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<document id="D7969BA26DF74B640CC93AA4A2306F1A" ID-DOI="10.11646/zootaxa.3847.3.1" ID-GBIF-Dataset="624e0a0e-4766-436b-a552-8aad3b411999" ID-ISSN="1175-5326" ID-Zenodo-Dep="253194" ID-ZooBank="19E58554-5B31-496B-B851-A26DEE4B7929" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460471539568" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N." docDate="2014" docId="03CA2572FFD4FF87858017C4C69EB0EB" docLanguage="en" docName="zt03847p332.pdf" docOrigin="Zootaxa 3847 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Morphna plana" docType="treatment" docVersion="13" lastPageNumber="311" masterDocId="FFF35D0AFFDEFF8D85171669C30FB71E" masterDocTitle="On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa" masterLastPageNumber="332" masterPageNumber="301" pageNumber="311" updateTime="1698305926497" updateUser="plazi">
<mods:mods id="E5A34F25C8FBAD9039E79794D2AB339B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1C6E71B0DC426B86BE60D72480E4C339">
<mods:title id="C0C80307D4458BACFDFB219268E2A2A1">On cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from South India and Sri Lanka, with descriptions of new taxa</mods:title>
</mods:titleInfo>
<mods:name id="6539465109E31A04E5448F5B6DD93D79" type="personal">
<mods:role id="606205E227C878F65A0D69D6FEC26920">
<mods:roleTerm id="2E41A003CCDA195AB07C2BBA3BC55AEB">Author</mods:roleTerm>
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<mods:namePart id="A713B78A82F98DDEA32DB773806EE62F">Anisyutkin, Leonid N.</mods:namePart>
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<mods:typeOfResource id="B7FF2B55A4D79033E2204574ADFA5C92">text</mods:typeOfResource>
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<mods:title id="545811480280B4993B3FD84B8E188C51">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E4DBFAFC73DE604EB4931223EECE327D">
<mods:date id="EFD0AF25F704BFA944A74B65F565CB1C">2014</mods:date>
<mods:detail id="08F7958073D2F05E61FFA27C4533A7E2" type="pubDate">
<mods:number id="C6498D702B9CAD1776A1E59A892DBE6B">2014-08-08</mods:number>
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<mods:detail id="607BB91BED5D6476F17D1E3F0B73932A" type="volume">
<mods:number id="626A664DC3213884E467F7BDC0D83677">3847</mods:number>
</mods:detail>
<mods:detail id="781082ACEBF68B786EE07575641E24B9" type="issue">
<mods:number id="26E4691F70A5B8F9432A91C12C0838F0">3</mods:number>
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<mods:classification id="538A83744B2C1BA76AD391395263ECEE">journal article</mods:classification>
<mods:identifier id="CEA84AB8249E29143F3D96103F551B5E" type="DOI">10.11646/zootaxa.3847.3.1</mods:identifier>
<mods:identifier id="D169D1333BB01A868DCB5A502A05CAEC" type="GBIF-Dataset">624e0a0e-4766-436b-a552-8aad3b411999</mods:identifier>
<mods:identifier id="D863905A53270121113914609400B763" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CA2572FFD4FF87858017C4C69EB0EB" ID-DOI="http://doi.org/10.5281/zenodo.6134126" ID-GBIF-Taxon="119515391" ID-Zenodo-Dep="6134126" LSID="urn:lsid:plazi:treatment:03CA2572FFD4FF87858017C4C69EB0EB" httpUri="http://treatment.plazi.org/id/03CA2572FFD4FF87858017C4C69EB0EB" lastPageNumber="311" pageId="10" pageNumber="311">
<subSubSection id="C379C7EFFFD4FF87858017C4C1E8B6D9" box="[151,743,429,455]" pageId="10" pageNumber="311" type="nomenclature">
<paragraph id="8BDC9464FFD4FF87858017C4C1E8B6D9" blockId="10.[151,743,429,488]" box="[151,743,429,455]" pageId="10" pageNumber="311">
<heading id="D0942308FFD4FF87858017C4C1E8B6D9" bold="true" box="[151,743,429,455]" fontSize="11" level="1" pageId="10" pageNumber="311" reason="1">
<taxonomicName id="4C63EFE7FFD4FF87858017C4C1E8B6D9" ID-CoL="742SL" authority="(Brunner von Wattenwyl, 1865)" baseAuthorityName="Brunner von Wattenwyl" baseAuthorityYear="1865" box="[151,743,429,455]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="B9174876FFD4FF87858017C4C1E8B6D9" bold="true" box="[151,743,429,455]" pageId="10" pageNumber="311">
<emphasis id="B9174876FFD4FF87858017C4C25AB6D9" bold="true" box="[151,341,429,455]" italics="true" pageId="10" pageNumber="311">Morphna plana</emphasis>
(
<bibRefCitation id="EFF2E995FFD4FF87847317C4C1D0B6D9" author="Brunner von Wattenwyl, C." bookContentInfo="426 pp." box="[356,735,429,455]" firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Charles Ueberreuter, Wien" pageId="10" pageNumber="311" refId="ref19622" refString="Brunner von Wattenwyl, C. (1865) Nouveau systeme des Blattaires. Charles Ueberreuter, Wien, 426 pp." title="Nouveau systeme des Blattaires" type="book" year="1865">Brunner von Wattenwyl, 1865</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD4FF87858017A6C27BB6F6" box="[151,372,463,488]" pageId="10" pageNumber="311" type="description">
<paragraph id="8BDC9464FFD4FF87858017A6C27BB6F6" blockId="10.[151,743,429,488]" box="[151,372,463,488]" pageId="10" pageNumber="311">
(
<figureCitation id="135888E1FFD4FF87858817A6C3E7B6F6" box="[159,232,463,488]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A – C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H – J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="10" pageNumber="311">Figs. 1</figureCitation>
F, G,
<figureCitation id="135888E1FFD4FF87843117A6C23CB6F6" box="[294,307,463,488]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R1T and R4, perpendicular to sight; J—sclerite L2D from above; K—apical part of sclerite L2D from above; L—sclerite L3; M–O—apical part of sclerite L3. Dotted area shows membranous parts. Abbreviations: c.p.R1T; d.o.; R1T; R2; R3; R4; R5—see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M–O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">5</figureCitation>
A–O)
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD4FF878580147EC7F2B52E" box="[151,1277,535,560]" pageId="10" pageNumber="311" type="materials_examined">
<paragraph id="8BDC9464FFD4FF878580147EC7F2B52E" blockId="10.[151,1437,535,2037]" box="[151,1277,535,560]" pageId="10" pageNumber="311">
<emphasis id="B9174876FFD4FF878580147EC3F0B52E" bold="true" box="[151,255,535,560]" pageId="10" pageNumber="311">Material</emphasis>
.
<materialsCitation id="3B0B9E39FFD4FF87841A1471C7F2B52E" ID-GBIF-Occurrence="3923855301" box="[269,1277,535,560]" collectionCode="MHNG" country="Sri Lanka" location="Penadewa Ceylan I. Humbert" pageId="10" pageNumber="311" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F374D4F4FFD4FF87841A1471C2ABB52E" box="[269,420,536,560]" name="Sri Lanka" pageId="10" pageNumber="311">SRI LANKA</collectingCountry>
, &quot;
<location id="8EBCC2BFFFD4FF8784AF147EC005B52E" LSID="urn:lsid:plazi:treatment:03CA2572FFD4FF87858017C4C69EB0EB:8EBCC2BFFFD4FF8784AF147EC005B52E" box="[440,778,535,560]" country="Sri Lanka" name="Penadewa Ceylan I. Humbert" pageId="10" pageNumber="311">Penadewa Ceylan I. Humbert</location>
&quot;, &quot;
<originalDetermination id="A87D2C98FFD4FF878630147EC71BB52E" box="[807,1044,535,560]" pageId="10" pageNumber="311">
<taxonomicName id="4C63EFE7FFD4FF878630147EC0F7B531" authority="Br." authorityName="Br." box="[807,1016,535,560]" class="Insecta" family="Blaberidae" genus="Molytria" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="plana">Molytria plana Br.</taxonomicName>
♂
</originalDetermination>
&quot;—
<specimenCount id="9D655FEDFFD4FF8781281471C789B52E" box="[1087,1158,535,560]" pageId="10" pageNumber="311" type="male">1 male</specimenCount>
(
<collectionCode id="ED720CA1FFD4FF8781821471C7FEB52E" box="[1173,1265,536,560]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="10" pageNumber="311" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C379C7EFFFD4FF8785D01455C69EB0EB" pageId="10" pageNumber="311" type="description">
<paragraph id="8BDC9464FFD4FF8785D01455C029B38B" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="311">
<emphasis id="B9174876FFD4FF8785D01455C25EB54B" bold="true" box="[199,337,572,597]" pageId="10" pageNumber="311">Description</emphasis>
. Male. General color yellowish brown (
<figureCitation id="135888E1FFD4FF8786211455C074B54A" box="[822,891,572,597]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A – C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H – J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="10" pageNumber="311">Fig. 1</figureCitation>
G); facial part of head dark yellowish, partly darkened at vertex, genae and between antennal sockets, labrum lighter (
<figureCitation id="135888E1FFD4FF8786C61436C71EB566" box="[977,1041,607,632]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A – C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H – J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="10" pageNumber="311">Fig. 1</figureCitation>
F); antennae brownish in proximal (about 16–18 basal segments), greyish in distal parts; 1st–4th segments of maxillary palps—brownish, 5th one—greyish; labial palps—yellowish; pronotum and, in lesser degree, tegmina densely speckled with small brown spots, tegmina lighter toward apex; legs,
<taxonomicName id="4C63EFE7FFD4FF8787C214A5C012B5FB" box="[725,797,716,741]" class="Insecta" family="Blaberidae" genus="Thorax" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="332" phylum="Arthropoda" rank="genus">thorax</taxonomicName>
and abdomen from below dirty yellow, partly brown. Surfaces smooth and lustrous, distal parts of antennae (approximately from 16–18th segments) and 5th segment of maxillary palps dull; punctation present only in basal half of tegmina; facial part of head with weak wrinkles (
<figureCitation id="135888E1FFD4FF878073157DC3AEB44E" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A – C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H – J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="10" pageNumber="311">Figs. 1</figureCitation>
F, 5A). Head longer than wide (
<figureCitation id="135888E1FFD4FF87871F155EC15EB44E" box="[520,593,823,848]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A – C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H – J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="10" pageNumber="311">Figs. 1</figureCitation>
F, 5A); ocellar spots large; facial part with weak impressions between eyes and above clypeus; distance between eyes 0.5 times eye length; distance between antennal sockets 1.8 times of the scape length (
<quantity id="4C9B3981FFD4FF87842515E9C282B486" box="[306,397,896,920]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="311" unit="mm" value="1.1">1.1 mm</quantity>
); approximate length ratio of 3rd–5th segments of maxillary palps 1: 1: 1.2. Pronotum as in
<figureCitation id="135888E1FFD4FF87858015CDC3EFB4A2" box="[151,224,932,957]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A – C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H – J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="10" pageNumber="311">Figs. 1</figureCitation>
G, 5B. Tegmina and wings completely developed, surpassing abdominal apex (
<figureCitation id="135888E1FFD4FF87817215CDC7AFB4A3" box="[1125,1184,932,957]" captionStart="FIGURE 1. A, D, F" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[170,1400,200,1869]" captionTargetId="figure@5.[167,1406,193,1876]" captionTargetPageId="5" captionText="FIGURE 1. A, D, F, H—facial part of head; B, E, G, I, J—general view from above; C—anterior part of body from below. A – C— Indoapterolampra rugosiuscula sp. nov. (male, holotype); D, E— Morphna lucida sp. nov. (male, holotype); F, G— M. plana (Brunner von Wattenwyl, 1865) (male); H – J— M. decolyi (Bolivar, 1897) (H, I—male, J—female). In Figs. B, I, J abdominal apex removed. All specimens are pinned." httpUri="https://zenodo.org/record/253195/files/figure.png" pageId="10" pageNumber="311">Fig.1</figureCitation>
G), similar to those of
<taxonomicName id="4C63EFE7FFD4FF87858015A0C20CB4FE" box="[151,259,967,992]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD4FF87858015A0C20CB4FE" box="[151,259,967,992]" italics="true" pageId="10" pageNumber="311">M. lucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD4FF87841D15AEC26AB4FE" box="[266,357,967,992]" pageId="10" pageNumber="311" rank="species">
<emphasis id="B9174876FFD4FF87841D15AEC26AB4FE" bold="true" box="[266,357,967,992]" pageId="10" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
, but rounded apically and
<emphasis id="B9174876FFD4FF87878515A0C1A6B4FE" box="[658,681,969,992]" italics="true" pageId="10" pageNumber="311">M</emphasis>
stem subobsolete in proximal 1/3 of tegmen length (visible as row of punctures). Anterior margin of fore femur (right femur broken off in specimen studied) armed according to the
<typeStatus id="54D82AC6FFD4FF8785801279C3C8B336" box="[151,199,1040,1064]" pageId="10" pageNumber="311">type</typeStatus>
B, with 6 spines, including 2 apical ones. Structure of hind tarsi (
<figureCitation id="135888E1FFD4FF8786BF1266C0E5B336" box="[936,1002,1039,1064]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
C) similar to that of
<taxonomicName id="4C63EFE7FFD4FF8781D81278C634B336" box="[1231,1339,1039,1064]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD4FF8781D81278C634B336" box="[1231,1339,1039,1064]" italics="true" pageId="10" pageNumber="311">M. lucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD4FF8780551266C693B336" box="[1346,1436,1039,1064]" pageId="10" pageNumber="311" rank="species">
<emphasis id="B9174876FFD4FF8780551266C693B336" bold="true" box="[1346,1436,1039,1064]" pageId="10" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
Anal plate (tergite X) triangular in shape, with triangular median incision on caudal margin (
<figureCitation id="135888E1FFD4FF8781FF125DC620B353" box="[1256,1327,1076,1101]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
D). Cerci slightly shorter than in
<taxonomicName id="4C63EFE7FFD4FF8784BA1230C111B36E" box="[429,542,1111,1136]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD4FF8784BA1230C111B36E" box="[429,542,1111,1136]" italics="true" pageId="10" pageNumber="311">M. lucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD4FF87873E123EC188B36E" box="[553,647,1111,1136]" pageId="10" pageNumber="311" rank="species">
<emphasis id="B9174876FFD4FF87873E123EC188B36E" bold="true" box="[553,647,1111,1136]" pageId="10" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
Paraprocts of the blaberid-type, as in
<figureCitation id="135888E1FFD4FF878144123EC796B36E" box="[1107,1177,1111,1136]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
E. Hypandrium nearly symmetrical (
<figureCitation id="135888E1FFD4FF8784261215C27CB38B" box="[305,371,1148,1173]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
F), similar to that of
<taxonomicName id="4C63EFE7FFD4FF87874C1214C1C9B38B" box="[603,710,1148,1173]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD4FF87874C1214C1C9B38B" box="[603,710,1148,1173]" italics="true" pageId="10" pageNumber="311">M. lucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD4FF8787DA1215C029B38B" box="[717,806,1148,1173]" pageId="10" pageNumber="311" rank="species">
<emphasis id="B9174876FFD4FF8787DA1215C029B38B" bold="true" box="[717,806,1148,1173]" pageId="10" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDC9464FFD4FF8785D012F6C06DB273" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="311">
Male genitalia (
<figureCitation id="135888E1FFD4FF87846112F6C2B8B3A6" box="[374,439,1183,1208]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
G–O). Right phallomere (R+N) as in
<figureCitation id="135888E1FFD4FF87864D12F6C095B3A6" box="[858,922,1183,1208]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
G–I, caudal part of sclerite R1T rounded (
<figureCitation id="135888E1FFD4FF87807812F6C3ABB3C3" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">
Fig.
<date id="FFDDB2A4FFD4FF87858012ADC3ABB3C3" box="[151,164,1220,1245]" pageId="10" pageNumber="311">5</date>
</figureCitation>
I–H,
<emphasis id="B9174876FFD4FF8785C812ADC238B3C2" box="[223,311,1220,1244]" italics="true" pageId="10" pageNumber="311">c.p.R1T</emphasis>
), densely covered with bristles; R2 weakly curved (
<figureCitation id="135888E1FFD4FF87869012ADC0C5B3C3" box="[903,970,1220,1245]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
G); R3 deeply forked caudally (
<figureCitation id="135888E1FFD4FF87802412ADC678B3C3" box="[1331,1399,1220,1245]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
H); R4 large (
<figureCitation id="135888E1FFD4FF878410128EC246B21E" box="[263,329,1255,1280]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">
Fig.
<date id="FFDDB2A4FFD4FF87842B128EC246B21E" box="[316,329,1255,1280]" pageId="10" pageNumber="311">5</date>
</figureCitation>
I–H); R5 large, fused with sclerite R3 (
<figureCitation id="135888E1FFD4FF878613128EC04AB21E" box="[772,837,1255,1280]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
G, H). Sclerite L2D (L1) with basal part very weakly widened cranially (
<figureCitation id="135888E1FFD4FF8784661365C2B9B23B" box="[369,438,1292,1317]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
J); apical part rounded, with distinct &quot;dorsal outgrows&quot;, bristles completely absent (
<figureCitation id="135888E1FFD4FF8780781365C3AAB256" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
K). Sclerite L3 (L2d) (
<figureCitation id="135888E1FFD4FF8784881346C2EFB256" box="[415,480,1327,1352]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
L) similar to that of
<taxonomicName id="4C63EFE7FFD4FF8787D71358C024B256" box="[704,811,1327,1352]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD4FF8787D71358C024B256" box="[704,811,1327,1352]" italics="true" pageId="10" pageNumber="311">M. lucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD4FF8786261346C085B256" box="[817,906,1327,1352]" pageId="10" pageNumber="311" rank="species">
<emphasis id="B9174876FFD4FF8786261346C085B256" bold="true" box="[817,906,1327,1352]" pageId="10" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
, but slightly differs in shape of apical part (
<figureCitation id="135888E1FFD4FF8780791346C3ABB273" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
M–O). Sclerite L4U (L3d) similar to that of
<taxonomicName id="4C63EFE7FFD4FF878781133CC00EB273" box="[662,769,1364,1389]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lucida" status="sp. nov.">
<emphasis id="B9174876FFD4FF878781133CC00EB273" box="[662,769,1364,1389]" italics="true" pageId="10" pageNumber="311">M. lucida</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A224F50DFFD4FF87861F133DC06DB273" box="[776,866,1364,1389]" pageId="10" pageNumber="311" rank="species">
<emphasis id="B9174876FFD4FF87861F133DC06DB273" bold="true" box="[776,866,1364,1389]" pageId="10" pageNumber="311">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BDC9464FFD4FF8785D01310C7A3B2AA" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="311">
Measurements (mm). Head length 4.5, head width 4.2, pronotum length 7.0, pronotum width 9.8, tegmen length 27.0, tegmen width (in place where
<emphasis id="B9174876FFD4FF87876113F4C1A7B2AA" box="[630,680,1437,1460]" italics="true" pageId="10" pageNumber="311">CuP</emphasis>
running into posterior margin of tegmen) 9.0.
</paragraph>
<paragraph id="8BDC9464FFD4FF8785D013D6C241B13E" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="311">
<emphasis id="B9174876FFD4FF8785D013D6C21EB2C6" bold="true" box="[199,273,1471,1496]" pageId="10" pageNumber="311">Notes.</emphasis>
<taxonomicName id="4C63EFE7FFD4FF87840E13D6C2C6B2C6" box="[281,457,1471,1496]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="B9174876FFD4FF87840E13D6C2C6B2C6" box="[281,457,1471,1496]" italics="true" pageId="10" pageNumber="311">Morphna plana</emphasis>
</taxonomicName>
was described in the genus
<taxonomicName id="4C63EFE7FFD4FF87861E13D6C741B2C6" authority="Burmeister, 1838" authorityName="Burmeister" authorityYear="1838" box="[777,1102,1471,1496]" class="Insecta" family="Blaberidae" genus="Epilampra" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD4FF87861E13D6C08EB2C6" box="[777,897,1471,1496]" italics="true" pageId="10" pageNumber="311">Epilampra</emphasis>
<bibRefCitation id="EFF2E995FFD4FF87869F13D6C741B2C6" author="Burmeister, H." box="[904,1102,1471,1496]" firstAuthor="Burmeister" journalOrPublisher="Band. 2, Abteilung 2. Berlin" pageId="10" pageNumber="311" pagination="397 - 1050" refId="ref19681" refString="Burmeister, H. (1838) Handbuch der Entomologie. Band. 2, Abteilung 2. Berlin, 397 - 1050." title="Handbuch der Entomologie" type="journal article" year="1838">Burmeister, 1838</bibRefCitation>
</taxonomicName>
. Later, this species was listed in the genus
<taxonomicName id="4C63EFE7FFD4FF878430138DC2B3B2E3" authority="Stal" authorityName="Stal" box="[295,444,1508,1533]" class="Insecta" family="Blaberidae" genus="Molytria" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD4FF878430138DC286B2E3" box="[295,393,1508,1533]" italics="true" pageId="10" pageNumber="311">Molytria</emphasis>
Stål
</taxonomicName>
(
<bibRefCitation id="EFF2E995FFD4FF8784DC138DC17BB2E3" author="Saussure, H. de" box="[459,628,1508,1533]" firstAuthor="Saussure" journalOrPublisher="Revue suisse de Zoologie" pageId="10" pageNumber="311" pagination="299 - 364" part="3" refId="ref20527" refString="Saussure, H. de (1895) Revision de la tribu des Panesthiens et de celle des Epilampriens (Orthopteres de la Famille des Blattides). Revue suisse de Zoologie, 3, 299 - 364." title="Revision de la tribu des Panesthiens et de celle des Epilampriens (Orthopteres de la Famille des Blattides)" type="journal article" year="1895">Saussure 1895</bibRefCitation>
;
<bibRefCitation id="EFF2E995FFD4FF878797138DC00DB2E2" author="Kirby, W. F." bookContentInfo="501 pp." box="[640,770,1508,1533]" firstAuthor="Kirby" journalOrPublisher="British Museum, London" pageId="10" pageNumber="311" refId="ref19953" refString="Kirby, W. F. (1904) A synonymic Catalogue of Orthoptera. Vol. I. Orthoptera Euplexoptera, Cursoria, et Gressoria (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae). British Museum, London, 501 pp." title="A synonymic Catalogue of Orthoptera. Vol. I. Orthoptera Euplexoptera, Cursoria, et Gressoria (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae)" type="book" year="1904">Kirby 1904</bibRefCitation>
) and finally has been transferred by
<bibRefCitation id="EFF2E995FFD4FF8781B5138DC65BB2E2" author="Shelford, R." box="[1186,1364,1508,1533]" editor="Wytsman, P." firstAuthor="Shelford" journalOrPublisher="Bruxelles" pageId="10" pageNumber="311" pagination="1 - 21" part="101" refId="ref20566" refString="Shelford, R. (1910) Orthoptera. Fam. Blattidae. Subfam. Epilamprinae. In: Wytsman, P. (Ed.), Genera Insectorum, 101. Bruxelles, pp. 1 - 21." title="Orthoptera. Fam. Blattidae. Subfam. Epilamprinae" type="journal article" volumeTitle="Genera Insectorum" year="1910">Shelford (1910)</bibRefCitation>
to the genus
<taxonomicName id="4C63EFE7FFD4FF8785C8106EC248B13E" box="[223,327,1543,1568]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD4FF8785C8106EC248B13E" box="[223,327,1543,1568]" italics="true" pageId="10" pageNumber="311">Morphna</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BDC9464FFD4FF8785D01045C3F3B0B3" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="311">
The male described above broadly corresponds to the original description by C.
<bibRefCitation id="EFF2E995FFD4FF8781551044C698B15B" author="Brunner von Wattenwyl, C." bookContentInfo="426 pp." box="[1090,1431,1580,1605]" firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Charles Ueberreuter, Wien" pageId="10" pageNumber="311" refId="ref19622" refString="Brunner von Wattenwyl, C. (1865) Nouveau systeme des Blattaires. Charles Ueberreuter, Wien, 426 pp." title="Nouveau systeme des Blattaires" type="book" year="1865">Brunner von Wattenwyl (1865</bibRefCitation>
, p. 183, 184). Some differences in coloration can be explained by an intraspecific variation and/or partial fading of the old dried specimen. Brunner von Wattenwyl did not list the
<typeStatus id="54D82AC6FFD4FF878696101CC0BEB193" box="[897,945,1653,1677]" pageId="10" pageNumber="311">type</typeStatus>
series in his original description. He only mentioned that it was based on both sexes from &quot;
<collectingCountry id="F374D4F4FFD4FF8787D110FEC014B1AE" box="[710,795,1687,1712]" name="Sri Lanka" pageId="10" pageNumber="311">Ceylan</collectingCountry>
&quot; and also that he had two males &quot;before my eyes&quot; (&quot;que j'ai devant moi&quot;—
<bibRefCitation id="EFF2E995FFD4FF87847F10D5C1CFB1CB" author="Brunner von Wattenwyl, C." bookContentInfo="426 pp." box="[360,704,1724,1749]" firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Charles Ueberreuter, Wien" pageId="10" pageNumber="311" refId="ref19622" refString="Brunner von Wattenwyl, C. (1865) Nouveau systeme des Blattaires. Charles Ueberreuter, Wien, 426 pp." title="Nouveau systeme des Blattaires" type="book" year="1865">Brunner von Wattenwyl 1865</bibRefCitation>
, p. 184). Both males had the unique stylus on the right side of hypandrium (&quot;La plaque sousgénitale du mâle, légèrement échancree, ne porte qu' un style unique du côté droit dans les deux individus, que j'ai devant moi&quot;—
<bibRefCitation id="EFF2E995FFD4FF8787A2116DC704B003" author="Brunner von Wattenwyl, C." bookContentInfo="426 pp." box="[693,1035,1796,1821]" firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Charles Ueberreuter, Wien" pageId="10" pageNumber="311" refId="ref19622" refString="Brunner von Wattenwyl, C. (1865) Nouveau systeme des Blattaires. Charles Ueberreuter, Wien, 426 pp." title="Nouveau systeme des Blattaires" type="book" year="1865">Brunner von Wattenwyl 1865</bibRefCitation>
, p. 184). The latter fact excludes a possibility that the specimen studied in the present work belonged to the original
<typeStatus id="54D82AC6FFD4FF87813E1140C756B05F" box="[1065,1113,1833,1857]" pageId="10" pageNumber="311">type</typeStatus>
series. If the structure of the hypandrium studied by Brunner von Wattenwyl was not an artefact, this would be evidence that these specimens belong to a different species. In order to resolve this problem, a re-examination of the original
<typeStatus id="54D82AC6FFD4FF8780151118C63DB097" box="[1282,1330,1905,1929]" pageId="10" pageNumber="311">type</typeStatus>
series is required.
</paragraph>
<paragraph id="8BDC9464FFD4FF8785D011D1C69EB0EB" blockId="10.[151,1437,535,2037]" pageId="10" pageNumber="311">
The complete absence of bristles on the apical part of the sclerite L2D (
<figureCitation id="135888E1FFD4FF8786E111D1C737B0CF" box="[1014,1080,1976,2001]" captionStart="FIGURE 5" captionStartId="11.[151,250,1819,1841]" captionTargetBox="[176,1412,211,1789]" captionTargetId="figure@11.[166,1420,195,1797]" captionTargetPageId="11" captionText="FIGURE 5. Morphna plana (Brunner von Wattenwyl, 1865), male. A—facial part of head; B—pronotum from above; C—left hind tarsus from outside; D—abdominal apex from above; E—paraprocts from caudal; F—hypandrium from below; G, H—right phallomere from above (G) and below (H); I—sclerites R 1 T and R 4, perpendicular to sight; J—sclerite L 2 D from above; K—apical part of sclerite L 2 D from above; L—sclerite L 3; M – O—apical part of sclerite L 3. Dotted area shows membranous parts. Abbreviations: c. p. R 1 T; d. o.; R 1 T; R 2; R 3; R 4; R 5 — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D, E; e = F; f = G, H; g = I; h = J; i = K; j = L; k = M – O." httpUri="https://zenodo.org/record/253199/files/figure.png" pageId="10" pageNumber="311">Fig. 5</figureCitation>
K) is an unusual features in the genus
<taxonomicName id="4C63EFE7FFD4FF8785C811B5C247B0EB" box="[223,328,2012,2037]" class="Insecta" family="Blaberidae" genus="Morphna" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="311" phylum="Arthropoda" rank="genus">
<emphasis id="B9174876FFD4FF8785C811B5C247B0EB" box="[223,328,2012,2037]" italics="true" pageId="10" pageNumber="311">Morphna</emphasis>
</taxonomicName>
and the tribe Morphnini as well. In my opinion, it could be explained by the secondary reduction.
</paragraph>
</subSubSection>
</treatment>
</document>
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