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<document id="CAA45973527385158F1EE86853BD346D" ID-DOI="10.18590/euscorpius.2007.vol2007.iss54.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="4673271" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617911562369" checkinUser="carolina" docAuthor="Kovařík, František" docDate="2007" docId="E65A542EFFD5FFEFFC75FB51FC0E82BF" docLanguage="en" docName="Euscorpius.54.1-7.pdf" docOrigin="Euscorpius 54 (54)" docSource="https://mds.marshall.edu/euscorpius/vol2007/iss54/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Tityus (Archaeotityus) mraceki Kovařík 2007, sp. n." docType="treatment" docUuid="14D883C0-E78B-42E3-87B7-00F5AAA84933" docUuidSource="ZooBank" docVersion="4" lastPageNumber="3" masterDocId="1A632C56FFD7FFEBFFE4FFDAFFF88409" masterDocTitle="Description of Tityus mraceki sp. n. from Colombia and synonymization of T. meridanus González-Sponga with T. nematochirus Mello-Leitão (Scorpiones: Buthidae)" masterLastPageNumber="7" masterPageNumber="1" pageNumber="1" updateTime="1698902843396" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="E9C3B2351B8B573A4F2598BD9B5655CE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DCA13AEEC9CF1E54CA4219721B74E18F">
<mods:title id="1D82B1BFEE1368F85D980C803CCDA95C">Description of Tityus mraceki sp. n. from Colombia and synonymization of T. meridanus González-Sponga with T. nematochirus Mello-Leitão (Scorpiones: Buthidae)</mods:title>
</mods:titleInfo>
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<mods:role id="BFA79691AFA2CE7F754B86341705D281">
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</mods:role>
<mods:namePart id="6D272EFFE0E2634AF0F234275E5DEDB8">Kovařík, František</mods:namePart>
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<mods:typeOfResource id="30B8B27BC78309C8238451F048321B67">text</mods:typeOfResource>
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<mods:title id="67AF7045F72CBEDEC489880C0C73637E">Euscorpius</mods:title>
</mods:titleInfo>
<mods:part id="654F5F236C8FF96C3D2CF66B20A978A7">
<mods:date id="256BB3E2F6AC6988D1A19102641DD128">2007</mods:date>
<mods:detail id="86DD20486FCE06A0230DEC50A99E3582" type="pubDate">
<mods:number id="3E1F2E7D064193BE821CEFBDD034CB13">2007-12-31</mods:number>
</mods:detail>
<mods:detail id="816141E4B7CBFFECC60AF5D527DCB576" type="volume">
<mods:number id="FA38781C3C23917E79D9ED6C379E75D2">54</mods:number>
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<mods:detail id="6AEF404BE4785147646C341F72C07C31" type="issue">
<mods:number id="6CC1454CDF6CFB149B6ACCBBA810F367">54</mods:number>
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<mods:start id="4ABDC908EFF60F84A044A46D474F3054">1</mods:start>
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<mods:url id="9F66C425E863F035E58F48C69E7ACFA6">https://mds.marshall.edu/euscorpius/vol2007/iss54/1/</mods:url>
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<mods:classification id="183B9ADC549E0AC32D1DE6EA430E1C0C">journal article</mods:classification>
<mods:identifier id="07AF51653B3D623FB277E54A36344FE8" type="DOI">10.18590/euscorpius.2007.vol2007.iss54.1</mods:identifier>
<mods:identifier id="CC47EA6DA1CFD607681D2CB546B3AFED" type="ISSN">1536-9307</mods:identifier>
<mods:identifier id="A2435DD39671AB9638E25B0457C8C155" type="Zenodo-Dep">4673271</mods:identifier>
<mods:identifier id="D0EC964FB7772756B4E428D6CBF6AE81" type="ZooBank">4C949910-48EF-46D9-88D5-877939939690</mods:identifier>
</mods:mods>
<treatment id="E65A542EFFD5FFEFFC75FB51FC0E82BF" LSID="urn:lsid:zoobank.org:act:14D883C0-E78B-42E3-87B7-00F5AAA84933" httpUri="http://treatment.plazi.org/id/E65A542EFFD5FFEFFC75FB51FC0E82BF" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1">
<subSubSection id="26E9B6B3FFD5FFE9FC75FB51FAB580AC" box="[913,1357,1163,1189]" pageId="2" pageNumber="1" type="nomenclature">
<paragraph id="6E4CE538FFD5FFE9FC75FB51FAB580AC" blockId="2.[913,1357,1163,1219]" box="[913,1357,1163,1189]" pageId="2" pageNumber="1">
<heading id="35045254FFD5FFE9FC75FB51FAB580AC" box="[913,1357,1163,1189]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="1" reason="4">
<taxonomicName id="A9F39EBBFFD5FFE9FC75FB51FAF080AC" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[913,1288,1163,1189]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov." subGenus="Archaeotityus">
<emphasis id="5C87392AFFD5FFE9FC75FB51FAF080AC" bold="true" box="[913,1288,1163,1189]" pageId="2" pageNumber="1">
<emphasis id="5C87392AFFD5FFE9FC75FB51FC2480AC" bold="true" box="[913,988,1163,1189]" italics="true" pageId="2" pageNumber="1">Tityus</emphasis>
(
<emphasis id="5C87392AFFD5FFE9FC09FB51FB6D80AC" bold="true" box="[1005,1173,1163,1189]" italics="true" pageId="2" pageNumber="1">Archaeotityus</emphasis>
)
<emphasis id="5C87392AFFD5FFE9FB42FB51FAF080AC" bold="true" box="[1190,1288,1163,1189]" italics="true" pageId="2" pageNumber="1">mraceki</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47B48451FFD5FFE9FAEBFB51FAB580AC" box="[1295,1357,1163,1189]" pageId="2" pageNumber="1" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="26E9B6B3FFD5FFE9FBE2FB76FB2180CA" box="[1030,1241,1196,1219]" pageId="2" pageNumber="1" type="description">
<paragraph id="6E4CE538FFD5FFE9FBE2FB76FB2180CA" blockId="2.[913,1357,1163,1219]" box="[1030,1241,1196,1219]" pageId="2" pageNumber="1">
(
<figureCitation id="F6C8F9BDFFD5FFE9FBE9FB76FB8B80CA" box="[1037,1139,1196,1219]" captionStart-0="Figures 1–3" captionStart-1="Figure 4" captionStartId-0="3.[192,278,1900,1923]" captionStartId-1="5.[192,268,1046,1069]" captionTargetBox-0="[191,1440,191,1866]" captionTargetBox-1="[191,1440,191,1014]" captionTargetId-0="figure-8@3.[192,1439,192,1865]" captionTargetId-1="figure-7@5.[192,1439,192,1013]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionText-0="Figures 1–3: Tityus mraceki sp. n., male holotype. 1. Dorsal view. 2. Ventral view. 3. Lateral view of metasoma." captionText-1="Figure 4: Type locality of Tityus mraceki sp. n., mangrove swamp on the Pacific shore, Colombia, Juanchaco near Buenaventura." figureDoi-0="http://doi.org/10.5281/zenodo.4673275" figureDoi-1="http://doi.org/10.5281/zenodo.4673277" httpUri-0="https://zenodo.org/record/4673275/files/figure.png" httpUri-1="https://zenodo.org/record/4673277/files/figure.png" pageId="2" pageNumber="1">Figs. 1–4</figureCitation>
,
<tableCitation id="2371D083FFD5FFE9FB9BFB76FB3680CA" box="[1151,1230,1196,1219]" captionStart="Table 1" captionStartId="4.[192,257,997,1020]" captionText="Table 1: Measurements (in mm) of the male holotype of Tityus mraceki sp. n. * Includes intersegment membranes." pageId="2" pageNumber="1">Table 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="26E9B6B3FFD5FFE9FCDAFB33FC6F81B1" pageId="2" pageNumber="1" type="materials_examined">
<paragraph id="6E4CE538FFD5FFE9FCDAFB33FAA08134" blockId="2.[830,1441,1257,1341]" pageId="2" pageNumber="1">
<materialsCitation id="DE9BEF65FFD5FFE9FCDAFB33FBCB80F6" box="[830,1075,1257,1280]" collectionCode="LOCALITY, AND" pageId="2" pageNumber="1" specimenCount="1" typeStatus="holotype">
<typeStatus id="B1485B9AFFD5FFE9FCDAFB33FC8F80F6" box="[830,887,1257,1280]" pageId="2" pageNumber="1">TYPE</typeStatus>
<collectionCode id="08E27DFDFFD5FFE9FC6DFB36FBCB80F6" box="[905,1075,1260,1279]" pageId="2" pageNumber="1">LOCALITY AND</collectionCode>
</materialsCitation>
<materialsCitation id="DE9BEF65FFD5FFE9FBA1FB36FB0D8134" collectionCode="REPOSITORY" country="Colombia" county="Pacific" location="Juanchaco" municipality="Buenaventura" pageId="2" pageNumber="1" specimenCount="1" typeStatus="holotype">
<typeStatus id="B1485B9AFFD5FFE9FBA1FB36FB8280F6" box="[1093,1146,1260,1279]" pageId="2" pageNumber="1">TYPE</typeStatus>
<collectionCode id="08E27DFDFFD5FFE9FB69FB36FAE880F6" box="[1165,1296,1260,1279]" pageId="2" pageNumber="1">REPOSITORY</collectionCode>
.
<emphasis id="5C87392AFFD5FFE9FACEFB33FA618109" bold="true" box="[1322,1433,1257,1280]" pageId="2" pageNumber="1">
<collectingCountry id="16E4A5A8FFD5FFE9FACEFB33FA618109" box="[1322,1433,1257,1280]" name="Colombia" pageId="2" pageNumber="1">Colombia</collectingCountry>
</emphasis>
,
<location id="6B2CB3E3FFD5FFE9FCDAFAD2FC558116" LSID="urn:lsid:plazi:treatment:E65A542EFFD5FFEFFC75FB51FC0E82BF:6B2CB3E3FFD5FFE9FCDAFAD2FC558116" box="[830,941,1288,1311]" country="Colombia" county="Pacific" municipality="Buenaventura" name="Juanchaco" pageId="2" pageNumber="1">Juanchaco</location>
near
<collectingMunicipality id="8E287F42FFD5FFE9FC0AFAD2FB7A8116" box="[1006,1154,1288,1311]" pageId="2" pageNumber="1">Buenaventura</collectingMunicipality>
, mangrove swamp on the
<collectingCounty id="872D9DB4FFD5FFE9FCDAFAFCFC708134" box="[830,904,1318,1341]" pageId="2" pageNumber="1">Pacific</collectingCounty>
shore (
<figureCitation id="F6C8F9BDFFD5FFE9FC3CFAFCFBE18134" box="[984,1049,1318,1341]" captionStart="Figure 4" captionStartId="5.[192,268,1046,1069]" captionTargetBox="[191,1440,191,1014]" captionTargetId="figure-7@5.[192,1439,192,1013]" captionTargetPageId="5" captionText="Figure 4: Type locality of Tityus mraceki sp. n., mangrove swamp on the Pacific shore, Colombia, Juanchaco near Buenaventura." figureDoi="http://doi.org/10.5281/zenodo.4673277" httpUri="https://zenodo.org/record/4673277/files/figure.png" pageId="2" pageNumber="1">Fig. 4</figureCitation>
); author´s collection
</materialsCitation>
(
<collectionCode id="08E27DFDFFD5FFE9FAE7FAFCFAB48134" box="[1283,1356,1318,1341]" pageId="2" pageNumber="1">FKCP</collectionCode>
).
</paragraph>
<paragraph id="6E4CE538FFD5FFE9FCDAFABEFC6F81B1" blockId="2.[830,1440,1380,1464]" pageId="2" pageNumber="1">
<materialsCitation id="DE9BEF65FFD5FFE9FCDAFABEFC6F81B1" collectingDate="1995-09-15" collectingDateMax="1995-09-20" collectingDateMin="1995-09-15" collectionCode="TYPE, MATERIAL" collectorName="Z. Mracek." country="Colombia" location="Juanchaco" municipality="Buenaventura" pageId="2" pageNumber="1" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectionCode id="08E27DFDFFD5FFE9FCDAFABEFC0D8173" box="[830,1013,1380,1403]" pageId="2" pageNumber="1">TYPE MATERIAL</collectionCode>
.
<emphasis id="5C87392AFFD5FFE9FBE9FABEFB848172" bold="true" box="[1037,1148,1380,1403]" pageId="2" pageNumber="1">
<collectingCountry id="16E4A5A8FFD5FFE9FBE9FABEFB848172" box="[1037,1148,1380,1403]" name="Colombia" pageId="2" pageNumber="1">Colombia</collectingCountry>
</emphasis>
,
<location id="6B2CB3E3FFD5FFE9FB70FABEFAFB8172" LSID="urn:lsid:plazi:treatment:E65A542EFFD5FFEFFC75FB51FC0E82BF:6B2CB3E3FFD5FFE9FB70FABEFAFB8172" box="[1172,1283,1380,1403]" country="Colombia" municipality="Buenaventura" name="Juanchaco" pageId="2" pageNumber="1">Juanchaco</location>
near
<collectingMunicipality id="8E287F42FFD5FFE9FAB7FABEFC698190" pageId="2" pageNumber="1">Buenaventura</collectingMunicipality>
,
<date id="1A4DC3F8FFD5FFE9FC47FA58FB528190" box="[931,1194,1410,1433]" pageId="2" pageNumber="1" value="1995-09-15" valueMax="1995-09-20" valueMin="1995-09-15">
<collectingDate id="0A093A10FFD5FFE9FC47FA58FB528190" box="[931,1194,1410,1433]" pageId="2" pageNumber="1" value="1995-09-15" valueMax="1995-09-20" valueMin="1995-09-15">15–20 September 1995</collectingDate>
</date>
,
<typeStatus id="B1485B9AFFD5FFE9FB5BFA58FAE48190" box="[1215,1308,1410,1433]" pageId="2" pageNumber="1" type="holotype">holotype</typeStatus>
<specimenCount id="78F52EB1FFD5FFE9FACFFA58FAC78190" box="[1323,1343,1410,1433]" pageId="2" pageNumber="1" type="male">♂</specimenCount>
, leg.
<collectorName id="C30680EEFFD5FFE9FA6DFA58FC6F81B1" pageId="2" pageNumber="1">Z. Mráček.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="26E9B6B3FFD5FFE9FCDAFA04FC19821D" pageId="2" pageNumber="1" type="etymology">
<paragraph id="6E4CE538FFD5FFE9FCDAFA04FC19821D" blockId="2.[830,1440,1502,1556]" pageId="2" pageNumber="1">
ETYMOLOGY. Named after Zdeněk Mráček, who collected the
<typeStatus id="B1485B9AFFD5FFE9FC48FA27FC22821D" box="[940,986,1533,1556]" pageId="2" pageNumber="1">type</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="26E9B6B3FFD5FFEFFCDAF9E3FD3C8076" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1" type="diagnosis">
<paragraph id="6E4CE538FFD5FFEFFCDAF9E3FD3C8076" blockId="2.[830,1441,1593,1892]" lastBlockId="4.[192,802,1097,1151]" lastPageId="4" lastPageNumber="3" pageId="2" pageNumber="1">
DIAGNOSIS: Adult male
<quantity id="A90B48DDFFD5FFE9FBADF9E3FB6E8259" box="[1097,1174,1593,1616]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="1" unit="mm" value="50.0">50 mm</quantity>
long. Female unknown. Color blotched. For habitus see
<figureCitation id="F6C8F9BDFFD5FFE9FB7EF98DFAD38267" box="[1178,1323,1623,1646]" captionStart="Figures 1–3" captionStartId="3.[192,278,1900,1923]" captionTargetBox="[191,1440,191,1866]" captionTargetId="figure-8@3.[192,1439,192,1865]" captionTargetPageId="3" captionText="Figures 1–3: Tityus mraceki sp. n., male holotype. 1. Dorsal view. 2. Ventral view. 3. Lateral view of metasoma." figureDoi="http://doi.org/10.5281/zenodo.4673275" httpUri="https://zenodo.org/record/4673275/files/figure.png" pageId="2" pageNumber="1">Figs. 1 and 2</figureCitation>
. Carapace, mesosoma and metasoma densely granulate. Pectines with 14 teeth. Stigmata are elongate. Rhomboidal subaculear tooth with four granules in two rows. Movable finger of pedipalp with 15 or 16 rows of granules, which include external and internal granules. First metasomal segment with 10 carinae, second to fourth metasomal segments with eight carinae, fifth metasomal segment with only three ventral carinae. All carinae parallel and composed of equally sized granules. Chela of pedipalp narrow. Male metasoma long.
</paragraph>
</subSubSection>
<caption id="3A8CB5B0FFD4FFE8FF24F8B6FAED838A" ID-DOI="http://doi.org/10.5281/zenodo.4673275" ID-Zenodo-Dep="4673275" box="[192,1301,1900,1923]" httpUri="https://zenodo.org/record/4673275/files/figure.png" pageId="3" pageNumber="2" startId="3.[192,278,1900,1923]" targetBox="[191,1440,191,1866]" targetPageId="3">
<paragraph id="6E4CE538FFD4FFE8FF24F8B6FAED838A" blockId="3.[192,1301,1900,1923]" box="[192,1301,1900,1923]" pageId="3" pageNumber="2">
<emphasis id="5C87392AFFD4FFE8FF24F8B6FEB5838A" bold="true" box="[192,333,1900,1923]" pageId="3" pageNumber="2">Figures 1–3:</emphasis>
<taxonomicName id="A9F39EBBFFD4FFE8FEBDF8B4FE1F838A" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[345,487,1902,1923]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD4FFE8FEBDF8B4FE1F838A" box="[345,487,1902,1923]" italics="true" pageId="3" pageNumber="2">Tityus mraceki</emphasis>
</taxonomicName>
<emphasis id="5C87392AFFD4FFE8FE0AF8B4FDDB838A" bold="true" box="[494,547,1902,1923]" pageId="3" pageNumber="2">
<taxonomicNameLabel id="47B48451FFD4FFE8FE0AF8B4FDDB838A" box="[494,547,1902,1923]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male holotype.
<emphasis id="5C87392AFFD4FFE8FD20F8B4FD2E838A" bold="true" box="[708,726,1902,1923]" pageId="3" pageNumber="2">1.</emphasis>
Dorsal view.
<emphasis id="5C87392AFFD4FFE8FCB9F8B4FC97838A" bold="true" box="[861,879,1902,1923]" pageId="3" pageNumber="2">2.</emphasis>
Ventral view.
<emphasis id="5C87392AFFD4FFE8FC1AF8B4FBE8838A" bold="true" box="[1022,1040,1902,1923]" pageId="3" pageNumber="2">3.</emphasis>
Lateral view of metasoma.
</paragraph>
</caption>
<paragraph id="6E4CE538FFD3FFEFFE96FF13FD7D87C9" pageId="4" pageNumber="3">
<table id="1CF31798FFD30014FF11FF13FD4187C9" box="[245,697,201,960]" gridcols="2" gridrows="27" pageId="4" pageNumber="3">
<tr id="D0C3E77AFFD30014FF11FF13FD4184EA" box="[245,697,201,227]" gridrow="0" pageId="4" pageNumber="3">
<th id="93128E06FFD30014FF11FF13FD4184EA" box="[245,697,201,227]" colspan="2" colspanRight="1" gridcol="0" gridrow="0" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE96FF13FD8884EA" bold="true" box="[370,624,201,227]" pageId="4" pageNumber="3">
<taxonomicName id="A9F39EBBFFD3FFEFFE96FF13FDDE84EA" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[370,550,201,227]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD3FFEFFE96FF13FDDE84EA" bold="true" box="[370,550,201,227]" italics="true" pageId="4" pageNumber="3">Tityus mraceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47B48451FFD3FFEFFDC9FF13FD8884EA" box="[557,624,201,227]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="D0C3E77AFFD30014FF11FF31FD41850B" box="[245,697,235,258]" gridrow="1" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FF31FD41850B" box="[245,697,235,258]" colspan="2" colspanRight="1" gridcol="0" gridrow="1" pageId="4" pageNumber="3">Male Holotype</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FED6FD418528" box="[245,697,268,289]" gridrow="2" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FED6FE348528" box="[245,460,268,289]" gridcol="0" gridrow="2" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFF11FED6FE348528" bold="true" box="[245,460,268,289]" pageId="4" pageNumber="3">Total length *</emphasis>
</td>
<td id="93128E06FFD30014FDAAFED6FD418528" box="[590,697,268,289]" gridcol="1" gridrow="2" pageId="4" pageNumber="3">50.7</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FEFDFD418534" box="[245,697,295,317]" gridrow="3" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FEFDFE348534" box="[245,460,295,317]" gridcol="0" gridrow="3" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFF11FEF2FE428534" bold="true" box="[245,442,296,317]" pageId="4" pageNumber="3">Carapace length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFEFDFD418534" box="[590,697,295,317]" gridcol="1" gridrow="3" pageId="4" pageNumber="3">4.5</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FE99FD418551" box="[245,697,323,344]" gridrow="4" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FE99FE348551" box="[245,460,323,344]" gridcol="0" gridrow="4" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE9EFE99FE4D8551" bold="true" box="[378,437,323,344]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFE99FD418551" box="[590,697,323,344]" gridcol="1" gridrow="4" pageId="4" pageNumber="3">4.6</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FEBAFD41857C" box="[245,697,352,373]" gridrow="5" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FEBAFD41857C" box="[245,697,352,373]" colspan="2" colspanRight="1" gridcol="0" gridrow="5" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFF11FEBAFE75857C" bold="true" box="[245,397,352,373]" pageId="4" pageNumber="3">Metasoma and</emphasis>
</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FEA1FD418598" box="[245,697,379,401]" gridrow="6" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FEA1FE348598" box="[245,460,379,401]" gridcol="0" gridrow="6" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFEC2FEA6FE348598" bold="true" box="[294,460,380,401]" pageId="4" pageNumber="3">Telson length *</emphasis>
</td>
<td id="93128E06FFD30014FDAAFEA1FD418598" box="[590,697,379,401]" gridcol="1" gridrow="6" pageId="4" pageNumber="3">32.7</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FE4DFD4185A5" box="[245,697,407,428]" gridrow="7" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FE4DFE3485A5" box="[245,460,407,428]" gridcol="0" gridrow="7" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFEE6FE4DFE4285A5" bold="true" box="[258,442,407,428]" pageId="4" pageNumber="3">Segment I length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFE4DFD4185A5" box="[590,697,407,428]" gridcol="1" gridrow="7" pageId="4" pageNumber="3">4.0</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FE69FD4185C1" box="[245,697,435,456]" gridrow="8" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FE69FE3485C1" box="[245,460,435,456]" gridcol="0" gridrow="8" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE9DFE69FE4B85C1" bold="true" box="[377,435,435,456]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFE69FD4185C1" box="[590,697,435,456]" gridcol="1" gridrow="8" pageId="4" pageNumber="3">2.0</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FE14FD4185ED" box="[245,697,462,484]" gridrow="9" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FE14FE3485ED" box="[245,460,462,484]" gridcol="0" gridrow="9" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFEE6FE15FE4285ED" bold="true" box="[258,442,463,484]" pageId="4" pageNumber="3">Segment II length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFE14FD4185ED" box="[590,697,462,484]" gridcol="1" gridrow="9" pageId="4" pageNumber="3">5.2</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FE30FD4185F6" box="[245,697,490,511]" gridrow="10" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FE30FE3485F6" box="[245,460,490,511]" gridcol="0" gridrow="10" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE9DFE30FE4B85F6" bold="true" box="[377,435,490,511]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFE30FD4185F6" box="[590,697,490,511]" gridcol="1" gridrow="10" pageId="4" pageNumber="3">1.9</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FDDFFD418612" box="[245,697,517,539]" gridrow="11" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FDDFFE348612" box="[245,460,517,539]" gridcol="0" gridrow="11" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFEE6FDDCFE3A8612" bold="true" box="[258,450,518,539]" pageId="4" pageNumber="3">Segment III length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFDDFFD418612" box="[590,697,517,539]" gridcol="1" gridrow="11" pageId="4" pageNumber="3">5.9</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FDFBFD41863E" box="[245,697,545,567]" gridrow="12" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FDFBFE34863E" box="[245,460,545,567]" gridcol="0" gridrow="12" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE9BFDF8FE41863E" bold="true" box="[383,441,546,567]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFDFBFD41863E" box="[590,697,545,567]" gridcol="1" gridrow="12" pageId="4" pageNumber="3">1.9</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FDE7FD41865B" box="[245,697,573,594]" gridrow="13" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FDE7FE34865B" box="[245,460,573,594]" gridcol="0" gridrow="13" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFEE6FDE7FE38865B" bold="true" box="[258,448,573,594]" pageId="4" pageNumber="3">Segment IV length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFDE7FD41865B" box="[590,697,573,594]" gridcol="1" gridrow="13" pageId="4" pageNumber="3">6.4</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FD82FD418667" box="[245,697,600,622]" gridrow="14" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FD82FE348667" box="[245,460,600,622]" gridcol="0" gridrow="14" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE64FD83FE438667" bold="true" box="[384,443,601,622]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFD82FD418667" box="[590,697,600,622]" gridcol="1" gridrow="14" pageId="4" pageNumber="3">1.9</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FDAEFD418680" box="[245,697,628,649]" gridrow="15" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FDAEFE348680" box="[245,460,628,649]" gridcol="0" gridrow="15" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFEE6FDAEFE428680" bold="true" box="[258,442,628,649]" pageId="4" pageNumber="3">Segment V length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFDAEFD418680" box="[590,697,628,649]" gridcol="1" gridrow="15" pageId="4" pageNumber="3">6.3</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FD55FD4186AC" box="[245,697,655,677]" gridrow="16" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FD55FE3486AC" box="[245,460,655,677]" gridcol="0" gridrow="16" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE9DFD4AFE4B86AC" bold="true" box="[377,435,656,677]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFD55FD4186AC" box="[590,697,655,677]" gridcol="1" gridrow="16" pageId="4" pageNumber="3">2.3</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FD71FD4186C9" box="[245,697,683,704]" gridrow="17" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FD71FE3486C9" box="[245,460,683,704]" gridcol="0" gridrow="17" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFF12FD71FE7886C9" bold="true" box="[246,384,683,704]" pageId="4" pageNumber="3">Telson length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFD71FD4186C9" box="[590,697,683,704]" gridcol="1" gridrow="17" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFD81FD71FD9586C9" bold="true" box="[613,621,683,704]" pageId="4" pageNumber="3">-</emphasis>
</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FD12FD4186D4" box="[245,697,712,733]" gridrow="18" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FD12FD4186D4" box="[245,697,712,733]" colspan="2" colspanRight="1" gridcol="0" gridrow="18" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFF11FD12FEA886D4" bold="true" box="[245,336,712,733]" pageId="4" pageNumber="3">Pedipalp</emphasis>
</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FD39FD4186F0" box="[245,697,739,761]" gridrow="19" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FD39FE3486F0" box="[245,460,739,761]" gridcol="0" gridrow="19" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFECFFD3EFE4F86F0" bold="true" box="[299,439,740,761]" pageId="4" pageNumber="3">Femur length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFD39FD4186F0" box="[590,697,739,761]" gridcol="1" gridrow="19" pageId="4" pageNumber="3">5.2</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FD25FD41871C" box="[245,697,767,789]" gridrow="20" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FD25FE34871C" box="[245,460,767,789]" gridcol="0" gridrow="20" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE90FCDAFE57871C" bold="true" box="[372,431,768,789]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFD25FD41871C" box="[590,697,767,789]" gridcol="1" gridrow="20" pageId="4" pageNumber="3">1.2</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FCC1FD418739" box="[245,697,795,816]" gridrow="21" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FCC1FE348739" box="[245,460,795,816]" gridcol="0" gridrow="21" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFECFFCC1FE408739" bold="true" box="[299,440,795,816]" pageId="4" pageNumber="3">Patella length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFCC1FD418739" box="[590,697,795,816]" gridcol="1" gridrow="21" pageId="4" pageNumber="3">5.7</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FCECFD418745" box="[245,697,822,844]" gridrow="22" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FCECFE348745" box="[245,460,822,844]" gridcol="0" gridrow="22" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE9DFCEDFE4B8745" bold="true" box="[377,435,823,844]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFCECFD418745" box="[590,697,822,844]" gridcol="1" gridrow="22" pageId="4" pageNumber="3">1.7</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FC88FD41876E" box="[245,697,850,871]" gridrow="23" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FC88FE34876E" box="[245,460,850,871]" gridcol="0" gridrow="23" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFEC8FC88FE56876E" bold="true" box="[300,430,850,871]" pageId="4" pageNumber="3">Chela length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFC88FD41876E" box="[590,697,850,871]" gridcol="1" gridrow="23" pageId="4" pageNumber="3">8.9</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FCB4FD41878A" box="[245,697,878,899]" gridrow="24" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FCB4FE34878A" box="[245,460,878,899]" gridcol="0" gridrow="24" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFE89FCB4FE5F878A" bold="true" box="[365,423,878,899]" pageId="4" pageNumber="3">width</emphasis>
</td>
<td id="93128E06FFD30014FDAAFCB4FD41878A" box="[590,697,878,899]" gridcol="1" gridrow="24" pageId="4" pageNumber="3">1.5</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FC53FD418796" box="[245,697,905,927]" gridrow="25" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FC53FE348796" box="[245,460,905,927]" gridcol="0" gridrow="25" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFF12FC50FE428796" bold="true" box="[246,442,906,927]" pageId="4" pageNumber="3">Mov. Finger length</emphasis>
</td>
<td id="93128E06FFD30014FDAAFC53FD418796" box="[590,697,905,927]" gridcol="1" gridrow="25" pageId="4" pageNumber="3">5.8</td>
</tr>
<tr id="D0C3E77AFFD30014FF11FC71FD4187C9" box="[245,697,939,960]" gridrow="26" pageId="4" pageNumber="3">
<td id="93128E06FFD30014FF11FC71FE3487C9" box="[245,460,939,960]" gridcol="0" gridrow="26" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFF11FC71FE7987C9" bold="true" box="[245,385,939,960]" pageId="4" pageNumber="3">Pectinal teeth</emphasis>
</td>
<td id="93128E06FFD30014FDAAFC71FD4187C9" box="[590,697,939,960]" gridcol="1" gridrow="26" pageId="4" pageNumber="3">14:14</td>
</tr>
</table>
</paragraph>
<caption id="3A8CB5B0FFD3FFEFFF24FC3FFD09801D" pageId="4" pageNumber="3" startId="4.[192,257,997,1020]">
<paragraph id="6E4CE538FFD3FFEFFF24FC3FFD09801D" blockId="4.[192,802,997,1044]" pageId="4" pageNumber="3">
<emphasis id="5C87392AFFD3FFEFFF24FC3FFEDC87F5" bold="true" box="[192,292,997,1020]" pageId="4" pageNumber="3">Table 1:</emphasis>
Measurements (in mm) of the male holotype of
<taxonomicName id="A9F39EBBFFD3FFEFFF24FC25FEB6801D" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[192,334,1023,1044]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD3FFEFFF24FC25FEB6801D" box="[192,334,1023,1044]" italics="true" pageId="4" pageNumber="3">Tityus mraceki</emphasis>
</taxonomicName>
<emphasis id="5C87392AFFD3FFEFFEB0FC25FE72801D" bold="true" box="[340,394,1023,1044]" pageId="4" pageNumber="3">
<taxonomicNameLabel id="47B48451FFD3FFEFFEB0FC25FE72801D" box="[340,394,1023,1044]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
* Includes intersegment membranes.
</paragraph>
</caption>
<subSubSection id="26E9B6B3FFD3FFEFFF24FB72FC0E82BF" pageId="4" pageNumber="3" type="description">
<paragraph id="6E4CE538FFD3FFEFFF24FB72FE3E8218" blockId="4.[192,802,1192,1890]" pageId="4" pageNumber="3">
DESCRIPTION: Measurements of the carapace, telson, segments of the metasoma and segments of the pedipalps, and numbers of pectinal teeth are given in Table 2. Coloration is shown in
<figureCitation id="F6C8F9BDFFD3FFEFFDC9FADEFD478112" box="[557,703,1284,1307]" captionStart="Figures 1–3" captionStartId="3.[192,278,1900,1923]" captionTargetBox="[191,1440,191,1866]" captionTargetId="figure-8@3.[192,1439,192,1865]" captionTargetPageId="3" captionText="Figures 1–3: Tityus mraceki sp. n., male holotype. 1. Dorsal view. 2. Ventral view. 3. Lateral view of metasoma." figureDoi="http://doi.org/10.5281/zenodo.4673275" httpUri="https://zenodo.org/record/4673275/files/figure.png" pageId="4" pageNumber="3">Figs. 1 and 2</figureCitation>
. Most of the body is blotched, including the ventral surface of metasomal segments. Only the fifth metasomal segment and telson are black. The manus of pedipalp is yellow and spotted, fingers of pedipalps are brown. Although the female is not known, sexual dimorphism can be assumed because the male has elongated metasomal segments and, conversely, lacks widening of the pedipalp manus (
<figureCitation id="F6C8F9BDFFD3FFEFFE93FA20FE4E8218" box="[375,438,1530,1553]" captionStart="Figures 1–3" captionStartId="3.[192,278,1900,1923]" captionTargetBox="[191,1440,191,1866]" captionTargetId="figure-8@3.[192,1439,192,1865]" captionTargetPageId="3" captionText="Figures 1–3: Tityus mraceki sp. n., male holotype. 1. Dorsal view. 2. Ventral view. 3. Lateral view of metasoma." figureDoi="http://doi.org/10.5281/zenodo.4673275" httpUri="https://zenodo.org/record/4673275/files/figure.png" pageId="4" pageNumber="3">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="6E4CE538FFD3FFEFFF24F9C2FE81836B" blockId="4.[192,802,1192,1890]" pageId="4" pageNumber="3">MESOSOMA AND CARAPACE: The carapace bears only posterior median carinae but is densely granulate. The chelicerae are reticulate in anterior half, their fingers are dark. The mesosomal tergites I–VI with a single carina, which is indistinct due to dense granulation. The sternites are densely granulated and the seventh sternite bears four carinae. The third and fifth sternites bear glossy expanded zones in posterior parts. On the fourth sternite the glossy zone is reduced and on the sixth sternite it is absent. Stigmata are elongate. The pectinal tooth count is 14.</paragraph>
<paragraph id="6E4CE538FFD3FFEFFCDAFF1DFA8285BC" blockId="4.[830,1441,199,590]" pageId="4" pageNumber="3">
METASOMA AND TELSON: The first metasomal segment has 10 carinae, the second through fourth segments have eight carinae, and the fifth segment has only three ventral carinae. All the carinae run in parallel and are composed of equally sized granules (
<figureCitation id="F6C8F9BDFFD3FFEFFB04FE98FADD8550" box="[1248,1317,322,345]" captionStart="Figures 1–3" captionStartId="3.[192,278,1900,1923]" captionTargetBox="[191,1440,191,1866]" captionTargetId="figure-8@3.[192,1439,192,1865]" captionTargetPageId="3" captionText="Figures 1–3: Tityus mraceki sp. n., male holotype. 1. Dorsal view. 2. Ventral view. 3. Lateral view of metasoma." figureDoi="http://doi.org/10.5281/zenodo.4673275" httpUri="https://zenodo.org/record/4673275/files/figure.png" pageId="4" pageNumber="3">Fig. 3</figureCitation>
). The fifth segment is laterally convex and therefore wider than other segments. The telson has a strong rhomboidal subaculear tooth and bears four granules in two rows.
</paragraph>
<paragraph id="6E4CE538FFD3FFEFFCDAFE66FA848647" blockId="4.[830,1441,199,590]" pageId="4" pageNumber="3">PEDIPALPS: The movable fingers have 15 and 16 rows of granules, which include external and internal granules. The entire pedipalps are densely granulated, namely on the dorsal surface. The chela is narrow, granulated and carinated. The patella bears a strong internal tubercle.</paragraph>
<paragraph id="6E4CE538FFD3FFEFFCDAFDB4FC6C8121" blockId="4.[830,1441,622,1718]" pageId="4" pageNumber="3">
AFFINITIES. The described features distinguish
<taxonomicName id="A9F39EBBFFD3FFEFFABBFDB4FC6C86AD" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD3FFEFFABBFDB4FC6C86AD" italics="true" pageId="4" pageNumber="3">Tityus mraceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47B48451FFD3FFEFFC40FD57FC1E86AD" box="[932,998,653,676]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
from all other species of the genus. Within the subgenus
<taxonomicName id="A9F39EBBFFD3FFEFFBD4FD71FB3C86CB" authorityName="Lourenco" authorityYear="2006" box="[1072,1220,683,706]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="5C87392AFFD3FFEFFBD4FD71FB3C86CB" box="[1072,1220,683,706]" italics="true" pageId="4" pageNumber="3">Archaeotityus</emphasis>
</taxonomicName>
its assumed sexual dimorphism is unique, as the male long metasoma and the narrow chela of pedipalp distinguish
<taxonomicName id="A9F39EBBFFD3FFEFFAE7FD33FA808709" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[1283,1400,745,768]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD3FFEFFAE7FD33FA808709" box="[1283,1400,745,768]" italics="true" pageId="4" pageNumber="3">T. mraceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47B48451FFD3FFEFFA66FD33FCAB8717" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
from all other species. Other species of the subgenus
<taxonomicName id="A9F39EBBFFD3FFEFFCDAFCFCFC2A8734" authorityName="Lourenco" authorityYear="2006" box="[830,978,806,829]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="subGenus" subGenus="Archaeotityus">
<emphasis id="5C87392AFFD3FFEFFCDAFCFCFC2A8734" box="[830,978,806,829]" italics="true" pageId="4" pageNumber="3">Archaeotityus</emphasis>
</taxonomicName>
differ in sexual dimorphism, which may be divided into three groups: Male pedipalps broader and more bulky than those of females, male metasoma not markedly longer than that of female (e.g.
<taxonomicName id="A9F39EBBFFD3FFEFFABBFC58FB8D87B1" authority="C. L. Koch, 1843" authorityName="C. L. Koch" authorityYear="1843" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="clathratus">
<emphasis id="5C87392AFFD3FFEFFABBFC58FC5287B1" italics="true" pageId="4" pageNumber="3">Tityus clathratus</emphasis>
C. L. Koch, 1843
</taxonomicName>
); male pedipalps broad and more bulky than those of females, male metasoma longer than that of female (e.g.
<taxonomicName id="A9F39EBBFFD3FFEFFB7BFC04FC8F801D" authority="Abalos, 1954" authorityName="Abalos" authorityYear="1954" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="birabeni">
<emphasis id="5C87392AFFD3FFEFFB7BFC04FABC87FC" box="[1183,1348,990,1013]" italics="true" pageId="4" pageNumber="3">Tityus birabeni</emphasis>
Abalos, 1954
</taxonomicName>
); shape of pedipalp manus and length of metasomal segments similar in both sexes (e.g.
<taxonomicName id="A9F39EBBFFD3FFEFFABBFBC1FBB98058" authority="Kovarik, 2005" authorityName="Kovarik" authorityYear="2005" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="kaderkai">
<emphasis id="5C87392AFFD3FFEFFABBFBC1FC648058" italics="true" pageId="4" pageNumber="3">Tityus kaderkai</emphasis>
Kovařík, 2005
</taxonomicName>
). The adult male is not known in two species of this subgenus.
<taxonomicName id="A9F39EBBFFD3FFEFFB6DFB83FA598079" authority="Lourenco, 1999" authorityName="Lourenco" authorityYear="1999" box="[1161,1441,1113,1136]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="5C87392AFFD3FFEFFB6DFB83FB138079" box="[1161,1259,1113,1136]" italics="true" pageId="4" pageNumber="3">T. erikae</emphasis>
Lourenço, 1999
</taxonomicName>
is known only from a single juvenile specimen, which shares with
<taxonomicName id="A9F39EBBFFD3FFEFFC27FB4CFBCF80A4" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[963,1079,1174,1197]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD3FFEFFC27FB4CFBCF80A4" box="[963,1079,1174,1197]" italics="true" pageId="4" pageNumber="3">T. mraceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47B48451FFD3FFEFFBA4FB4CFB8480A4" box="[1088,1148,1174,1197]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
glossy expanded zone on the posterior part of the fifth sternite (fig.
<quantity id="A90B48DDFFD3FFEFFAE6FB6FFAD480C5" box="[1282,1324,1205,1228]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="3" unit="in" value="6.0">6 in</quantity>
Lourenço, 1999: 3); however,
<taxonomicName id="A9F39EBBFFD3FFEFFBFBFB09FB6E80E3" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[1055,1174,1235,1258]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD3FFEFFBFBFB09FB6E80E3" box="[1055,1174,1235,1258]" italics="true" pageId="4" pageNumber="3">T. mraceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47B48451FFD3FFEFFB46FB09FB1880E3" box="[1186,1248,1235,1258]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
has the sternites densely granulated, whereas
<taxonomicName id="A9F39EBBFFD3FFEFFB64FB28FB1B8100" authorityName="Lourenco" authorityYear="1999" box="[1152,1251,1266,1289]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="erikae">
<emphasis id="5C87392AFFD3FFEFFB64FB28FB1B8100" box="[1152,1251,1266,1289]" italics="true" pageId="4" pageNumber="3">T. erikae</emphasis>
</taxonomicName>
has them nearly smooth.
</paragraph>
<paragraph id="6E4CE538FFD3FFEFFC8AFAF5FC0E82BF" blockId="4.[830,1441,622,1718]" pageId="4" pageNumber="3">
<taxonomicName id="A9F39EBBFFD3FFEFFC8AFAF5FB64814F" authority="Lourenco, 1992" authorityName="Lourenco" authorityYear="1992" box="[878,1180,1327,1350]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="betschi">
<emphasis id="5C87392AFFD3FFEFFC8AFAF5FC26814F" box="[878,990,1327,1350]" italics="true" pageId="4" pageNumber="3">T. betschi</emphasis>
Lourenço, 1992
</taxonomicName>
is based on a female characterized as having a moderate and feebly rhomboidal subaculear tooth, which according to
<bibRefCitation id="0A6298C9FFD3FFEFFCDAFA51FBA981AB" author="LOURENCO" box="[830,1105,1419,1442]" pageId="4" pageNumber="3" pagination="449 - 461" refId="ref3007" refString="LOURENCO, W. R. 2000. Synopsis of the Colombian species of Tityus Koch (Chelicerata, Scorpiones, Buthidae), with descriptions of three new species. Journal of Natural History, 34: 449 - 461." type="journal article" year="2000">Lourenço (2000: 458)</bibRefCitation>
distinguishes it from
<taxonomicName id="A9F39EBBFFD3FFEFFA6FFA51FC3E81C8" authorityName="Thorell" authorityYear="1876" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="columbianus">
<emphasis id="5C87392AFFD3FFEFFA6FFA51FC3E81C8" italics="true" pageId="4" pageNumber="3">T. columbianus</emphasis>
</taxonomicName>
. However, the relatively common
<taxonomicName id="A9F39EBBFFD3FFEFFA6EFA70FC3E81E9" authorityName="Thorell" authorityYear="1876" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="columbianus">
<emphasis id="5C87392AFFD3FFEFFA6EFA70FC3E81E9" italics="true" pageId="4" pageNumber="3">T. columbianus</emphasis>
</taxonomicName>
also has a moderate and feebly rhomboidal subaculear tooth, thus the possibility cannot be excluded that
<taxonomicName id="A9F39EBBFFD3FFEFFC9EF9DCFC178214" authorityName="Lourenco" authorityYear="1992" box="[890,1007,1542,1565]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="betschi">
<emphasis id="5C87392AFFD3FFEFFC9EF9DCFC178214" box="[890,1007,1542,1565]" italics="true" pageId="4" pageNumber="3">T. betschi</emphasis>
</taxonomicName>
is a synonym of
<taxonomicName id="A9F39EBBFFD3FFEFFB0CF9DCFA618214" authorityName="Thorell" authorityYear="1876" box="[1256,1433,1542,1565]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="columbianus">
<emphasis id="5C87392AFFD3FFEFFB0CF9DCFA618214" box="[1256,1433,1542,1565]" italics="true" pageId="4" pageNumber="3">T. columbianus</emphasis>
</taxonomicName>
. Confusion between
<taxonomicName id="A9F39EBBFFD3FFEFFBF0F9FFFB418235" authorityName="Thorell" authorityYear="1876" box="[1044,1209,1573,1596]" class="Insecta" family="Hepialidae" genus="Phassus" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="columbianus">
<emphasis id="5C87392AFFD3FFEFFBF0F9FFFB418235" box="[1044,1209,1573,1596]" italics="true" pageId="4" pageNumber="3">T. columbianus</emphasis>
</taxonomicName>
and
<taxonomicName id="A9F39EBBFFD3FFEFFB0AF9FFFA988235" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[1262,1376,1573,1596]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD3FFEFFB0AF9FFFA988235" box="[1262,1376,1573,1596]" italics="true" pageId="4" pageNumber="3">T. mraceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47B48451FFD3FFEFFA83F9FFFA588235" box="[1383,1440,1573,1596]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
is not possible, the sexual dimorphism and proportions make these two species easy to separate. In addition, the telson of
<taxonomicName id="A9F39EBBFFD3FFEFFC55F95BFBD38291" authority="Kovařík, 2007" authorityName="Kovařík" authorityYear="2007" box="[945,1067,1665,1688]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mraceki" status="sp. nov.">
<emphasis id="5C87392AFFD3FFEFFC55F95BFBD38291" box="[945,1067,1665,1688]" italics="true" pageId="4" pageNumber="3">T. mraceki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="47B48451FFD3FFEFFBDDF95BFB828291" box="[1081,1146,1665,1688]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
has a strong rhomboidal subaculear tooth.
</paragraph>
</subSubSection>
</treatment>
</document>
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