treatments-xml/data/03/DB/41/03DB41541F64FFEAF1CBCE79D62CF9B0.xml

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<document id="E9390CBF347187E2BCCEBF9E13383800" ID-CLB-Dataset="30756" ID-DOI="10.11646/zootaxa.4379.1.5" ID-GBIF-Dataset="2c7fe9b1-f5a7-4169-bcc0-d544c884e01f" ID-ISSN="1175-5326" ID-Zenodo-Dep="1172356" ID-ZooBank="E860555F-9CED-4A38-AAF8-B7AB5C1A7E71" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1518511506799" checkinUser="plazi" docAuthor="Rasool, Iftekhar, Abdel-Dayem, Mahmoud S., Felix, Ron F. F. L. &amp; Aldhafer, Hathal M." docDate="2018" docId="03DB41541F64FFEAF1CBCE79D62CF9B0" docLanguage="en" docName="zootaxa.4379.1.5.pdf" docOrigin="Zootaxa 4379 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Matabele arabica Mateu 1986" docType="treatment" docVersion="7" lastPageNumber="100" masterDocId="FFE2392C1F69FFE7F15CCF32D75CFF96" masterDocTitle="A review of the Subtribe Lebiina Bonelli (Lebiini, Carabidae, Coleoptera) from Southwest of Saudi Arabia" masterLastPageNumber="102" masterPageNumber="87" pageNumber="100" updateTime="1698521292142" updateUser="ExternalLinkService">
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<mods:title id="826C65789D701C6A6419FB15D4C561DC">A review of the Subtribe Lebiina Bonelli (Lebiini, Carabidae, Coleoptera) from Southwest of Saudi Arabia</mods:title>
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<mods:namePart id="36B02F49BD405D1A1FB9605D1E00681C">Rasool, Iftekhar</mods:namePart>
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<mods:namePart id="1C922758A4BFB7F6AC3AD45A301654A8">Abdel-Dayem, Mahmoud S.</mods:namePart>
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<mods:namePart id="904CD3C41C2ED7588BC376ED9492B3C6">Felix, Ron F. F. L.</mods:namePart>
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<mods:namePart id="9F061C6241222121F6880B7C7CA2D95D">Aldhafer, Hathal M.</mods:namePart>
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<mods:title id="297FC91E623C60E83E12BB735E74BC43">Zootaxa</mods:title>
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<mods:date id="705B9B471E5E2CC62000EA4D014E22A2">2018</mods:date>
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<treatment id="03DB41541F64FFEAF1CBCE79D62CF9B0" ID-DOI="http://doi.org/10.5281/zenodo.5979222" ID-GBIF-Taxon="140583191" ID-Zenodo-Dep="5979222" LSID="urn:lsid:plazi:treatment:03DB41541F64FFEAF1CBCE79D62CF9B0" httpUri="http://treatment.plazi.org/id/03DB41541F64FFEAF1CBCE79D62CF9B0" lastPageNumber="100" pageId="13" pageNumber="100">
<subSubSection id="C368A3C91F64FFEAF1CBCE79D550FEF0" box="[151,524,331,358]" pageId="13" pageNumber="100" type="nomenclature">
<paragraph id="8BCDF0421F64FFEAF1CBCE79D550FEF0" blockId="13.[151,524,331,358]" box="[151,524,331,358]" pageId="13" pageNumber="100">
<heading id="D085472E1F64FFEAF1CBCE79D550FEF0" bold="true" box="[151,524,331,358]" fontSize="11" level="1" pageId="13" pageNumber="100" reason="1">
<taxonomicName id="4C728BC11F64FFEAF1CBCE79D550FEF0" ID-CoL="3YDWC" authority="Mateu, 1986" authorityName="Mateu" authorityYear="1986" box="[151,524,331,358]" class="Insecta" family="Carabidae" genus="Matabele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="B9062C501F64FFEAF1CBCE79D550FEF0" bold="true" box="[151,524,331,358]" pageId="13" pageNumber="100">
<emphasis id="B9062C501F64FFEAF1CBCE79D635FEF3" bold="true" box="[151,361,331,357]" italics="true" pageId="13" pageNumber="100">Matabele arabica</emphasis>
<bibRefCitation id="EFE38DB31F64FFEAF02CCE7ED550FEF0" box="[368,524,332,358]" pageId="13" pageNumber="100" refId="ref11942" refString="Mateu, J. (1986) Lebiinae and Brachininae of Saudi Arabia (Coleoptera: Carabidae). Fauna of Saudi Arabia, 8, 198 - 217." type="journal article">Mateu, 1986</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C368A3C91F64FFEAF1CBCEA0D559FE3E" box="[151,517,402,424]" pageId="13" pageNumber="100" type="reference_group">
<paragraph id="8BCDF0421F64FFEAF1CBCEA0D559FE3E" blockId="13.[151,1438,402,1574]" box="[151,517,402,424]" pageId="13" pageNumber="100">
<treatmentCitationGroup id="AB62D76C1F64FFEAF1CBCEA0D559FE3E" box="[151,517,402,424]" pageId="13" pageNumber="100">
<taxonomicName id="4C728BC11F64FFEAF1CBCEA0D559FE3E" ID-CoL="3YDWC" authority="Mateu, 1986: 201" authorityName="Mateu" authorityPageNumber="201" authorityYear="1986" box="[151,517,402,424]" class="Insecta" family="Carabidae" genus="Matabele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="arabica">
<emphasis id="B9062C501F64FFEAF1CBCEA0D610FE3E" box="[151,332,402,424]" italics="true" pageId="13" pageNumber="100">Matabele arabica</emphasis>
<treatmentCitation id="0AD3D6531F64FFEAF00ECEA1D559FE3E" author="" box="[338,517,402,424]" page="201" pageId="13" pageNumber="100">
<bibRefCitation id="EFE38DB31F64FFEAF00ECEA1D689FE3E" box="[338,469,402,424]" pageId="13" pageNumber="100" refId="ref11942" refString="Mateu, J. (1986) Lebiinae and Brachininae of Saudi Arabia (Coleoptera: Carabidae). Fauna of Saudi Arabia, 8, 198 - 217." type="journal article">Mateu, 1986</bibRefCitation>
: 201
</treatmentCitation>
</taxonomicName>
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</paragraph>
</subSubSection>
<subSubSection id="C368A3C91F64FFEAF1CBCE83D243FB2B" pageId="13" pageNumber="100" type="materials_examined">
<paragraph id="8BCDF0421F64FFEAF1CBCE83D502FE5F" blockId="13.[151,1438,402,1574]" box="[151,606,433,458]" pageId="13" pageNumber="100">
<materialsCitation id="3B1AFA1F1F64FFEAF1CBCE83D502FE5F" ID-GBIF-Occurrence="1806563519" box="[151,606,433,458]" country="Saudi Arabia" location="Saudi Arabia" pageId="13" pageNumber="100" specimenCount="1" typeStatus="holotype">
<emphasis id="B9062C501F64FFEAF1CBCE83D669FE5C" bold="true" box="[151,309,433,458]" pageId="13" pageNumber="100">
<typeStatus id="54C94EE01F64FFEAF1CBCE83D78EFE5C" box="[151,210,433,458]" pageId="13" pageNumber="100">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F365B0D21F64FFEAF060CE83D68EFE5C" box="[316,466,433,458]" name="Saudi Arabia" pageId="13" pageNumber="100">Saudi Arabia</collectingCountry>
, An-Namas.
</materialsCitation>
</paragraph>
<paragraph id="8BCDF0421F64FFEAF19BCEE7D59CFE7B" blockId="13.[151,1438,402,1574]" box="[199,704,469,494]" pageId="13" pageNumber="100">
<emphasis id="B9062C501F64FFEAF19BCEE7D6D0FE78" bold="true" box="[199,396,469,494]" pageId="13" pageNumber="100">Type depository.</emphasis>
Holotype; male in BMNH.
</paragraph>
<paragraph id="8BCDF0421F64FFEAF19BCECBD3C8FDCF" blockId="13.[151,1438,402,1574]" pageId="13" pageNumber="100">
<materialsCitation id="3B1AFA1F1F64FFEAF19BCECBD612FDA0" ID-GBIF-Occurrence="1806563522" country="Saudi Arabia" latitude="19.838818" location="Shada Al Ala" longLatPrecision="1" longitude="41.310066" municipality="Al Makhwa" pageId="13" pageNumber="100" specimenCount="1" specimenCount-female="1" stateProvince="Baha">
<emphasis id="B9062C501F64FFEAF19BCECBD6E9FD84" bold="true" box="[199,437,505,530]" pageId="13" pageNumber="100">Material examined:</emphasis>
Total two specimens:
<specimenCount id="9D743BCB1F64FFEAF3E0CECBD5B5FD87" box="[700,745,505,529]" pageId="13" pageNumber="100" type="female">1 ♀</specimenCount>
, “
<collectingCountry id="F365B0D21F64FFEAF258CECBD41CFD87" box="[772,832,505,529]" name="Saudi Arabia" pageId="13" pageNumber="100">KSA</collectingCountry>
, Al
<collectingRegion id="49B63EA01F64FFEAF229CECBD4EDFD84" box="[885,945,505,530]" country="Saudi Arabia" name="Al Bahah" pageId="13" pageNumber="100">Baha</collectingRegion>
,
<collectingMunicipality id="6BA96A381F64FFEAF2E3CECBD316FD84" box="[959,1098,505,530]" pageId="13" pageNumber="100">Al Makhwa</collectingMunicipality>
,
<location id="8EADA6991F64FFEAF504CECBD3ABFD84" LSID="urn:lsid:plazi:treatment:03DB41541F64FFEAF1CBCE79D62CF9B0:8EADA6991F64FFEAF504CECBD3ABFD84" box="[1112,1271,505,530]" country="Saudi Arabia" latitude="19.838818" longLatPrecision="1" longitude="41.310066" municipality="Al Makhwa" name="Shada Al Ala" pageId="13" pageNumber="100" stateProvince="Baha">Shada Al Ala</location>
,
<geoCoordinate id="EE4696851F64FFEAF459CEC8D2C2FD84" box="[1285,1438,505,530]" direction="north" orientation="latitude" pageId="13" pageNumber="100" precision="1" value="19.838818">N 19°50.329'</geoCoordinate>
<geoCoordinate id="EE4696851F64FFEAF1CBCD2FD67EFDA0" box="[151,290,541,566]" direction="east" orientation="longitude" pageId="13" pageNumber="100" precision="1" value="41.310066">E41°18.604'</geoCoordinate>
Alt
</materialsCitation>
.
<quantity id="4C8A5DA71F64FFEAF005CD2FD6F3FDA3" box="[345,431,541,566]" metricMagnitude="3" metricUnit="m" metricValue="1.563" pageId="13" pageNumber="100" unit="m" value="1563.0">1563 m</quantity>
,
<date id="FFCCD6821F64FFEAF0E6CD2CD507FDA0" box="[442,603,541,566]" pageId="13" pageNumber="100" value="2015-08-23">23. VIII. 2015</date>
, (LT)., 1 ♀, “N 19°51.762' E 41°18.089' Alt.
<materialsCitation id="3B1AFA1F1F64FFEAF50ECD2FD3C8FDCF" ID-GBIF-Occurrence="1806563517" collectingDate="2015-11-14" collectorName="Al Dafer H. &amp; Abdel-Dayem, H. H &amp; Elgharbawy &amp; Soliman &amp; Rasool, I." country="Turkey" elevation="1225" location="A" pageId="13" pageNumber="100" specimenCount="1" stateProvince="A">
<elevation id="005F17711F64FFEAF50ECD2FD3F4FDA3" box="[1106,1192,541,566]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="13" pageNumber="100" unit="m" value="1225.0">
<quantity id="4C8A5DA71F64FFEAF50ECD2FD3F4FDA3" box="[1106,1192,541,566]" metricMagnitude="3" metricUnit="m" metricValue="1.225" pageId="13" pageNumber="100" unit="m" value="1225.0">1225 m</quantity>
</elevation>
,
<collectingDate id="EF882F6A1F64FFEAF5E8CD2FD26AFDA0" box="[1204,1334,541,566]" pageId="13" pageNumber="100" value="2015-11-14">
<date id="FFCCD6821F64FFEAF5E8CD2FD26AFDA0" box="[1204,1334,541,566]" pageId="13" pageNumber="100" value="2015-11-14">14.XI.2015</date>
</collectingDate>
, (LT),
<collectorName id="268795941F64FFEAF4DCCD2FD7A7FDCF" pageId="13" pageNumber="100">Al Dafer H.</collectorName>
, M.S.
<collectorName id="268795941F64FFEAF018CD73D576FDCF" box="[324,554,577,602]" pageId="13" pageNumber="100">Abdel-Dayem, H. H</collectorName>
. Fadl,
<collectorName id="268795941F64FFEAF329CD73D5A1FDCC" box="[629,765,577,602]" pageId="13" pageNumber="100">Elgharbawy</collectorName>
, El
<collectingCountry id="F365B0D21F64FFEAF275CD73D426FDCC" box="[809,890,577,602]" name="Turkey" pageId="13" pageNumber="100">Turkey</collectingCountry>
,
<collectorName id="268795941F64FFEAF2D9CD73D4B9FDCC" box="[901,997,577,602]" pageId="13" pageNumber="100">Soliman</collectorName>
,
<collectingRegion id="49B63EA01F64FFEAF2ACCD73D358FDCF" box="[1008,1028,577,601]" country="Gambia" name="Upper River" pageId="13" pageNumber="100">A</collectingRegion>
&amp;
<collectorName id="268795941F64FFEAF574CD73D3C8FDCF" box="[1064,1172,577,602]" pageId="13" pageNumber="100">Rasool, I.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BCDF0421F64FFEAF19BCD57D243FB2B" blockId="13.[151,1438,402,1574]" pageId="13" pageNumber="100">
<emphasis id="B9062C501F64FFEAF19BCD57D601FDE8" bold="true" box="[199,349,613,638]" pageId="13" pageNumber="100">Recognition.</emphasis>
Large body size (
<figureCitation id="1349ECC71F64FFEAF37ACD57D526FDE8" box="[550,634,613,638]" captionStart="FIGURES 2729" captionStartId="11.[151,264,730,752]" captionTargetBox="[205,1352,205,707]" captionTargetId="figure@11.[173,1367,193,714]" captionTargetPageId="11" captionText="FIGURES 2729. Habitus of Lebia and Matabele: (27) Syntype. L. natalensis Chaudoir, 1871 (28) L. spec. (29) M. arabica Mateu, 1986." httpUri="https://zenodo.org/record/1172366/files/figure.png" pageId="13" pageNumber="100">Fig. 29</figureCitation>
), TBL
<quantity id="4C8A5DA71F64FFEAF397CD54D41AFDEB" box="[715,838,614,638]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" metricValueMax="1.4" metricValueMin="1.2" pageId="13" pageNumber="100" unit="mm" value="13.0" valueMax="14.0" valueMin="12.0">1214 mm</quantity>
,
<emphasis id="B9062C501F64FFEAF20ECD57D4F6FDEB" box="[850,938,613,637]" italics="true" pageId="13" pageNumber="100">Colour:</emphasis>
Head, pronotum and elytra black; antennae, mouthparts, femora and tibiae brunneous; tarsomeres red; elytra with two transverse testaceous bands, one on the post humeri extending from intervals IIIVIII and second on pre-apical of elytra extending from intervals III-VI.
<emphasis id="B9062C501F64FFEAF1CBCDE3D616FD7F" box="[151,330,721,745]" italics="true" pageId="13" pageNumber="100">Microsculpture:</emphasis>
More or less depressed isodiametric meshes on the head and pronotum surface; elytra with strongly isodiametric mesh pattern.
<emphasis id="B9062C501F64FFEAF318CDC7D5D5FC9B" box="[580,649,757,781]" italics="true" pageId="13" pageNumber="100">Head:</emphasis>
convex, HL
<quantity id="4C8A5DA71F64FFEAF26CCDC7D486FC9B" box="[816,986,757,782]" metricMagnitude="-3" metricUnit="m" metricValue="2.61" metricValueMax="2.75" metricValueMin="2.47" pageId="13" pageNumber="100" unit="mm" value="2.61" valueMax="2.75" valueMin="2.47">2.472.75 mm</quantity>
and HW
<quantity id="4C8A5DA71F64FFEAF506CDC7D259FC9B" box="[1114,1285,757,782]" metricMagnitude="-3" metricUnit="m" metricValue="2.485" metricValueMax="2.6" metricValueMin="2.37" pageId="13" pageNumber="100" unit="mm" value="2.485" valueMax="2.6" valueMin="2.37">2.372.60 mm</quantity>
, with bulky, outstanding and convex eyes, tempora short, narrowed behind at right angle (
<figureCitation id="1349ECC71F64FFEAF2AFCC2BD31AFCA4" box="[1011,1094,793,818]" captionStart="FIGURES3037" captionStartId="11.[151,264,1938,1960]" captionTargetBox="[347,1211,836,1903]" captionTargetId="figure@11.[347,1211,824,1910]" captionTargetPageId="11" captionText="FIGURES3037. Head and pronotum of Lebia: (30) L.auberti (Fairmaire,1892), (31) L.melanacra (Chaudoir,1878), (32) L. melanura (Dejean, 1831), (33) L. raeesae sp. n. (34) L. natalensis Chaudoir, 1871, (35) L. nilotica Chaudoir, 1871, (36) L. spec.(37) Matabele arabica Mateu, 1986." httpUri="https://zenodo.org/record/1172368/files/figure.png" pageId="13" pageNumber="100">Fig. 37</figureCitation>
), dorsum of head punctate and wrinkled on the frons and neck; labrum rounded anteriorly, not transverse; mentum emarginated with bifid tooth and well developed epilobes, last labial palpi fusiform, finely and sparsely pubescent; first segment of labial palpi longer than the second which with four setae; antennae elongate extending beyond the base of post-humeral testaceous band.
<emphasis id="B9062C501F64FFEAF00BCC98D693FC57" box="[343,463,938,961]" italics="true" pageId="13" pageNumber="100">Pronotum:</emphasis>
transverse and wider than head with eyes, PL
<quantity id="4C8A5DA71F64FFEAF2BFCC98D3DBFC57" box="[995,1159,938,962]" metricMagnitude="-3" metricUnit="m" metricValue="1.995" metricValueMax="2.12" metricValueMin="1.87" pageId="13" pageNumber="100" unit="mm" value="1.995" valueMax="2.12" valueMin="1.87">1.872.12 mm</quantity>
and PW
<quantity id="4C8A5DA71F64FFEAF5AECC9BD2C4FC57" box="[1266,1432,937,962]" metricMagnitude="-3" metricUnit="m" metricValue="2.875" metricValueMax="3.1" metricValueMin="2.65" pageId="13" pageNumber="100" unit="mm" value="2.875" valueMax="3.1" valueMin="2.65">2.653.10 mm</quantity>
, weakly convex (
<figureCitation id="1349ECC71F64FFEAF007CCFFD6E9FC70" box="[347,437,973,998]" captionStart="FIGURES3037" captionStartId="11.[151,264,1938,1960]" captionTargetBox="[347,1211,836,1903]" captionTargetId="figure@11.[347,1211,824,1910]" captionTargetPageId="11" captionText="FIGURES3037. Head and pronotum of Lebia: (30) L.auberti (Fairmaire,1892), (31) L.melanacra (Chaudoir,1878), (32) L. melanura (Dejean, 1831), (33) L. raeesae sp. n. (34) L. natalensis Chaudoir, 1871, (35) L. nilotica Chaudoir, 1871, (36) L. spec.(37) Matabele arabica Mateu, 1986." httpUri="https://zenodo.org/record/1172368/files/figure.png" pageId="13" pageNumber="100">Fig. 37</figureCitation>
); rounded anteriorly and moderately narrowed near basal angles; base of pronotum projecting strongly in the middle; median impression of pronotum fairly deep; dorsum strongly punctate, sculptured and wrinkled.
<emphasis id="B9062C501F64FFEAF0E4CB27D55BFBBB" box="[440,519,1045,1069]" italics="true" pageId="13" pageNumber="100">Elytra:</emphasis>
Elongate but fairly broad, EL
<quantity id="4C8A5DA71F64FFEAF23BCB27D350FBBB" box="[871,1036,1045,1070]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="7.75" metricValueMin="7.25" pageId="13" pageNumber="100" unit="mm" value="7.5" valueMax="7.75" valueMin="7.25">7.257.75 mm</quantity>
and EW 5.0
<quantity id="4C8A5DA71F64FFEAF5F7CB27D258FBBB" box="[1195,1284,1045,1070]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="13" pageNumber="100" unit="mm" value="5.5">5.5 mm</quantity>
, convex with bordered base; apex of elytra obliquely truncate, sinuate and external angles pointed; striae deep and punctate; intervals convex, finely and sparsely punctuate. All tarsomeres with spines on inner side, tarsomeres I III expended and tarsomeres IV markedly excavate; tarsomere IV bilobed in fore legs; claws pectinate with 89 teeth.
<emphasis id="B9062C501F64FFEAF1CBCB97D656FB2B" box="[151,266,1189,1213]" italics="true" pageId="13" pageNumber="100">Abdomen:</emphasis>
Ventrum of the body pubescent; abdominal sternum VII truncate and bi-setose in both sexes.
</paragraph>
</subSubSection>
<subSubSection id="C368A3C91F64FFEAF19BCBFBD62CF9B0" pageId="13" pageNumber="100" type="description">
<paragraph id="8BCDF0421F64FFEAF19BCBFBD663FABC" blockId="13.[151,1438,402,1574]" pageId="13" pageNumber="100">
<emphasis id="B9062C501F64FFEAF19BCBFBD661FB77" box="[199,317,1225,1249]" italics="true" pageId="13" pageNumber="100">Aedeagus:</emphasis>
In lateral view aedeagus is robust and almost straight ventrally, scarcely sinuate near base, apex slightly inclined and drawn out into only a short point; when viewed from above it looks broad and rounded distally (
<bibRefCitation id="EFE38DB31F64FFEAF1C2CA20D66EFABC" box="[158,306,1297,1322]" pageId="13" pageNumber="100" refId="ref11942" refString="Mateu, J. (1986) Lebiinae and Brachininae of Saudi Arabia (Coleoptera: Carabidae). Fauna of Saudi Arabia, 8, 198 - 217." type="journal article">Mateu, 1986</bibRefCitation>
).
</paragraph>
<paragraph id="8BCDF0421F64FFEAF19BCA07D53CFA2C" blockId="13.[151,1438,402,1574]" pageId="13" pageNumber="100">
<emphasis id="B9062C501F64FFEAF19BCA07D676FAD8" bold="true" box="[199,298,1333,1358]" pageId="13" pageNumber="100">Affinity.</emphasis>
This species resembles to
<taxonomicName id="4C728BC11F64FFEAF30ACA04D587FADB" authorityName="Peringuey" authorityYear="1896" box="[598,731,1333,1357]" class="Insecta" family="Carabidae" genus="Matabele" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="miranda">
<emphasis id="B9062C501F64FFEAF30ACA04D587FADB" box="[598,731,1333,1357]" italics="true" pageId="13" pageNumber="100">M. miranda</emphasis>
</taxonomicName>
, the only other member of the genus, in colouration, body size and markings of elytra, but can be distinguished by smooth intervals of elytra, rounded anterior margin of labrum, obtuse head, strongly pointed external angles of elytra, tarsomeres rather more pubescent and colour of tarsomeres and femur reddish brown (
<bibRefCitation id="EFE38DB31F64FFEAF09CCA90D508FA2C" box="[448,596,1441,1466]" pageId="13" pageNumber="100" refId="ref11942" refString="Mateu, J. (1986) Lebiinae and Brachininae of Saudi Arabia (Coleoptera: Carabidae). Fauna of Saudi Arabia, 8, 198 - 217." type="journal article">Mateu, 1986</bibRefCitation>
).
</paragraph>
<paragraph id="8BCDF0421F64FFEAF19BCAF7D62CF9B0" blockId="13.[151,1438,402,1574]" pageId="13" pageNumber="100">
<emphasis id="B9062C501F64FFEAF19BCAF7D55BFA48" bold="true" box="[199,519,1477,1502]" pageId="13" pageNumber="100">Geographical distribution.</emphasis>
The species was described from
<collectingCountry id="F365B0D21F64FFEAF2D5CAF7D342FA48" box="[905,1054,1477,1502]" name="Saudi Arabia" pageId="13" pageNumber="100">Saudi Arabia</collectingCountry>
(
<bibRefCitation id="EFE38DB31F64FFEAF573CAF4D39EFA48" box="[1071,1218,1477,1502]" pageId="13" pageNumber="100" refId="ref11942" refString="Mateu, J. (1986) Lebiinae and Brachininae of Saudi Arabia (Coleoptera: Carabidae). Fauna of Saudi Arabia, 8, 198 - 217." type="journal article">Mateu, 1986</bibRefCitation>
) and then reported from
<collectingCountry id="F365B0D21F64FFEAF187CAD8D67CF994" box="[219,288,1514,1538]" name="Oman" pageId="13" pageNumber="100">Oman</collectingCountry>
(Anichtchenko, 2016). Only found in Arabian Peninsula (Anichtchenko, 2016) and exemplifies Arabian chorotype.
</paragraph>
</subSubSection>
</treatment>
</document>