCA3AF7FCD5FF5F" attach="left" box="[813,858,219,236]" fontSize="5" pageId="5" pageNumber="188">TABLE</subScript>
2. — Mean width/length ratio of the p2, p3 and p4 respectively in the 
<taxonomicName id="4C694D17FFDEFFE5FCCA3ADEFC07FEB0" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[813,904,242,259]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDEFFE5FCCA3ADEFC07FEB0" box="[813,904,242,259]" italics="true" pageId="5" pageNumber="188">Matthodon</emphasis>
</taxonomicName>
species.
</paragraph>
</caption>
<paragraph id="8BD63694FFDEFFE5FF613B11FD2BFCE2" pageId="5" pageNumber="188">
<table id="F969C434FFDE001FFF613B11FD33FCE2" box="[134,700,317,849]" gridcols="5" gridrows="10" pageId="5" pageNumber="188">
<tr id="355934D6FFDE001FFF613B11FD33FEE2" box="[134,700,317,337]" gridrow="0" pageId="5" pageNumber="188" rowspan-1="1">
<th id="76885DAAFFDE001FFF613B11FF4BFEE2" box="[134,196,317,337]" gridcol="0" gridrow="0" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FF613B11FF4BFEE2" bold="true" box="[134,196,317,337]" pageId="5" pageNumber="188">Locus</emphasis>
</th>
<th id="76885DAAFFDE001FFEA13B11FEDCFEE2" box="[326,339,317,337]" gridcol="2" gridrow="0" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FEA13B11FEDCFEE2" bold="true" box="[326,339,317,337]" pageId="5" pageNumber="188">n</emphasis>
</th>
<th id="76885DAAFFDE001FFE703B11FD85FEE2" box="[407,522,317,337]" gridcol="3" gridrow="0" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FE273B11FE6FFEE2" bold="true" box="[448,480,317,337]" pageId="5" pageNumber="188">OR</emphasis>
</th>
<th id="76885DAAFFDE001FFD663B11FD33FEE2" box="[641,700,317,337]" gridcol="4" gridrow="0" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FD633B11FD33FEE2" bold="true" box="[644,700,317,337]" pageId="5" pageNumber="188">Mean</emphasis>
</th>
</tr>
<tr id="355934D6FFDE001FFF613B4DFD33FE3E" box="[134,700,353,397]" gridrow="1" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFF613B4DFF4BFE3E" box="[134,196,353,397]" gridcol="0" gridrow="1" pageId="5" pageNumber="188">c</th>
<td id="76885DAAFFDE001FFEE13B4DFE95FE3E" box="[262,282,353,397]" gridcol="1" gridrow="1" pageId="5" pageNumber="188">L W</td>
<td id="76885DAAFFDE001FFEA13B4DFEDCFE3E" box="[326,339,353,397]" gridcol="2" gridrow="1" pageId="5" pageNumber="188">1 1</td>
<td id="76885DAAFFDE001FFE703B4DFD85FE3E" box="[407,522,353,397]" gridcol="3" gridrow="1" pageId="5" pageNumber="188">14.4 11.46</td>
<td id="76885DAAFFDE001FFD663B4DFD33FE3E" box="[641,700,353,397]" gridcol="4" gridrow="1" pageId="5" pageNumber="188">– –</td>
</tr>
<tr id="355934D6FFDE001FFF613BB0FD33FE7A" box="[134,700,412,457]" gridrow="2" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFF613BB0FF4BFE7A" box="[134,196,412,457]" gridcol="0" gridrow="2" pageId="5" pageNumber="188">p1</th>
<td id="76885DAAFFDE001FFEE13BB0FE95FE7A" box="[262,282,412,457]" gridcol="1" gridrow="2" pageId="5" pageNumber="188">L W</td>
<td id="76885DAAFFDE001FFEA13BB0FEDCFE7A" box="[326,339,412,457]" gridcol="2" gridrow="2" pageId="5" pageNumber="188">1 1</td>
<td id="76885DAAFFDE001FFE703BB0FD85FE7A" box="[407,522,412,457]" gridcol="3" gridrow="2" pageId="5" pageNumber="188">6.06 4.76</td>
<td id="76885DAAFFDE001FFD663BB0FD33FE7A" box="[641,700,412,457]" gridcol="4" gridrow="2" pageId="5" pageNumber="188">– –</td>
</tr>
<tr id="355934D6FFDE001FFF613BF4FD33FDB7" box="[134,700,472,516]" gridrow="3" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFF613BF4FF4BFDB7" box="[134,196,472,516]" gridcol="0" gridrow="3" pageId="5" pageNumber="188">p2</th>
<td id="76885DAAFFDE001FFEE13BF4FE95FDB7" box="[262,282,472,516]" gridcol="1" gridrow="3" pageId="5" pageNumber="188">L W</td>
<td id="76885DAAFFDE001FFEA13BF4FEDCFDB7" box="[326,339,472,516]" gridcol="2" gridrow="3" pageId="5" pageNumber="188">1 1</td>
<td id="76885DAAFFDE001FFE703BF4FD85FDB7" box="[407,522,472,516]" gridcol="3" gridrow="3" pageId="5" pageNumber="188">9.32 5.66</td>
<td id="76885DAAFFDE001FFD663BF4FD33FDB7" box="[641,700,472,516]" gridcol="4" gridrow="3" pageId="5" pageNumber="188">– –</td>
</tr>
<tr id="355934D6FFDE001FFF61383FFD33FDF3" box="[134,700,531,576]" gridrow="4" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFF61383FFF4BFDF3" box="[134,196,531,576]" gridcol="0" gridrow="4" pageId="5" pageNumber="188">p3</th>
<td id="76885DAAFFDE001FFEE1383FFE95FDF3" box="[262,282,531,576]" gridcol="1" gridrow="4" pageId="5" pageNumber="188">L W</td>
<td id="76885DAAFFDE001FFEA1383FFEDCFDF3" box="[326,339,531,576]" gridcol="2" gridrow="4" pageId="5" pageNumber="188">1 1</td>
<td id="76885DAAFFDE001FFE70383FFD85FDF3" box="[407,522,531,576]" gridcol="3" gridrow="4" pageId="5" pageNumber="188">9.67 5.92</td>
<td id="76885DAAFFDE001FFD66383FFD33FDF3" box="[641,700,531,576]" gridcol="4" gridrow="4" pageId="5" pageNumber="188">– –</td>
</tr>
<tr id="355934D6FFDE001FFF613863FD33FDC8" box="[134,700,591,635]" gridrow="5" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFF613863FF4BFDC8" box="[134,196,591,635]" gridcol="0" gridrow="5" pageId="5" pageNumber="188">p4</th>
<td id="76885DAAFFDE001FFEE13863FE95FDC8" box="[262,282,591,635]" gridcol="1" gridrow="5" pageId="5" pageNumber="188">L W</td>
<td id="76885DAAFFDE001FFEA13863FEDCFDC8" box="[326,339,591,635]" gridcol="2" gridrow="5" pageId="5" pageNumber="188">1 1</td>
<td id="76885DAAFFDE001FFE703863FD85FDC8" box="[407,522,591,635]" gridcol="3" gridrow="5" pageId="5" pageNumber="188">11.81 6.85</td>
<td id="76885DAAFFDE001FFD663863FD33FDC8" box="[641,700,591,635]" gridcol="4" gridrow="5" pageId="5" pageNumber="188">– –</td>
</tr>
<tr id="355934D6FFDE001FFF6138A6FD33FD04" box="[134,700,650,695]" gridrow="6" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFF6138A6FF4BFD04" box="[134,196,650,695]" gridcol="0" gridrow="6" pageId="5" pageNumber="188">m1</th>
<td id="76885DAAFFDE001FFEE138A6FE95FD04" box="[262,282,650,695]" gridcol="1" gridrow="6" pageId="5" pageNumber="188">L W</td>
<td id="76885DAAFFDE001FFEA138A6FEDCFD04" box="[326,339,650,695]" gridcol="2" gridrow="6" pageId="5" pageNumber="188">1 1</td>
<td id="76885DAAFFDE001FFE7038A6FD85FD04" box="[407,522,650,695]" gridcol="3" gridrow="6" pageId="5" pageNumber="188">11.59 6.22</td>
<td id="76885DAAFFDE001FFD6638A6FD33FD04" box="[641,700,650,695]" gridcol="4" gridrow="6" pageId="5" pageNumber="188">– –</td>
</tr>
<tr id="355934D6FFDE001FFF6138EAFD33FD41" box="[134,700,710,754]" gridrow="7" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFF6138EAFF4BFD41" box="[134,196,710,754]" gridcol="0" gridrow="7" pageId="5" pageNumber="188">m2</th>
<td id="76885DAAFFDE001FFEE138EAFE95FD41" box="[262,282,710,754]" gridcol="1" gridrow="7" pageId="5" pageNumber="188">L W</td>
<td id="76885DAAFFDE001FFEA138EAFEDCFD41" box="[326,339,710,754]" gridcol="2" gridrow="7" pageId="5" pageNumber="188">2 2</td>
<td id="76885DAAFFDE001FFE7038EAFD85FD41" box="[407,522,710,754]" gridcol="3" gridrow="7" pageId="5" pageNumber="188">12.56-12.89 7.33-8.14</td>
<td id="76885DAAFFDE001FFD6638EAFD33FD41" box="[641,700,710,754]" gridcol="4" gridrow="7" pageId="5" pageNumber="188">12.75 7.74</td>
</tr>
<tr id="355934D6FFDE001FFF61392EFD33FC9D" box="[134,700,770,814]" gridrow="8" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFF61392EFF4BFC9D" box="[134,196,770,814]" gridcol="0" gridrow="8" pageId="5" pageNumber="188">m3</th>
<td id="76885DAAFFDE001FFEE1392EFE95FC9D" box="[262,282,770,814]" gridcol="1" gridrow="8" pageId="5" pageNumber="188">L W</td>
<td id="76885DAAFFDE001FFEA1392EFEDCFC9D" box="[326,339,770,814]" gridcol="2" gridrow="8" pageId="5" pageNumber="188">2 2</td>
<td id="76885DAAFFDE001FFE70392EFD85FC9D" box="[407,522,770,814]" gridcol="3" gridrow="8" pageId="5" pageNumber="188">12.31-13.00 7.88-8.26</td>
<td id="76885DAAFFDE001FFD66392EFD33FC9D" box="[641,700,770,814]" gridcol="4" gridrow="8" pageId="5" pageNumber="188">12.66 8.07</td>
</tr>
<tr id="355934D6FFDE001FFF613911FD33FCE2" box="[134,700,829,849]" gridrow="9" pageId="5" pageNumber="188" rowspan-1="1">
<th id="76885DAAFFDE001FFF613911FF4BFCE2" box="[134,196,829,849]" gridcol="0" gridrow="9" pageId="5" pageNumber="188">MD</th>
<td id="76885DAAFFDE001FFEA13911FEDCFCE2" box="[326,339,829,849]" gridcol="2" gridrow="9" pageId="5" pageNumber="188">1</td>
<td id="76885DAAFFDE001FFE703911FD85FCE2" box="[407,522,829,849]" gridcol="3" gridrow="9" pageId="5" pageNumber="188">26.27</td>
<td id="76885DAAFFDE001FFD663911FD33FCE2" box="[641,700,829,849]" gridcol="4" gridrow="9" pageId="5" pageNumber="188">–</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD63694FFDEFFE5FCC93B11FA0FFDEF" pageId="5" pageNumber="188">
<table id="F969C434FFDE001FFCC93B11FA12FDEF" box="[814,1437,317,604]" gridcols="4" gridrows="10" pageId="5" pageNumber="188">
<tr id="355934D6FFDE001FFCC93B11FA12FEE2" box="[814,1437,317,337]" gridrow="0" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFCC93B11FC6EFEE2" box="[814,993,317,337]" gridcol="0" gridrow="0" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FCC93B11FCF0FEE2" bold="true" box="[814,895,317,337]" pageId="5" pageNumber="188">Species</emphasis>
</th>
<th id="76885DAAFFDE001FFBF33B11FBF0FEE2" box="[1044,1151,317,337]" gridcol="1" gridrow="0" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FBDA3B11FBD9FEE2" bold="true" box="[1085,1110,317,337]" pageId="5" pageNumber="188">p2</emphasis>
</th>
<th id="76885DAAFFDE001FFB443B11FA81FEE2" box="[1187,1294,317,337]" gridcol="2" gridrow="0" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FB2B3B11FB6AFEE2" bold="true" box="[1228,1253,317,337]" pageId="5" pageNumber="188">p3</emphasis>
</th>
<th id="76885DAAFFDE001FFAD43B11FA12FEE2" box="[1331,1437,317,337]" gridcol="3" gridrow="0" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FABB3B11FAFBFEE2" bold="true" box="[1372,1396,317,337]" pageId="5" pageNumber="188">p4</emphasis>
</th>
</tr>
<tr id="355934D6FFDE001FFCC93B4DFA12FEC6" box="[814,1437,353,373]" gridrow="1" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFCC93B4DFC6EFEC6" box="[814,993,353,373]" gridcol="0" gridrow="1" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FCC93B4DFC59FEC6" box="[814,982,353,373]" italics="true" pageId="5" pageNumber="188">
<taxonomicName id="4C694D17FFDEFFE5FCC93B4DFC19FEC6" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[814,918,353,373]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
menui
</emphasis>
</th>
<td id="76885DAAFFDE001FFBF33B4DFBF0FEC6" box="[1044,1151,353,373]" gridcol="1" gridrow="1" pageId="5" pageNumber="188">0.65</td>
<td id="76885DAAFFDE001FFB443B4DFA81FEC6" box="[1187,1294,353,373]" gridcol="2" gridrow="1" pageId="5" pageNumber="188">0.66</td>
<td id="76885DAAFFDE001FFAD43B4DFA12FEC6" box="[1331,1437,353,373]" gridcol="3" gridrow="1" pageId="5" pageNumber="188">0.6</td>
</tr>
<tr id="355934D6FFDE001FFCC93B55FA12FE3E" box="[814,1437,377,397]" gridrow="2" pageId="5" pageNumber="188" rowspan-0="1">
<td id="76885DAAFFDE001FFBF33B55FBF0FE3E" box="[1044,1151,377,397]" gridcol="1" gridrow="2" pageId="5" pageNumber="188">(OR: 0.63-</td>
<td id="76885DAAFFDE001FFB443B55FA81FE3E" box="[1187,1294,377,397]" gridcol="2" gridrow="2" pageId="5" pageNumber="188">(OR: 0.65-</td>
<td id="76885DAAFFDE001FFAD43B55FA12FE3E" box="[1331,1437,377,397]" gridcol="3" gridrow="2" pageId="5" pageNumber="188">(OR: 0.57-</td>
</tr>
<tr id="355934D6FFDE001FFCC93BBFFA12FE14" box="[814,1437,403,423]" gridrow="3" pageId="5" pageNumber="188" rowspan-0="1">
<td id="76885DAAFFDE001FFBF33BBFFBF0FE14" box="[1044,1151,403,423]" gridcol="1" gridrow="3" pageId="5" pageNumber="188">0.67, n=2)</td>
<td id="76885DAAFFDE001FFB443BBFFA81FE14" box="[1187,1294,403,423]" gridcol="2" gridrow="3" pageId="5" pageNumber="188">0.67, n=2)</td>
<td id="76885DAAFFDE001FFAD43BBFFA12FE14" box="[1331,1437,403,423]" gridcol="3" gridrow="3" pageId="5" pageNumber="188">0.63, n=2)</td>
</tr>
<tr id="355934D6FFDE001FFCC93B9AFA12FE79" box="[814,1437,438,458]" gridrow="4" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFCC93B9AFC6EFE79" box="[814,993,438,458]" gridcol="0" gridrow="4" pageId="5" pageNumber="188">
<emphasis id="B91DEA86FFDEFFE5FCC93B9AFC58FE79" box="[814,983,438,458]" italics="true" pageId="5" pageNumber="188">
<taxonomicName id="4C694D17FFDEFFE5FCC93B9AFC19FE79" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[814,918,438,458]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
tritens
</emphasis>
</th>
<td id="76885DAAFFDE001FFBF33B9AFBF0FE79" box="[1044,1151,438,458]" gridcol="1" gridrow="4" pageId="5" pageNumber="188">0.71</td>
<td id="76885DAAFFDE001FFB443B9AFA81FE79" box="[1187,1294,438,458]" gridcol="2" gridrow="4" pageId="5" pageNumber="188">0.67</td>
<td id="76885DAAFFDE001FFAD43B9AFA12FE79" box="[1331,1437,438,458]" gridcol="3" gridrow="4" pageId="5" pageNumber="188">0.65</td>
</tr>
<tr id="355934D6FFDE001FFCC93BE2FA12FE51" box="[814,1437,462,482]" gridrow="5" pageId="5" pageNumber="188" rowspan-0="1">
<td id="76885DAAFFDE001FFBF33BE2FBF0FE51" box="[1044,1151,462,482]" gridcol="1" gridrow="5" pageId="5" pageNumber="188">(OR: 0.69-</td>
<td id="76885DAAFFDE001FFB443BE2FA81FE51" box="[1187,1294,462,482]" gridcol="2" gridrow="5" pageId="5" pageNumber="188">(OR: 0.54-</td>
<td id="76885DAAFFDE001FFAD43BE2FA12FE51" box="[1331,1437,462,482]" gridcol="3" gridrow="5" pageId="5" pageNumber="188">(OR: 0.59-</td>
</tr>
<tr id="355934D6FFDE001FFCC93BC4FA12FE4F" box="[814,1437,488,508]" gridrow="6" pageId="5" pageNumber="188" rowspan-0="1">
<td id="76885DAAFFDE001FFBF33BC4FBF0FE4F" box="[1044,1151,488,508]" gridcol="1" gridrow="6" pageId="5" pageNumber="188">0.73, n = 4)</td>
<td id="76885DAAFFDE001FFB443BC4FA81FE4F" box="[1187,1294,488,508]" gridcol="2" gridrow="6" pageId="5" pageNumber="188">0.75, n = 5)</td>
<td id="76885DAAFFDE001FFAD43BC4FA12FE4F" box="[1331,1437,488,508]" gridcol="3" gridrow="6" pageId="5" pageNumber="188">0.68, n = 4)</td>
</tr>
<tr id="355934D6FFDE001FFCC93827FA12FDAC" box="[814,1437,523,543]" gridrow="7" pageId="5" pageNumber="188">
<th id="76885DAAFFDE001FFCC93827FC6EFDAC" box="[814,993,523,543]" gridcol="0" gridrow="7" pageId="5" pageNumber="188">
<taxonomicName id="4C694D17FFDEFFE5FCC93827FC6EFDAC" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[814,993,523,543]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="188" phylum="Chordata" rank="species" species="peignei">
<emphasis id="B91DEA86FFDEFFE5FCC93827FC6EFDAC" box="[814,993,523,543]" italics="true" pageId="5" pageNumber="188">Matthodon peignei</emphasis>
</taxonomicName>
</th>
<td id="76885DAAFFDE001FFBF33827FBF0FDAC" box="[1044,1151,523,543]" gridcol="1" gridrow="7" pageId="5" pageNumber="188">0.61</td>
<td id="76885DAAFFDE001FFB443827FA81FDAC" box="[1187,1294,523,543]" gridcol="2" gridrow="7" pageId="5" pageNumber="188">0.61</td>
<td id="76885DAAFFDE001FFAD43827FA12FDAC" box="[1331,1437,523,543]" gridcol="3" gridrow="7" pageId="5" pageNumber="188">0.58</td>
</tr>
<tr id="355934D6FFDE001FFCC93809FA12FD8A" box="[814,1437,549,569]" gridrow="8" pageId="5" pageNumber="188" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="76885DAAFFDE001FFCC93809FC6EFD8A" box="[814,993,549,569]" gridcol="0" gridrow="8" pageId="5" pageNumber="188">n. sp.</th>
</tr>
<tr id="355934D6FFDE001FFCC93864FA12FDEF" box="[814,1437,584,604]" gridrow="9" pageId="5" pageNumber="188" rowspan-0="1">
<td id="76885DAAFFDE001FFBF33864FBF0FDEF" box="[1044,1151,584,604]" gridcol="1" gridrow="9" pageId="5" pageNumber="188">(n=1)</td>
<td id="76885DAAFFDE001FFB443864FA81FDEF" box="[1187,1294,584,604]" gridcol="2" gridrow="9" pageId="5" pageNumber="188">(n=1)</td>
<td id="76885DAAFFDE001FFAD43864FA12FDEF" box="[1331,1437,584,604]" gridcol="3" gridrow="9" pageId="5" pageNumber="188">(n=1)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD63694FFDEFFE5FF6339BAFE9AFC1D" blockId="5.[132,776,918,2030]" box="[132,277,918,944]" pageId="5" pageNumber="188">DESCRIPTION</paragraph>
<paragraph id="8BD63694FFDEFFE5FF63399BFA88FD61" blockId="5.[132,776,918,2030]" lastBlockId="5.[813,1455,664,722]" pageId="5" pageNumber="188">
UM-AUM 694 is almost a complete mandible that bears well-preserved teeth, except the i3, canine and p1, which are represented by their alveoli (
<figureCitation id="13522A11FFDEFFE5FE4C39DAFE69FBA2" box="[427,486,1014,1041]" captionStart="FIG" captionStartId="6.[132,143,1962,1979]" captionTargetBox="[132,1455,215,1878]" captionTargetId="figure@6.[132,1455,215,1878]" captionTargetPageId="6" captionText="FIG. 1. — Matthodon peignei n. sp. from Aumelas (MP10-MP11?): A-C, UM-AUM694 (Holotype), left mandible bearing the p2-m3 and the alveoli of the canine and p1; A, occlusal view; B, lingual view; C, labial view; D-F, UM-AUM268, left mandible bearing the m2 and m3; D, occlusal view; E, lingual view; F, labial view. Scale bars: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3894812" httpUri="https://zenodo.org/record/3894812/files/figure.png" pageId="5" pageNumber="188">Fig. 1</figureCitation>
A-C). The mandible is deep (
<quantity id="4C919B71FFDEFFE5FF6B3E3AFF7EFB82" box="[140,241,1046,1073]" metricMagnitude="-2" metricUnit="m" metricValue="2.63" pageId="5" pageNumber="188" unit="mm" value="26.3">26.3 mm</quantity>
below the m1). Two mental foramina are present: the first one opens below the posterior root of p2, while the second is present below the anterior root of p4. The masseteric fossa is very deep. A deep fossa for insertion of the temporal muscle is present along the anterior margin of the crest. The angular process is curved sharply dorsally and is relatively wide. The mandibular condyle is robust, and is mediolaterally wide. The presence of an incisor (possibly the i3) is indicated by its tiny alveolus in front of the canine. The canine is buccolingually wide. The canine, premolars and molars are very close together. The p1 is single-rooted. The teeth display crenulated enamel.The p2 is simple with only the hypoconid present on the talonid. A small fold indicates the position of the paraconid. The p3 is mesiodistally longer and buccolingually wider than the p2. It differs by the presence of a paraconid, and by having three cuspids on the talonid: the larger corresponds to the hypoconid, while the other two are situated in the position of the entoconid. The p4 is more symmetrical in lateral view than the p3. The paraconid is much more developed. On the talonid, two cuspids are present: the hypoconid and the entoconid, which form a postfossid. The three molars display similar morphologies. The m3 is slightly mesiodistally shorter than the m2, but this is due to the reduction of the talonid. The paraconid is prominent and located mesially. The anterior keel is well developed. The metaconid is lower than the paraconid and almost completely fused with the protoconid. The latter is distinctly the highest cusp. The talonid is mesiodistally short and narrow. The postfossid is narrow and surrounded by three bulbous cusps: entoconid, hypoconid and hypoconulid; the entoconid being the tallest one. The m3 differs from m1 and m 
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having a much more reduced talonid in which the talonid cusps can- not be distinguished. The precingulid is short (limited to the paraconid) but is very wide, notably on m3.
</paragraph>
</subSubSection>
<subSubSection id="C373651FFFDEFFE7FCCA38DBFDC6FBA2" lastPageId="7" lastPageNumber="190" pageId="5" pageNumber="188" type="discussion">
<paragraph id="8BD63694FFDEFFE5FCCA38DBFC21FCBD" blockId="5.[813,1456,759,2030]" box="[813,942,759,785]" pageId="5" pageNumber="188">DISCUSSION</paragraph>
<paragraph id="8BD63694FFDEFFE5FCCA393BFAA5FBA2" blockId="5.[813,1456,759,2030]" pageId="5" pageNumber="188">
The new fossils share with 
<taxonomicName id="4C694D17FFDEFFE5FBBD393BFB5FFC82" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[1114,1232,791,817]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDEFFE5FBBD393BFB5FFC82" box="[1114,1232,791,817]" italics="true" pageId="5" pageNumber="188">Matthodon</emphasis>
</taxonomicName>
species the peculiar distal positions of the mental foramina (the first is inferior to the distal margin of p1 and the second is inferior to p4), the presence of a single-rooted p1, a closely packed toothrow, and the poorly developed metaconid on molars. All these features distinguish 
<taxonomicName id="4C694D17FFDEFFE5FBE3399BFBF7FC62" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[1028,1144,951,977]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDEFFE5FBE3399BFBF7FC62" box="[1028,1144,951,977]" italics="true" pageId="5" pageNumber="188">Matthodon</emphasis>
</taxonomicName>
from the European Ypresian and Lutetian hyaenodonts. UM-AUM694 is among the best preserved mandibles ever found for 
<taxonomicName id="4C694D17FFDEFFE5FB4839DAFAACFBA3" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[1199,1315,1014,1040]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDEFFE5FB4839DAFAACFBA3" box="[1199,1315,1014,1040]" italics="true" pageId="5" pageNumber="188">Matthodon</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD63694FFDEFFE5FCA43E3AFAB4FB23" blockId="5.[813,1456,759,2030]" pageId="5" pageNumber="188">
The genus 
<taxonomicName id="4C694D17FFDEFFE5FC5E3E3AFBA2FB83" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[953,1069,1046,1072]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDEFFE5FC5E3E3AFBA2FB83" box="[953,1069,1046,1072]" italics="true" pageId="5" pageNumber="188">Matthodon</emphasis>
</taxonomicName>
is represented by two species: 
<emphasis id="B91DEA86FFDEFFE5FA9E3E3BFC36FBE3" italics="true" pageId="5" pageNumber="188">
<taxonomicName id="4C694D17FFDEFFE5FA9E3E3BFCFBFBE3" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
menui
</emphasis>
from Chavot, Monthelon, Mancy, and Grauves (Ypresian, MP10; 
<collectingCountry id="F37E7604FFDEFFE5FC013E7BFBA3FBC2" box="[998,1068,1111,1137]" name="France" pageId="5" pageNumber="188">France</collectingCountry>
) and 
<emphasis id="B91DEA86FFDEFFE5FB843E7AFA9DFBC3" box="[1123,1298,1110,1136]" italics="true" pageId="5" pageNumber="188">
<taxonomicName id="4C694D17FFDEFFE5FB843E7AFB5DFBC3" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[1123,1234,1110,1136]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
tritens
</emphasis>
from Geiseltal- Untere Unterkohle (Lutetian, MP11; 
<collectingCountry id="F37E7604FFDEFFE5FB223E5AFABFFB23" box="[1221,1328,1142,1168]" name="Germany" pageId="5" pageNumber="188">Germany</collectingCountry>
).
</paragraph>
<paragraph id="8BD63694FFDEFFE5FCA43EBAFA0AFAC3" blockId="5.[813,1456,759,2030]" pageId="5" pageNumber="188">
The fossils from Aumelas differ from those of the Lutetian species, 
<emphasis id="B91DEA86FFDEFFE5FC983E9AFBBCFB7C" box="[895,1075,1206,1232]" italics="true" pageId="5" pageNumber="188">
<taxonomicName id="4C694D17FFDEFFE5FC983E9AFC7FFB63" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[895,1008,1206,1232]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
tritens
</emphasis>
, by having less developed paraconids on p2, p3 and p4; the larger and more individualized metaconid on the molars; the longer talonid on the m1 and m2; and by shorter molars (4% for the m1, 8% for the m2 and m3), and longer p2 and p4 (respectively 9% and 5%). These differences are reminiscent of the Ypresian species 
<emphasis id="B91DEA86FFDEFFE5FB213F79FAF1FADC" box="[1222,1406,1365,1391]" italics="true" pageId="5" pageNumber="188">
<taxonomicName id="4C694D17FFDEFFE5FB213F79FAB8FADC" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[1222,1335,1365,1391]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
menui
</emphasis>
.
</paragraph>
<paragraph id="8BD63694FFDEFFE5FCA43F59FA20F9FC" blockId="5.[813,1456,759,2030]" pageId="5" pageNumber="188">
The fossils from Aumelas, are distinct from those referred to 
<emphasis id="B91DEA86FFDEFFE5FCCA3FB9FC6AFA1C" box="[813,997,1429,1455]" italics="true" pageId="5" pageNumber="188">
<taxonomicName id="4C694D17FFDEFFE5FCCA3FB9FC2FFA1C" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[813,928,1429,1455]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="210" phylum="Chordata" rank="genus">Matthodon</taxonomicName>
menui
</emphasis>
by having longer premolars and molars: the molars are 6% longer, the p2 9% longer, and the p3 and p4 4% longer. The mesio-distal diameter of the alveolus of the p1 is also distinctly longer than those measured for 
<emphasis id="B91DEA86FFDEFFE5FAC43FD9FA07F9BD" box="[1315,1416,1525,1550]" italics="true" pageId="5" pageNumber="188">M. menui</emphasis>
. As indicated above, the teeth from Aumelas are morphologically more similar to those of 
<emphasis id="B91DEA86FFDEFFE5FBCB3C19FB1CF9FD" box="[1068,1171,1589,1614]" italics="true" pageId="5" pageNumber="188">M. menui</emphasis>
than to those of 
<emphasis id="B91DEA86FFDEFFE5FAA33C19FA27F9FD" box="[1348,1448,1589,1614]" italics="true" pageId="5" pageNumber="188">M. tritens</emphasis>
.
</paragraph>
<paragraph id="8BD63694FFDEFFE5FCA43C78FB50F91C" blockId="5.[813,1456,759,2030]" pageId="5" pageNumber="188">
The mean width/length ratio of the p2, p3 and p4 differentiates 
<taxonomicName id="4C694D17FFDEFFE5FC8C3C58FBA2F93D" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[875,1069,1652,1678]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="5" pageNumber="188" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="B91DEA86FFDEFFE5FC8C3C58FBA2F93D" box="[875,1069,1652,1678]" italics="true" pageId="5" pageNumber="188">Matthodon peignei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDEFFE5FBD43C59FBE2F93C" box="[1075,1133,1653,1679]" pageId="5" pageNumber="188" rank="species">n. sp.</taxonomicNameLabel>
from 
<emphasis id="B91DEA86FFDEFFE5FB4A3C59FA9AF93D" box="[1197,1301,1653,1678]" italics="true" pageId="5" pageNumber="188">M. menui</emphasis>
and 
<emphasis id="B91DEA86FFDEFFE5FAAE3C59FA20F93D" box="[1353,1455,1653,1678]" italics="true" pageId="5" pageNumber="188">M. tritens</emphasis>
by having narrower premolars (
<tableCitation id="C6EB032FFFDEFFE5FB983CB8FB40F91D" box="[1151,1231,1684,1710]" captionStart="TABLE" captionStartId="5.[813,824,219,236]" captionText="TABLE 2. — Mean width/length ratio of the p2, p3 and p4 respectively in the Matthodon species." pageId="5" pageNumber="188">Table 2</tableCitation>
).
</paragraph>
<paragraph id="8BD63694FFDEFFE5FCA43C98FA9AF85E" blockId="5.[813,1456,759,2030]" pageId="5" pageNumber="188">
The fossils from Aumelas appear intermediate both metrically and morphologically between Ypresian 
<emphasis id="B91DEA86FFDEFFE5FAF23CF9FAF0F95D" box="[1301,1407,1749,1774]" italics="true" pageId="5" pageNumber="188">M. menui</emphasis>
and Lutetian 
<emphasis id="B91DEA86FFDEFFE5FC6D3CD9FC63F8BD" box="[906,1004,1781,1806]" italics="true" pageId="5" pageNumber="188">M. tritens</emphasis>
. It differs from the two previously described taxa by the relatively large size of the p2. This feature is highly characteristic because the relative size of the p2, which decreases from 
<emphasis id="B91DEA86FFDEFFE5FC363D78FBB5F8DE" box="[977,1082,1876,1901]" italics="true" pageId="5" pageNumber="188">M. menui</emphasis>
to 
<emphasis id="B91DEA86FFDEFFE5FBB93D78FB4AF8DE" box="[1118,1221,1876,1901]" italics="true" pageId="5" pageNumber="188">M. tritens</emphasis>
, can be considered as an indication of the reduction of the anterior dentition (p1 and p2). The larger size of the p 
<quantity id="4C919B71FFDEFFE5FB9D3DBFFB2BF81E" box="[1146,1188,1939,1965]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="188" unit="in" value="2.0">2 in</quantity>
UM/AUM 694 thus can be regarded as a primitive feature. The same observation can be made for the narrowness of the premolars.
</paragraph>
<caption id="DF16661CFFDDFFE6FF633D86FE98F85A" ID-DOI="http://doi.org/10.5281/zenodo.3894812" ID-Zenodo-Dep="3894812" httpUri="https://zenodo.org/record/3894812/files/figure.png" pageId="6" pageNumber="189" startId="6.[132,143,1962,1979]" targetBox="[132,1455,215,1878]" targetPageId="6">
<paragraph id="8BD63694FFDDFFE6FF633D86FE98F85A" blockId="6.[132,1457,1962,2025]" pageId="6" pageNumber="189">
<subScript id="17ED34D1FFDDFFE6FF633D86FF12F808" attach="left" box="[132,157,1962,1979]" fontSize="5" pageId="6" pageNumber="189">FIG</subScript>
. 1. — 
<taxonomicName id="4C694D17FFDDFFE6FF323D86FEFDF808" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[213,370,1962,1979]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="6" pageNumber="189" phylum="Chordata" rank="species" species="peignei" status="sp. nov.">
<emphasis id="B91DEA86FFDDFFE6FF323D86FEFDF808" box="[213,370,1962,1979]" italics="true" pageId="6" pageNumber="189">Matthodon peignei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDDFFE6FE9F3D86FE26F808" box="[376,425,1962,1979]" pageId="6" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
from Aumelas (MP10-MP11?): 
<emphasis id="B91DEA86FFDDFFE6FD483D86FD5EF808" bold="true" box="[687,721,1962,1979]" pageId="6" pageNumber="189">A -C</emphasis>
, UM-AUM694 (Holotype), left mandible bearing the p2-m3 and the alveoli of the canine and p1; 
<emphasis id="B91DEA86FFDDFFE6FF2F3DEDFF5AF861" bold="true" box="[200,213,1985,2002]" pageId="6" pageNumber="189">A</emphasis>
, occlusal view; 
<emphasis id="B91DEA86FFDDFFE6FEBF3DEDFEEAF861" bold="true" box="[344,357,1985,2002]" pageId="6" pageNumber="189">B</emphasis>
, lingual view; 
<emphasis id="B91DEA86FFDDFFE6FE3E3DEDFE68F861" bold="true" box="[473,487,1985,2002]" pageId="6" pageNumber="189">C</emphasis>
, labial view; 
<emphasis id="B91DEA86FFDDFFE6FDB63DEDFDFEF861" bold="true" box="[593,625,1985,2002]" pageId="6" pageNumber="189">D -F</emphasis>
, UM-AUM268, left mandible bearing the m2 and m3; 
<emphasis id="B91DEA86FFDDFFE6FBCA3DEDFBB4F861" bold="true" box="[1069,1083,1985,2002]" pageId="6" pageNumber="189">D</emphasis>
, occlusal view; 
<emphasis id="B91DEA86FFDDFFE6FB593DEDFB45F861" bold="true" box="[1214,1226,1985,2002]" pageId="6" pageNumber="189">E</emphasis>
, lingual view; 
<emphasis id="B91DEA86FFDDFFE6FAD83DEDFAC5F861" bold="true" box="[1343,1354,1985,2002]" pageId="6" pageNumber="189">F</emphasis>
, labial view. Scale bars: 1 cm.
</paragraph>
</caption>
<caption id="DF16661CFFDCFFE7FF633AF7FEFAFEA9" pageId="7" pageNumber="190">
<paragraph id="8BD63694FFDCFFE7FF633AF7FEFAFEA9" blockId="7.[132,775,219,282]" pageId="7" pageNumber="190">
<subScript id="17ED34D1FFDCFFE7FF633AF7FF3DFF5F" attach="left" box="[132,178,219,236]" fontSize="5" pageId="7" pageNumber="190">TABLE</subScript>
3. — Measurements (in mm) of 
<taxonomicName id="4C694D17FFDCFFE7FE263AF7FD2AFF5F" authority="Solé &amp; Marandat &amp; Lihoreau, 2020" authorityName="Solé &amp; Marandat &amp; Lihoreau" authorityYear="2020" box="[449,677,219,236]" class="Mammalia" family="Hyaenodontidae" genus="Oxyaenoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Creodonta" pageId="7" pageNumber="190" phylum="Chordata" rank="species" species="aumelasiensis" status="sp. nov.">
<emphasis id="B91DEA86FFDCFFE7FE263AF7FD2AFF5F" box="[449,677,219,236]" italics="true" pageId="7" pageNumber="190">Oxyaenoides aumelasiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22E57FDFFDCFFE7FD4C3AF7FD53FF5F" box="[683,732,219,236]" pageId="7" pageNumber="190" rank="species">n. sp.</taxonomicNameLabel>
from Aumelas (MP10-MP11?). 
<emphasis id="B91DEA86FFDCFFE7FEB53ADEFED6FEB0" bold="true" box="[338,345,242,259]" pageId="7" pageNumber="190">*</emphasis>
, based on roots and/or alveoli;Body mass (
<emphasis id="B91DEA86FFDCFFE7FD523ADEFD4BFEB0" box="[693,708,242,259]" italics="true" pageId="7" pageNumber="190">c.</emphasis>
4.24 kg) estimated after 
<bibRefCitation id="EFF84B65FFDCFFE7FEE03B25FEFEFEA9" author="MORLO M." box="[263,369,265,282]" journalOrPublisher="Geobios" pageId="7" pageNumber="190" pagination="297 - 305" part="32" refId="ref15217" refString="MORLO M. 1999. - Niche structure and evolution in creodont (Mammalia) faunas of the European and North American Eocene. Geobios 32: 297 - 305. https: // doi. org / 10.1016 / S 0016 - 6995 (99) 80043 - 6" title="Niche structure and evolution in creodont (Mammalia) faunas of the European and North American Eocene" type="journal article" year="1999">Morlo (1999)</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="8BD63694FFDCFFE7FF613B78FD34FCDB" pageId="7" pageNumber="190">
<table id="F969C434FFDC001FFF613B78FD5EFCDB" box="[134,721,340,872]" gridcols="3" gridrows="10" pageId="7" pageNumber="190">
<tr id="355934D6FFDC001FFF613B78FD5EFEDB" box="[134,721,340,360]" gridrow="0" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF613B78FF4BFEDB" box="[134,196,340,360]" gridcol="0" gridrow="0" pageId="7" pageNumber="190">
<emphasis id="B91DEA86FFDCFFE7FF613B78FF4BFEDB" bold="true" box="[134,196,340,360]" pageId="7" pageNumber="190">Locus</emphasis>
</th>
<th id="76885DAAFFDC001FFEE13B78FDC5FEDB" box="[262,586,340,360]" gridcol="1" gridrow="0" pageId="7" pageNumber="190">
<emphasis id="B91DEA86FFDCFFE7FE853B78FDAFFEDB" bold="true" box="[354,544,340,360]" pageId="7" pageNumber="190">n OR</emphasis>
</th>
<th id="76885DAAFFDC001FFD7E3B78FD5EFEDB" box="[665,721,340,360]" gridcol="2" gridrow="0" pageId="7" pageNumber="190">
<emphasis id="B91DEA86FFDCFFE7FD7E3B78FD5EFEDB" bold="true" box="[665,721,340,360]" pageId="7" pageNumber="190">Mean</emphasis>
</th>
</tr>
<tr id="355934D6FFDC001FFF613B54FD5EFE17" box="[134,721,376,420]" gridrow="1" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF613B54FF4BFE17" box="[134,196,376,420]" gridcol="0" gridrow="1" pageId="7" pageNumber="190">P3</th>
<td id="76885DAAFFDC001FFEE13B54FDC5FE17" box="[262,586,376,420]" gridcol="1" gridrow="1" pageId="7" pageNumber="190">L 2 9.88-11.10 W 2 6.39-6.78</td>
<td id="76885DAAFFDC001FFD7E3B54FD5EFE17" box="[665,721,376,420]" gridcol="2" gridrow="1" pageId="7" pageNumber="190">10.49 6.59</td>
</tr>
<tr id="355934D6FFDC001FFF613B9FFD5EFE6C" box="[134,721,435,479]" gridrow="2" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF613B9FFF4BFE6C" box="[134,196,435,479]" gridcol="0" gridrow="2" pageId="7" pageNumber="190">P4</th>
<td id="76885DAAFFDC001FFEE13B9FFDC5FE6C" box="[262,586,435,479]" gridcol="1" gridrow="2" pageId="7" pageNumber="190">L 2 11.28-11.55 W 2 7.63-8.75</td>
<td id="76885DAAFFDC001FFD7E3B9FFD5EFE6C" box="[665,721,435,479]" gridcol="2" gridrow="2" pageId="7" pageNumber="190">11.42 8.19</td>
</tr>
<tr id="355934D6FFDC001FFF613BC3FD5EFDA8" box="[134,721,495,539]" gridrow="3" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF613BC3FF4BFDA8" box="[134,196,495,539]" gridcol="0" gridrow="3" pageId="7" pageNumber="190">M1</th>
<td id="76885DAAFFDC001FFEE13BC3FDC5FDA8" box="[262,586,495,539]" gridcol="1" gridrow="3" pageId="7" pageNumber="190">L 2 9.39-9.52 W 1 10.54</td>
<td id="76885DAAFFDC001FFD7E3BC3FD5EFDA8" box="[665,721,495,539]" gridcol="2" gridrow="3" pageId="7" pageNumber="190">9.46 –</td>
</tr>
<tr id="355934D6FFDC001FFF613806FD5EFDE5" box="[134,721,554,598]" gridrow="4" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF613806FF4BFDE5" box="[134,196,554,598]" gridcol="0" gridrow="4" pageId="7" pageNumber="190">M2</th>
<td id="76885DAAFFDC001FFEE13806FDC5FDE5" box="[262,586,554,598]" gridcol="1" gridrow="4" pageId="7" pageNumber="190">L 1 8.32 W 1 10.11</td>
<td id="76885DAAFFDC001FFD7E3806FD5EFDE5" box="[665,721,554,598]" gridcol="2" gridrow="4" pageId="7" pageNumber="190">– –</td>
</tr>
<tr id="355934D6FFDC001FFF61384AFD5EFD21" box="[134,721,614,658]" gridrow="5" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF61384AFF4BFD21" box="[134,196,614,658]" gridcol="0" gridrow="5" pageId="7" pageNumber="190">p2</th>
<td id="76885DAAFFDC001FFEE1384AFDC5FD21" box="[262,586,614,658]" gridcol="1" gridrow="5" pageId="7" pageNumber="190">L 1 5.88 W 1 2.31</td>
<td id="76885DAAFFDC001FFD7E384AFD5EFD21" box="[665,721,614,658]" gridcol="2" gridrow="5" pageId="7" pageNumber="190">– –</td>
</tr>
<tr id="355934D6FFDC001FFF61388DFD5EFD7E" box="[134,721,673,717]" gridrow="6" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF61388DFF4BFD7E" box="[134,196,673,717]" gridcol="0" gridrow="6" pageId="7" pageNumber="190">m1</th>
<td id="76885DAAFFDC001FFEE1388DFDC5FD7E" box="[262,586,673,717]" gridcol="1" gridrow="6" pageId="7" pageNumber="190">L 1 8.71* W 1 4.65*</td>
<td id="76885DAAFFDC001FFD7E388DFD5EFD7E" box="[665,721,673,717]" gridcol="2" gridrow="6" pageId="7" pageNumber="190">– –</td>
</tr>
<tr id="355934D6FFDC001FFF6138F1FD5EFCBA" box="[134,721,733,777]" gridrow="7" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF6138F1FF4BFCBA" box="[134,196,733,777]" gridcol="0" gridrow="7" pageId="7" pageNumber="190">m2</th>
<td id="76885DAAFFDC001FFEE138F1FDC5FCBA" box="[262,586,733,777]" gridcol="1" gridrow="7" pageId="7" pageNumber="190">L 1 7.74* W 1 5.16*</td>
<td id="76885DAAFFDC001FFD7E38F1FD5EFCBA" box="[665,721,733,777]" gridcol="2" gridrow="7" pageId="7" pageNumber="190">– –</td>
</tr>
<tr id="355934D6FFDC001FFF613934FD5EFCF7" box="[134,721,792,836]" gridrow="8" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF613934FF4BFCF7" box="[134,196,792,836]" gridcol="0" gridrow="8" pageId="7" pageNumber="190">m3</th>
<td id="76885DAAFFDC001FFEE13934FDC5FCF7" box="[262,586,792,836]" gridcol="1" gridrow="8" pageId="7" pageNumber="190">L 1 9.24 W 1 5.37</td>
<td id="76885DAAFFDC001FFD7E3934FD5EFCF7" box="[665,721,792,836]" gridcol="2" gridrow="8" pageId="7" pageNumber="190">– –</td>
</tr>
<tr id="355934D6FFDC001FFF613978FD5EFCDB" box="[134,721,852,872]" gridrow="9" pageId="7" pageNumber="190">
<th id="76885DAAFFDC001FFF613978FF4BFCDB" box="[134,196,852,872]" gridcol="0" gridrow="9" pageId="7" pageNumber="190">MD</th>
<td id="76885DAAFFDC001FFEE13978FDC5FCDB" box="[262,586,852,872]" gridcol="1" gridrow="9" pageId="7" pageNumber="190">1 16.68</td>
<td id="76885DAAFFDC001FFD7E3978FD5EFCDB" box="[665,721,852,872]" gridcol="2" gridrow="9" pageId="7" pageNumber="190">–</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD63694FFDCFFE7FF7C399BFDC6FBA2" blockId="7.[132,775,951,1041]" pageId="7" pageNumber="190">
Based on these comparisons the Aumelas specimen represents a new species of 
<taxonomicName id="4C694D17FFDCFFE7FE9639FAFE6AFC43" authorityName="Lange-Badre &amp; Haubold" authorityYear="1990" box="[369,485,982,1008]" class="Mammalia" genus="Matthodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Deltatheroida" pageId="7" pageNumber="210" phylum="Chordata" rank="genus">
<emphasis id="B91DEA86FFDCFFE7FE9639FAFE6AFC43" box="[369,485,982,1008]" italics="true" pageId="7" pageNumber="190">Matthodon</emphasis>
</taxonomicName>
, the first occurrence of the genus in the southwestern part of Europe.
</paragraph>
</subSubSection>
</treatment>
</document>