treatments-xml/data/03/F5/87/03F58791D32F0C06C9CF9FFEFA98FE4E.xml

<document id="4736E2C8D29455C2567D374088EEA39A" ID-DOI="10.5281/zenodo.191909" ID-GBIF-Dataset="b5235354-ef06-4a49-b9d1-5498e97c3639" ID-ISSN="1175-5326" ID-Zenodo-Dep="191909" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461137864308" checkinUser="plazi" docAuthor="Ng, Peter K. L., Rahayu, Dwi Listyo &amp; Naser, Murtada D." docDate="2009" docId="03F58791D32F0C06C9CF9FFEFA98FE4E" docLanguage="en" docName="zt02312p026.pdf" docOrigin="Zootaxa 2312" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptochryseus Al-Khayat &amp; Jones 1996" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="FFCCFFE9D3200C12C9589874FFC1FFD4" masterDocTitle="The Camptandriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-Khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura)" masterLastPageNumber="26" masterPageNumber="1" pageNumber="16" updateTime="1698598725934" updateUser="plazi">
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<mods:title id="EDE1300E3C5E170620D22FE63FC62A4C">The Camptandriidae of Iraq, with description of a new genus and notes on Leptochryseus Al-Khayat &amp; Jones, 1996 (Crustacea: Decapoda: Brachyura)</mods:title>
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<mods:namePart id="8D5A070ACC059B91C008127A40237E14">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="8CDCD61C729302430F21864EB3AB5E23">Rahayu, Dwi Listyo</mods:namePart>
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<subSubSection id="C346650CD32F0C1DC9CF9FFEFD1DF870" box="[151,732,1930,1956]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="8BE33687D32F0C1DC9CF9FFEFD1DF870" blockId="15.[151,732,1930,1956]" box="[151,732,1930,1956]" pageId="15" pageNumber="16">
<heading id="D0AB81EBD32F0C1DC9CF9FFEFD1DF870" bold="true" box="[151,732,1930,1956]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
<emphasis id="B928EA95D32F0C1DC9CF9FFEFD1DF870" bold="true" box="[151,732,1930,1956]" pageId="15" pageNumber="16">
Genus 
<taxonomicName id="4C5C4D04D32F0C1DC9B69FFEFD1DF870" ID-CoL="7PMN4" authority="Al-Khayat &amp; Jones, 1996" authorityName="Al-Khayat &amp; Jones" authorityYear="1996" box="[238,732,1930,1956]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D32F0C1DC9B69FFEFE5DF870" bold="true" box="[238,412,1930,1956]" italics="true" pageId="15" pageNumber="16">Leptochryseus</emphasis>
<bibRefCitation id="EFCD4B76D32F0C1DC8FB9FFEFD1DF870" author="Al-Khayat" box="[419,732,1930,1956]" pageId="15" pageNumber="16" refString="Al-Khayat, J. A. &amp; Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat &amp; Jones, 1996</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C346650CD32F0C02C9CF9FA0FC09FF7B" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="reference_group">
<paragraph id="8BE33687D32F0C1DC9CF9FA0FD44F83F" blockId="15.[151,645,2004,2027]" box="[151,645,2004,2027]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="AB4C11A9D32F0C1DC9CF9FA0FD44F83F" box="[151,645,2004,2027]" pageId="15" pageNumber="16">
<taxonomicName id="4C5C4D04D32F0C1DC9CF9FA0FD40F83F" ID-CoL="7PMN4" authority="Al-Khayat &amp; Jones, 1996: 798" authorityName="Al-Khayat &amp; Jones" authorityPageNumber="798" authorityYear="1996" box="[151,641,2004,2027]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D32F0C1DC9CF9FA0FEF0F83F" box="[151,305,2004,2027]" italics="true" pageId="15" pageNumber="16">Leptochryseus</emphasis>
<treatmentCitation id="0AFD1096D32F0C1DC86F9FA0FD40F83F" author="Al-Khayat" box="[311,641,2004,2027]" page="798" pageId="15" pageNumber="16" year="1996">
<bibRefCitation id="EFCD4B76D32F0C1DC86F9FA0FD8DF83F" author="Al-Khayat" box="[311,588,2004,2027]" pageId="15" pageNumber="16" refString="Al-Khayat, J. A. &amp; Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat &amp; Jones, 1996</bibRefCitation>
: 798
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.
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</paragraph>
<paragraph id="8BE33687D3300C02C9CF98ECFC09FF7B" blockId="16.[151,968,152,175]" box="[151,968,152,175]" pageId="16" pageNumber="17">
<taxonomicName id="4C5C4D04D3300C02C9CF98ECFEE2FF7B" ID-CoL="7NZVB" box="[151,291,152,175]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3300C02C9CF98ECFEE2FF7B" box="[151,291,152,175]" italics="true" pageId="16" pageNumber="17">Cleistostoma</emphasis>
</taxonomicName>
– Ng 
<emphasis id="B928EA95D3300C02C83C98ECFE5BFF7B" box="[356,410,152,175]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2008: 233 (part) (not 
<taxonomicName id="4C5C4D04D3300C02CBDD98ECFC7DFF7B" ID-CoL="7NZVB" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" box="[645,956,152,175]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3300C02CBDD98ECFCD1FF7B" box="[645,784,152,175]" italics="true" pageId="16" pageNumber="17">Cleistostoma</emphasis>
De Haan, 1833
</taxonomicName>
).
</paragraph>
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<subSubSection id="C346650CD3300C02C9CF98AAFBE6FF2C" box="[151,1063,222,248]" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BE33687D3300C02C9CF98AAFBE6FF2C" blockId="16.[151,1437,222,2027]" box="[151,1063,222,248]" pageId="16" pageNumber="17">
<emphasis id="B928EA95D3300C02C9CF98AAFEF8FF2C" bold="true" box="[151,313,222,248]" pageId="16" pageNumber="17">
<typeStatus id="54E78825D3300C02C9CF98AAFF15FF2C" box="[151,212,222,248]" pageId="16" pageNumber="17">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C5C4D04D3300C02C81898ABFCB8FF2C" authority="Jones &amp; Clayton, 1983" authorityName="Jones &amp; Clayton" authorityYear="1983" box="[320,889,222,248]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="kuwaitense">
<emphasis id="B928EA95D3300C02C81898ABFDA2FF2C" box="[320,611,223,248]" italics="true" pageId="16" pageNumber="17">Cleistostoma kuwaitense</emphasis>
<bibRefCitation id="EFCD4B76D3300C02CB3298AAFCB8FF2C" author="Jones" box="[618,889,222,248]" pageId="16" pageNumber="17" refString="Jones, D. A. &amp; Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983">Jones &amp; Clayton, 1983</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C346650CD3300C02C99E9971FDDFFAE7" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8BE33687D3300C02C99E9971FDDFFAE7" blockId="16.[151,1437,222,2027]" pageId="16" pageNumber="17">
<emphasis id="B928EA95D3300C02C99E9971FE87FECB" bold="true" box="[198,326,261,287]" pageId="16" pageNumber="17">Diagnosis.</emphasis>
Carapace average width to length ratio 1.32; quadrangular in outline, wider than long; dorsal surface moderately convex, without visible ridges dorsally, regions not well demarcated, epigastric region granular, gently convex, separated by longitudinal groove, gastro-cardiac groove shallow but visible (
<figureCitation id="13672A02D3300C02CC389926FF66FE47" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
A, 8, 11G). Anterolateral margin of carapace unarmed, gently tuberculate; dorsal surface posterior to external orbital tooth depressed; posterolateral margin convex, distinct dorsolateral margin present; posterolateral margin sinuous; posterior carapace margin gently sinuous, with distinct submarginal ridge of granules (
<figureCitation id="13672A02D3300C02C8529999FE9AFDD3" box="[266,347,493,519]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
A, 8, 11G). Front with anterior margin gently concave from dorsal view, entire; frontomedial portion not projected, margin vaguely bilobed (
<figureCitation id="13672A02D3300C02CB919A60FCDDFDFA" box="[713,796,532,558]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
A, 8, 11G). Supraorbital margin gently sinuous, notch separating it from base of front indistinct (
<figureCitation id="13672A02D3300C02CBC39A4EFD2EFD80" box="[667,751,570,596]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
A, 8, 11G); infraorbital margin receded, on orbital shelf, inner infraorbital tooth triangular, not separated from rest of margin by notch or fissure, suborbital ridge well developed, protruding beyond infraorbital margin, visible dorsally, suborbital sulcus adjacent to suborbital ridge, separating infraorbital margin from suborbital ridge. Epistome longitudinally narrow, interantennular septum broad, posterior medial tooth short, broad (
<figureCitation id="13672A02D3300C02CA799AA1FCBAFD3B" box="[801,891,725,751]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
B, 11H). Pterygostomian region with deep, relatively broad Y-shaped sulcus. Third maxilliped merus with outer margin rounded, expanded beyond outer margin of ischium, inner margins thick; merus with anteroexternal margin produced to form small lobe; inverted, distinct V-shaped groove on outer part; ischium with inner distal angle produced, with row of submarginal setae along distal margin; outer surface of basal segment of palp concave, forming spatulate structure (
<figureCitation id="13672A02D3300C02C84A9BE2FEABFC64" box="[274,362,918,944]" captionStart="FIGURE 11" captionStartId="23.[151,257,1868,1892]" captionTargetBox="[156,1409,201,1868]" captionTargetId="figure@23.[156,1430,171,1886]" captionTargetPageId="23" captionText="FIGURE 11. Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000.9), China; Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait. A, G, carapaces; B, H, frontal views of carapaces showing orbits, eyes and epistomes; C, outer view of right third maxilliped; D, outer view of right chela; J, outer view of left chela; I, outer view of left third maxilliped; E, K, left G 1; F, L, distal part of left G 1. Scales: A, B, G – J = 2.0 mm; C, D, E, K = 1.0 mm; F, L = 0.5 mm." httpUri="https://zenodo.org/record/191919/files/figure.png" pageId="16" pageNumber="17">
Fig. 
<date id="FFE21047D3300C02C8139BE2FEABFC64" box="[331,362,918,944]" pageId="16" pageNumber="17" value="1911-01">11</date>
</figureCitation>
I). Chelipeds equal; sexually dimorphic in large adult males; dactylus of male chela with distinct molariform tooth on cutting edge; finger tips in both sexes spatulate 
<figureCitation id="13672A02D3300C02CD679BC9FB54FC03" box="[1087,1173,957,983]" captionStart="FIGURE 11" captionStartId="23.[151,257,1868,1892]" captionTargetBox="[156,1409,201,1868]" captionTargetId="figure@23.[156,1430,171,1886]" captionTargetPageId="23" captionText="FIGURE 11. Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000.9), China; Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait. A, G, carapaces; B, H, frontal views of carapaces showing orbits, eyes and epistomes; C, outer view of right third maxilliped; D, outer view of right chela; J, outer view of left chela; I, outer view of left third maxilliped; E, K, left G 1; F, L, distal part of left G 1. Scales: A, B, G – J = 2.0 mm; C, D, E, K = 1.0 mm; F, L = 0.5 mm." httpUri="https://zenodo.org/record/191919/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
J), more developed in females. Ambulatory legs relatively broad; ventral margins of P3, P4 meri with sharp spines; P2–5 meri laterally flattened, expanded, appears foliaceous in smaller males, females; large males with disproportionately long P3, merus, propodus very long, slender, ventral margin of dactylus with tuft of setae (
<figureCitation id="13672A02D3300C02C9C79C2CFF33FBA6" box="[159,242,1112,1138]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
A, 8). Male abdomen with subtruncate somite 1, not reaching P5 coxae, separated by part of thoracic sternite 8; somite 2 subequal to width of somite 1; somites 2–5 immobile, sutures between somites incomplete to shallow; somite 5 broadly hexagonal with gently convex lateral margins; somite 6 immobile or with limited mobility in somite 5 although suture still present; telson free (
<figureCitation id="13672A02D3300C02CA2D9CB8FC09FB32" box="[885,968,1228,1254]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
B, 9B, 10B, C). G1 gradually recurved, no distinct junction between recurved, proximal portions, tip broad, truncate, with strong, robust, subterminal spines on one edge (
<figureCitation id="13672A02D3300C02C8DD9D6DFE16FAE7" box="[389,471,1305,1331]" captionStart="FIGURE 11" captionStartId="23.[151,257,1868,1892]" captionTargetBox="[156,1409,201,1868]" captionTargetId="figure@23.[156,1430,171,1886]" captionTargetPageId="23" captionText="FIGURE 11. Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000.9), China; Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait. A, G, carapaces; B, H, frontal views of carapaces showing orbits, eyes and epistomes; C, outer view of right third maxilliped; D, outer view of right chela; J, outer view of left chela; I, outer view of left third maxilliped; E, K, left G 1; F, L, distal part of left G 1. Scales: A, B, G – J = 2.0 mm; C, D, E, K = 1.0 mm; F, L = 0.5 mm." httpUri="https://zenodo.org/record/191919/files/figure.png" pageId="16" pageNumber="17">Fig. 11</figureCitation>
K, L).
</paragraph>
</subSubSection>
<subSubSection id="C346650CD3300C06C99E9D4BFB8CFF0C" lastPageId="20" lastPageNumber="21" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BE33687D3300C02C99E9D4BFAE1F95B" blockId="16.[151,1437,222,2027]" pageId="16" pageNumber="17">
<emphasis id="B928EA95D3300C02C99E9D4BFE81FA8D" bold="true" box="[198,320,1343,1369]" pageId="16" pageNumber="17">Remarks.</emphasis>
<bibRefCitation id="EFCD4B76D3300C02C8129D34FDADFA8E" author="Jones" box="[330,620,1344,1370]" pageId="16" pageNumber="17" refString="Jones, D. A. &amp; Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983">Jones &amp; Clayton (1983)</bibRefCitation>
originally placed this species in 
<taxonomicName id="4C5C4D04D3300C02CAA69D34FB56FA8D" box="[1022,1175,1344,1369]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3300C02CAA69D34FB56FA8D" box="[1022,1175,1344,1369]" italics="true" pageId="16" pageNumber="17">Cleistostoma</emphasis>
</taxonomicName>
on the basis of these characters: the convex lateral margins of the carapace which are broader than long; operculiform third maxillipeds with the merus as long as, or longer than ischium; stout chelae in males; absence of tufts of setae between legs; reflexed G1; and male abdomen with six somites and telson. None of these characters are useful at the generic level, and a re-examination of the material on hand indicates that not all the male abdominal somites are free. The description and figure by 
<bibRefCitation id="EFCD4B76D3300C02CB9D9E75FBB0F9CF" author="Jones" box="[709,1137,1537,1563]" pageId="16" pageNumber="17" refString="Jones, D. A. &amp; Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983" yearSuffix="i">
Jones &amp; Clayton (1983: 187, fig. 
<date id="FFE21047D3300C02CD089E75FBA6F9CF" box="[1104,1127,1537,1563]" pageId="16" pageNumber="17">2i</date>
)
</bibRefCitation>
of the male abdomen are incorrect. The male abdominal somites 2–5 are functionally fused even if parts of the suture are visible (
<figureCitation id="13672A02D3300C02CC399E5CFF67F9BC" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Figs. 7</figureCitation>
C, 9B, 10B). The median sutures are visible but the lateral parts are very shallow; and even between somites 4 and 5, where the suture is relatively more prominent, somites 2–5 cannot be flexed (
<figureCitation id="13672A02D3300C02CDDD9E01FB0AF95B" box="[1157,1227,1653,1679]" captionStart="FIGURE 7" captionStartId="17.[151,256,1433,1457]" captionTargetBox="[151,1435,351,1408]" captionTargetId="figure@17.[151,1436,351,1408]" captionTargetPageId="17" captionText="FIGURE 7. Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen." httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="16" pageNumber="17">Fig. 7</figureCitation>
C, 9B).
</paragraph>
<paragraph id="8BE33687D3300C03C99E9EE8FF28FEF2" blockId="16.[151,1437,222,2027]" lastBlockId="17.[151,1437,152,294]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<bibRefCitation id="EFCD4B76D3300C02C99E9EE8FE3CF962" author="Al-Khayat" box="[198,509,1692,1718]" pageId="16" pageNumber="17" refString="Al-Khayat, J. A. &amp; Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat &amp; Jones (1996)</bibRefCitation>
subsequently argued that 
<taxonomicName id="4C5C4D04D3300C02CA759EE8FC13F961" box="[813,978,1692,1717]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="kuwaitense">
<emphasis id="B928EA95D3300C02CA759EE8FC13F961" box="[813,978,1692,1717]" italics="true" pageId="16" pageNumber="17">C. kuwaitense</emphasis>
</taxonomicName>
should be referred to a new monotypic genus, 
<taxonomicName id="4C5C4D04D3300C02C9B59EB7FE5AF908" box="[237,411,1731,1756]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3300C02C9B59EB7FE5AF908" box="[237,411,1731,1756]" italics="true" pageId="16" pageNumber="17">Leptochryseus</emphasis>
</taxonomicName>
, with the following comments: “Originally diagnosed as 
<taxonomicName id="4C5C4D04D3300C02CD019EB7FB36F908" box="[1113,1271,1731,1756]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3300C02CD019EB7FB36F908" box="[1113,1271,1731,1756]" italics="true" pageId="16" pageNumber="17">Cleistostoma</emphasis>
</taxonomicName>
by 
<bibRefCitation id="EFCD4B76D3300C02CC7D9EB7FE9CF8D7" author="Jones" pageId="16" pageNumber="17" refString="Jones, D. A. &amp; Clayton, D. (1983) The systematics and ecology of crabs belonging to the genera Cleistostoma de Haan and Paracleistostoma de Man on Kuwait mudflats. Crustaceana, 45 (2), 183 - 199." type="journal article" year="1983">Jones and Clayton (1983)</bibRefCitation>
, but now excluded, since all abdominal segments are free ….. 
<taxonomicName id="4C5C4D04D3300C02CDD19E9DFA80F8D6" box="[1161,1345,1769,1794]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3300C02CDD19E9DFA80F8D6" box="[1161,1345,1769,1794]" italics="true" pageId="16" pageNumber="17">Leptochryseus</emphasis>
</taxonomicName>
is also distinguished from 
<taxonomicName id="4C5C4D04D3300C02C8249F64FE38F8FD" box="[380,505,1808,1833]" class="Malacostraca" family="Camptandriidae" genus="Manningis" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3300C02C8249F64FE38F8FD" box="[380,505,1808,1833]" italics="true" pageId="16" pageNumber="17">Manningis</emphasis>
</taxonomicName>
and other related genera, since the first abdominal somite reaches the coxae of the fifth pereopods, but is not wider than remaining abdominal somites” (
<bibRefCitation id="EFCD4B76D3300C02CAA89F42FA51F884" author="Al-Khayat" box="[1008,1424,1846,1872]" pageId="16" pageNumber="17" refString="Al-Khayat, J. A. &amp; Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat &amp; Jones 1996: 798, 799</bibRefCitation>
). As discussed earlier, the male abdominal somites 2–5 are in fact fused. With regards to male abdominal somite 1 reaching the coxae of the P5, this is also incorrect. The sides of the male abdomen are lined with setae, which obscure the surfaces of the sternites. Male abdominal somite 1 is not much wider than the next few somites but there remains a clear gap between the edge of this somite and the P5 coxae (
<figureCitation id="13672A02D3300C02CD839FA5FAF0F83F" box="[1243,1329,2001,2027]" captionStart="FIGURE 10" captionStartId="21.[151,259,1814,1838]" captionText="FIGURE 10. Posterior part of male carapace, thoracic sternum and anterior abdominal somites. A, Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000. 9), China; B, Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait; C, Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (10.7 x 15.1 mm) (ZRC 2009.834), Iraq. Abbreviations: a 1 – 3 = male abdominal somites 1 – 3, respectively; cx 5 = coxa of P 5; s 7, 8 = thoracic sternites 7 and 8, respectively." pageId="16" pageNumber="17">Fig. 10</figureCitation>
B). Ng 
<emphasis id="B928EA95D3300C03CCDF9FA5FF76FF65" italics="true" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">et al.</emphasis>
(2008: 234) commented that 
<taxonomicName id="4C5C4D04D3310C03CB7E98ECFBDFFF66" authority="Al-Khayat &amp; Jones, 1996" authorityName="Al-Khayat &amp; Jones" authorityYear="1996" box="[550,1054,152,178]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3310C03CB7E98ECFD16FF65" box="[550,727,152,177]" italics="true" pageId="17" pageNumber="18">Leptochryseus</emphasis>
<bibRefCitation id="EFCD4B76D3310C03CBB898ECFBDFFF66" author="Al-Khayat" box="[736,1054,152,178]" pageId="17" pageNumber="18" refString="Al-Khayat, J. A. &amp; Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat &amp; Jones, 1996</bibRefCitation>
</taxonomicName>
, was only a junior synonym of 
<taxonomicName id="4C5C4D04D3310C03C9CF98CBFE30FF0C" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" box="[151,497,190,216]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3310C03C9CF98CBFEF3FF0C" box="[151,306,191,216]" italics="true" pageId="17" pageNumber="18">Cleistostoma</emphasis>
De Haan, 1833
</taxonomicName>
, and retained the name 
<taxonomicName id="4C5C4D04D3310C03CA5598CBFBF5FF0C" box="[781,1076,191,216]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kuwaitense">
<emphasis id="B928EA95D3310C03CA5598CBFBF5FF0C" box="[781,1076,191,216]" italics="true" pageId="17" pageNumber="18">Cleistostoma kuwaitense</emphasis>
</taxonomicName>
. They based their decision on the diagnostic characters provided by 
<bibRefCitation id="EFCD4B76D3310C03CB3D9891FC6BFF2B" author="Al-Khayat" box="[613,938,229,255]" pageId="17" pageNumber="18" refString="Al-Khayat, J. A. &amp; Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat &amp; Jones (1996)</bibRefCitation>
, which, as has been discussed above, are wrong.
</paragraph>
<caption id="DF23660FD3310C03C9CF9DEDFEC1FA2E" httpUri="https://zenodo.org/record/191916/files/figure.png" pageId="17" pageNumber="18" targetBox="[151,1435,351,1408]" targetPageId="17">
<paragraph id="8BE33687D3310C03C9CF9DEDFEC1FA2E" blockId="17.[151,1437,1433,1530]" pageId="17" pageNumber="18">
<emphasis id="B928EA95D3310C03C9CF9DEDFEDAFA65" bold="true" box="[151,283,1433,1457]" pageId="17" pageNumber="18">FIGURE 7.</emphasis>
<taxonomicName id="4C5C4D04D3310C03C87C9DEEFCA1FA65" authority="Jones &amp; Clayton, 1983" authorityName="Jones &amp; Clayton" authorityYear="1983" box="[292,864,1434,1457]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kuwaitensis">
<emphasis id="B928EA95D3310C03C87C9DEEFD89FA65" box="[292,584,1434,1457]" italics="true" pageId="17" pageNumber="18">Leptochryseus kuwaitensis</emphasis>
(Jones &amp; Clayton, 1983)
</taxonomicName>
, holotype male (18.0 x 24.1 mm) (NHM 1981.498), Kuwait. A, overall dorsal view; B, frontal view of carapace; C, ventral view showing anterior thoracic sternum and abdomen.
</paragraph>
</caption>
<paragraph id="8BE33687D3310C03C99E9E22FB86F826" blockId="17.[151,1438,1622,2034]" pageId="17" pageNumber="18">
Comparisons of 
<taxonomicName id="4C5C4D04D3310C03C8CA9E22FD30F9A4" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" box="[402,753,1622,1648]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dilatatum">
<emphasis id="B928EA95D3310C03C8CA9E22FDE8F9BB" box="[402,553,1622,1647]" italics="true" pageId="17" pageNumber="18">C. dilatatum</emphasis>
De Haan, 1833
</taxonomicName>
(
<typeStatus id="54E78825D3310C03CA5D9E22FCF8F9A4" box="[773,825,1622,1648]" pageId="17" pageNumber="18">type</typeStatus>
species of 
<taxonomicName id="4C5C4D04D3310C03CA939E22FAEDF9A4" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" box="[971,1324,1622,1648]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3310C03CA939E22FBA5F9BB" box="[971,1124,1622,1647]" italics="true" pageId="17" pageNumber="18">Cleistostoma</emphasis>
De Haan, 1833
</taxonomicName>
) with 
<taxonomicName id="4C5C4D04D3310C03CCD99E22FEE3F942" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kuwaitense">
<emphasis id="B928EA95D3310C03CCD99E22FEE3F942" italics="true" pageId="17" pageNumber="18">C. kuwaitense</emphasis>
</taxonomicName>
, however, now indicate that 
<taxonomicName id="4C5C4D04D3310C03CBCA9E09FC86F942" box="[658,839,1661,1686]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3310C03CBCA9E09FC86F942" box="[658,839,1661,1686]" italics="true" pageId="17" pageNumber="18">Leptochryseus</emphasis>
</taxonomicName>
can be recognised by using a different suite of characters not previously reported. In 
<taxonomicName id="4C5C4D04D3310C03CB0B9ED7FD25F968" box="[595,740,1699,1724]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dilatatum">
<emphasis id="B928EA95D3310C03CB0B9ED7FD25F968" box="[595,740,1699,1724]" italics="true" pageId="17" pageNumber="18">C. dilatatum</emphasis>
</taxonomicName>
, the lateral edges of male abdominal somite 1 taper to form a distinctly triangular structure and reach beyond the edge of the P5 coxae, partially covering part of it (
<figureCitation id="13672A02D3310C03CC349EBEFF74F8DE" captionStart="FIGURE 10" captionStartId="21.[151,259,1814,1838]" captionText="FIGURE 10. Posterior part of male carapace, thoracic sternum and anterior abdominal somites. A, Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000. 9), China; B, Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait; C, Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (10.7 x 15.1 mm) (ZRC 2009.834), Iraq. Abbreviations: a 1 – 3 = male abdominal somites 1 – 3, respectively; cx 5 = coxa of P 5; s 7, 8 = thoracic sternites 7 and 8, respectively." pageId="17" pageNumber="18">Fig. 10</figureCitation>
A). In 
<taxonomicName id="4C5C4D04D3310C03C8599E85FE68F8DE" box="[257,425,1777,1802]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kuwaitensis">
<emphasis id="B928EA95D3310C03C8599E85FE68F8DE" box="[257,425,1777,1802]" italics="true" pageId="17" pageNumber="18">L. kuwaitensis</emphasis>
</taxonomicName>
, the lateral edges of the male abdominal somite 1 are more truncate and less wide, not reaching the P5 coxae. As a result, a part of thoracic sternite 8 remains exposed between the edge of male abdominal somite 1 and the P5 coxae (
<figureCitation id="13672A02D3310C03CB399F4AFD79F88C" box="[609,696,1854,1880]" captionStart="FIGURE 10" captionStartId="21.[151,259,1814,1838]" captionText="FIGURE 10. Posterior part of male carapace, thoracic sternum and anterior abdominal somites. A, Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000. 9), China; B, Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait; C, Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (10.7 x 15.1 mm) (ZRC 2009.834), Iraq. Abbreviations: a 1 – 3 = male abdominal somites 1 – 3, respectively; cx 5 = coxa of P 5; s 7, 8 = thoracic sternites 7 and 8, respectively." pageId="17" pageNumber="18">Fig. 10</figureCitation>
B). The structure of the G1 is also distinct. In 
<taxonomicName id="4C5C4D04D3310C03CD8E9F4AFAA8F883" box="[1238,1385,1854,1879]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="dilatatum">
<emphasis id="B928EA95D3310C03CD8E9F4AFAA8F883" box="[1238,1385,1854,1879]" italics="true" pageId="17" pageNumber="18">C. dilatatum</emphasis>
</taxonomicName>
, the G1 apex is drawn out into a point and has long, slender mushroom-like tubercles positioned subterminally on one edge (
<figureCitation id="13672A02D3310C03C8409FFFFEB1F871" box="[280,368,1931,1957]" captionStart="FIGURE 11" captionStartId="23.[151,257,1868,1892]" captionTargetBox="[156,1409,201,1868]" captionTargetId="figure@23.[156,1430,171,1886]" captionTargetPageId="23" captionText="FIGURE 11. Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000.9), China; Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait. A, G, carapaces; B, H, frontal views of carapaces showing orbits, eyes and epistomes; C, outer view of right third maxilliped; D, outer view of right chela; J, outer view of left chela; I, outer view of left third maxilliped; E, K, left G 1; F, L, distal part of left G 1. Scales: A, B, G – J = 2.0 mm; C, D, E, K = 1.0 mm; F, L = 0.5 mm." httpUri="https://zenodo.org/record/191919/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
E, F). The G1 apex is broad and truncate in 
<taxonomicName id="4C5C4D04D3310C03CACE9FFFFB85F870" box="[918,1092,1931,1956]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="kuwaitense">
<emphasis id="B928EA95D3310C03CACE9FFFFB85F870" box="[918,1092,1931,1956]" italics="true" pageId="17" pageNumber="18">C. kuwaitense</emphasis>
</taxonomicName>
, with strong, robust, spines situated subterminally on one edge (
<figureCitation id="13672A02D3310C03CB009FC6FD6EF818" box="[600,687,1970,1996]" captionStart="FIGURE 11" captionStartId="23.[151,257,1868,1892]" captionTargetBox="[156,1409,201,1868]" captionTargetId="figure@23.[156,1430,171,1886]" captionTargetPageId="23" captionText="FIGURE 11. Cleistostoma dilatatum (De Haan, 1833), male (19.5 x 28.3 mm) (ZRC 2000.9), China; Leptochryseus kuwaitensis (Jones &amp; Clayton, 1983), male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait. A, G, carapaces; B, H, frontal views of carapaces showing orbits, eyes and epistomes; C, outer view of right third maxilliped; D, outer view of right chela; J, outer view of left chela; I, outer view of left third maxilliped; E, K, left G 1; F, L, distal part of left G 1. Scales: A, B, G – J = 2.0 mm; C, D, E, K = 1.0 mm; F, L = 0.5 mm." httpUri="https://zenodo.org/record/191919/files/figure.png" pageId="17" pageNumber="18">Fig. 11</figureCitation>
K, L). These characters argue for recognizing 
<taxonomicName id="4C5C4D04D3310C03CDB49FC6FE04F826" authority="Al-Khayat &amp; Jones, 1996" authorityName="Al-Khayat &amp; Jones" authorityYear="1996" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3310C03CDB49FC6FA5DF81F" box="[1260,1436,1970,1995]" italics="true" pageId="17" pageNumber="18">Leptochryseus</emphasis>
<bibRefCitation id="EFCD4B76D3310C03C9CF9FACFE04F826" author="Al-Khayat" box="[151,453,2008,2034]" pageId="17" pageNumber="18" refString="Al-Khayat, J. A. &amp; Jones, D. A. (1996) Two new genera, Manningis and Leptochryseus (Decapoda: Camptandriinae), and descriptions of the first zoea of six brachyurans from the Arabian Gulf. Journal of Crustacean Biology, 16 (4), 797 - 813." type="journal article" year="1996">Al-Khayat &amp; Jones, 1996</bibRefCitation>
</taxonomicName>
, as a genus distinct from 
<taxonomicName id="4C5C4D04D3310C03CBB69FADFB83F826" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" box="[750,1090,2008,2034]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3310C03CBB69FADFC46F826" box="[750,903,2009,2034]" italics="true" pageId="17" pageNumber="18">Cleistostoma</emphasis>
De Haan, 1833
</taxonomicName>
.
</paragraph>
<caption id="DF23660FD3320C00C9CF9F29FF2CF86D" httpUri="https://zenodo.org/record/191917/files/figure.png" pageId="18" pageNumber="19" targetBox="[151,1436,197,1858]" targetPageId="18">
<paragraph id="8BE33687D3320C00C9CF9F29FF2CF86D" blockId="18.[151,1437,1885,1977]" pageId="18" pageNumber="19">
<emphasis id="B928EA95D3320C00C9CF9F29FEE1F8A1" bold="true" box="[151,288,1885,1909]" pageId="18" pageNumber="19">FIGURE 8.</emphasis>
<taxonomicName id="4C5C4D04D3320C00C8709F29FCB9F8A0" authority="Jones &amp; Clayton, 1983" authorityName="Jones &amp; Clayton" authorityYear="1983" box="[296,888,1885,1908]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kuwaitensis">
<emphasis id="B928EA95D3320C00C8709F29FD97F8A0" box="[296,598,1885,1908]" italics="true" pageId="18" pageNumber="19">Leptochryseus kuwaitensis</emphasis>
(Jones &amp; Clayton, 1983)
</taxonomicName>
, overall views. A, male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait; B, male (20.5 x 15.5 mm) (NHM 1979.269); C, female (16.8 x 22.1 mm) (ZRC 1999.1208), Kuwait.
</paragraph>
</caption>
<caption id="DF23660FD3330C01C9CF9FEDFEB5F821" httpUri="https://zenodo.org/record/191918/files/figure.png" pageId="19" pageNumber="20" targetBox="[151,1436,186,1927]" targetPageId="19">
<paragraph id="8BE33687D3330C01C9CF9FEDFEB5F821" blockId="19.[151,1436,1945,2037]" pageId="19" pageNumber="20">
<emphasis id="B928EA95D3330C01C9CF9FEDFEDAF865" bold="true" box="[151,283,1945,1969]" pageId="19" pageNumber="20">FIGURE 9.</emphasis>
Male thoracic sternum and abdomen. A, 
<taxonomicName id="4C5C4D04D3330C01CB829FEDFB53F864" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" box="[730,1170,1945,1968]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="dilatatum">
<emphasis id="B928EA95D3330C01CB829FEDFC15F864" box="[730,980,1945,1968]" italics="true" pageId="19" pageNumber="20">Cleistostoma dilatatum</emphasis>
(De Haan, 1833)
</taxonomicName>
, male, (14.7 x 20.4 mm) (ZRC 1965.71.15.59), China; B, 
<taxonomicName id="4C5C4D04D3330C01CB5E9FC8FB8BF807" authority="Jones &amp; Clayton, 1983" authorityName="Jones &amp; Clayton" authorityYear="1983" box="[518,1098,1980,2003]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="kuwaitensis">
<emphasis id="B928EA95D3330C01CB5E9FC8FCEFF807" box="[518,814,1980,2003]" italics="true" pageId="19" pageNumber="20">Leptochryseus kuwaitensis</emphasis>
(Jones &amp; Clayton, 1983)
</taxonomicName>
, male (22.7 x 28.6 mm) (ZRC 1999.1208), Kuwait.
</paragraph>
</caption>
<paragraph id="8BE33687D3340C06C99E98ECFB8CFF0C" blockId="20.[151,1437,152,410]" pageId="20" pageNumber="21">
The close similarity between 
<taxonomicName id="4C5C4D04D3340C06CB6398ECFC9EFF66" authority="Manning, 1991" authorityName="Manning" authorityYear="1991" box="[571,863,152,178]" class="Malacostraca" family="Camptandriidae" genus="Nasima" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3340C06CB6398ECFD54FF65" box="[571,661,152,177]" italics="true" pageId="20" pageNumber="21">Nasima</emphasis>
<bibRefCitation id="EFCD4B76D3340C06CBFB98ECFC9EFF66" author="Manning" box="[675,863,152,178]" pageId="20" pageNumber="21" refString="Manning, R. B. (1991) The status of Cleistostoma dotilliforme Alcock, 1900 (Crustacea: Decapoda: Ocypodidae), with the description of a new genus. Raffles Bulletin of Zoology, 39 (2), 299 - 306." type="journal article" year="1991">Manning, 1991</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C5C4D04D3340C06CAF398ECFB95FF65" box="[939,1108,152,177]" class="Malacostraca" family="Camptandriidae" genus="Leptochryseus" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B928EA95D3340C06CAF398ECFB95FF65" box="[939,1108,152,177]" italics="true" pageId="20" pageNumber="21">Leptochryseus</emphasis>
</taxonomicName>
, as well as their possible synonymy, has been discussed at length under the remarks of the previous genus.
</paragraph>
</subSubSection>
<subSubSection id="C346650CD3340C06C99E9891FA98FE4E" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BE33687D3340C06C99E9891FA98FE4E" blockId="20.[151,1437,152,410]" pageId="20" pageNumber="21">
<emphasis id="B928EA95D3340C06C99E9891FD92FF2B" bold="true" box="[198,595,229,255]" pageId="20" pageNumber="21">
Comparative material. 
<collectingCountry id="F34B7617D3340C06C8A99891FD92FF2B" box="[497,595,229,255]" name="China" pageId="20" pageNumber="21">CHINA</collectingCountry>
</emphasis>
: 
<taxonomicName id="4C5C4D04D3340C06CB3D9891FB93FF2B" authority="De Haan, 1833" authorityName="De Haan" authorityYear="1833" box="[613,1106,229,255]" class="Malacostraca" family="Camptandriidae" genus="Cleistostoma" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="dilatatum">
<emphasis id="B928EA95D3340C06CB3D9891FCB9FF2A" box="[613,888,229,254]" italics="true" pageId="20" pageNumber="21">Cleistostoma dilatatum</emphasis>
(De Haan, 1833)
</taxonomicName>
: 
<specimenCount id="9D5AFD0ED3340C06CD389891FB7CFF2B" box="[1120,1213,229,255]" pageId="20" pageNumber="21" type="male">2 males</specimenCount>
(14.7 x 
<quantity id="4CA49B62D3340C06CC719891FA59FF2B" box="[1321,1432,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" pageId="20" pageNumber="21" unit="mm" value="20.4">20.4 mm</quantity>
, 16.3 x 
<quantity id="4CA49B62D3340C06C9B79978FEA5FEF2" box="[239,356,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="20" pageNumber="21" unit="mm" value="22.9">22.9 mm</quantity>
), 
<specimenCount id="9D5AFD0ED3340C06C8229978FE35FEF2" box="[378,500,268,294]" pageId="20" pageNumber="21" type="female">2 females</specimenCount>
(13.6 x 
<quantity id="4CA49B62D3340C06CB389978FD13FEF2" box="[608,722,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="1.96" pageId="20" pageNumber="21" unit="mm" value="19.6">19.6 mm</quantity>
, 14.2 x 
<quantity id="4CA49B62D3340C06CA609978FC6BFEF2" box="[824,938,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="2.01" pageId="20" pageNumber="21" unit="mm" value="20.1">20.1 mm</quantity>
) (
<collectionCode id="ED4DAE42D3340C06CA9B9978FBC0FEF2" box="[963,1025,268,294]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="21">ZRC</collectionCode>
1965.71.15.59–62), Yangmatao, Shantung Peninsula, no date; 
<specimenCount id="9D5AFD0ED3340C06CB599946FD95FE98" box="[513,596,306,332]" pageId="20" pageNumber="21" type="male">1 male</specimenCount>
(19.5 x 
<quantity id="4CA49B62D3340C06CBE39946FCEDFE98" box="[699,812,306,332]" metricMagnitude="-2" metricUnit="m" metricValue="2.83" pageId="20" pageNumber="21" unit="mm" value="28.3">28.3 mm</quantity>
), 
<specimenCount id="9D5AFD0ED3340C06CA1A9946FC6AFE98" box="[834,939,306,332]" pageId="20" pageNumber="21" type="female">1 female</specimenCount>
(19.2 x 
<quantity id="4CA49B62D3340C06CD4B9946FB43FE98" box="[1043,1154,306,332]" metricMagnitude="-2" metricUnit="m" metricValue="3.06" pageId="20" pageNumber="21" unit="mm" value="30.6">30.6 mm</quantity>
) (
<collectionCode id="ED4DAE42D3340C06CDC29946FB17FE98" box="[1178,1238,306,332]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="20" pageNumber="21">ZRC</collectionCode>
2000.9), Hebei, Tangu, coll. 
<date id="FFE21047D3340C06C871992DFE02FEA7" box="[297,451,345,371]" pageId="20" pageNumber="21" value="1913-07-23">23 July 1913</date>
. 
<emphasis id="B928EA95D3340C06C897992CFD69FEA6" bold="true" box="[463,680,344,370]" pageId="20" pageNumber="21">
<collectingCountry id="F34B7617D3340C06C897992CFD69FEA6" box="[463,680,344,370]" name="South Korea" pageId="20" pageNumber="21">SOUTH KOREA</collectingCountry>
</emphasis>
: 
<specimenCount id="9D5AFD0ED3340C06CBE0992DFCE9FEA7" box="[696,808,345,371]" pageId="20" pageNumber="21" type="female">8 females</specimenCount>
(10.0– 
<quantity id="4CA49B62D3340C06CA23992DFC25FEA7" box="[891,996,345,371]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="20" pageNumber="21" unit="mm" value="15.4">15.4 mm</quantity>
x 
<quantity id="4CA49B62D3340C06CD59992DFB6CFEA7" box="[1025,1197,345,371]" metricMagnitude="-2" metricUnit="m" metricValue="1.855" metricValueMax="2.26" metricValueMin="1.45" pageId="20" pageNumber="21" unit="mm" value="18.55" valueMax="22.6" valueMin="14.5">14.5–22.6 mm</quantity>
) (
<collectionCode id="ED4DAE42D3340C06CD9C992DFADFFEA7" LSID="urn:lsid:biocol.org:col:34992" box="[1220,1310,345,371]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="20" pageNumber="21">RMNH</collectionCode>
D 39236), in mud, Asan Bay, Yellow Sea, on west coast, southwest of Seoul, coll. C. Swennen, 
<date id="FFE21047D3340C06CD2499F4FA95FE4E" box="[1148,1364,384,410]" pageId="20" pageNumber="21" value="1989-09-09">9 September 1989</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>