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<document id="5A68634834D4C8857BD13939A0A3CD05" ID-CLB-Dataset="29802" ID-DOI="10.11646/zootaxa.4442.1.3" ID-GBIF-Dataset="51315b1a-dcc4-41e2-aff7-f90478fed976" ID-ISSN="1175-5326" ID-Zenodo-Dep="1301390" ID-ZooBank="973323F0-BA57-4ACE-9DB6-0C0237B2E487" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530293601868" checkinUser="plazi" docAuthor="Attasopa, Korrawat, Bänziger, Hans, Disayathanoowat, Terd &amp; Packer, Laurence" docDate="2018" docId="A71E87EE8B7FFFD7DDF8C6E0FE8DFC7E" docLanguage="en" docName="zootaxa.4442.1.3.pdf" docOrigin="Zootaxa 4442 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidotrigona flavibasis Attasopa, Bänziger, Disayathanoowat &amp; Packer, 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="79" masterDocId="5B27FF968B71FFC7DD6FC678FF8BFFBA" masterDocTitle="A new species of Lepidotrigona (Hymenoptera: Apidae) from Thailand with the description of males of L. flavibasis and L. doipaensis and comments on asymmetrical genitalia in bees" masterLastPageNumber="82" masterPageNumber="63" pageNumber="77" updateTime="1698551722399" updateUser="ExternalLinkService">
<mods:mods id="A39C2BCD619CC069FEE1E2D02EA375AA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="2CE38AEB85EAD65C3C58D5F4CD5208C5">A new species of Lepidotrigona (Hymenoptera: Apidae) from Thailand with the description of males of L. flavibasis and L. doipaensis and comments on asymmetrical genitalia in bees</mods:title>
</mods:titleInfo>
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<mods:namePart id="3EC75BD81CE2FC3F8E8D97B017A25FF2">Attasopa, Korrawat</mods:namePart>
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<mods:namePart id="950527163790B00C6D15420F8C75689D">Bänziger, Hans</mods:namePart>
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<mods:namePart id="238A9B9659AF20F4580753B6B4CFBB23">Disayathanoowat, Terd</mods:namePart>
</mods:name>
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<mods:namePart id="3AA09BD7F865A673B9D4480541536AA5">Packer, Laurence</mods:namePart>
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<mods:typeOfResource id="E4B2BC94E2B2E87F723497E76C10B254">text</mods:typeOfResource>
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<mods:title id="F8955F45BD07458489922C9A7EA36B5C">Zootaxa</mods:title>
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<mods:part id="5FD1FAC1D6464928AA34542679AB4EA4">
<mods:date id="8CA40BF44A42DB7326ED7288BFB62D1C">2018</mods:date>
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<mods:number id="D866ECA0ACCBB6D1DA431FD3FEB289E5">2018-06-29</mods:number>
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<mods:detail id="168DC59CCBE8A5C996EF825C3389D565" type="volume">
<mods:number id="3EFEB5D461DC3A951FD524806546771B">4442</mods:number>
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<mods:classification id="DBF8D9F61F692162ACF5556ED4B04FE4">journal article</mods:classification>
<mods:identifier id="57E9137F73E34005281E0661A30D5E4E" type="CLB-Dataset">29802</mods:identifier>
<mods:identifier id="801313150841C45E0CCC5AE08CA7E3EF" type="DOI">10.11646/zootaxa.4442.1.3</mods:identifier>
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<treatment id="A71E87EE8B7FFFD7DDF8C6E0FE8DFC7E" ID-DOI="http://doi.org/10.5281/zenodo.6489333" ID-Zenodo-Dep="6489333" LSID="urn:lsid:plazi:treatment:A71E87EE8B7FFFD7DDF8C6E0FE8DFC7E" httpUri="http://treatment.plazi.org/id/A71E87EE8B7FFFD7DDF8C6E0FE8DFC7E" lastPageId="16" lastPageNumber="79" pageId="14" pageNumber="77">
<subSubSection id="67AD65738B7FFFC9DDF8C6E0FE04FF0B" box="[151,399,151,178]" pageId="14" pageNumber="77" type="description">
<paragraph id="2F0836F88B7FFFC9DDF8C6E0FE04FF0B" blockId="14.[151,399,151,178]" box="[151,399,151,178]" pageId="14" pageNumber="77">
<heading id="744081948B7FFFC9DDF8C6E0FE04FF0B" bold="true" box="[151,399,151,178]" fontSize="11" level="1" pageId="14" pageNumber="77" reason="1">
<emphasis id="1DC3EAEA8B7FFFC9DDF8C6E0FE04FF0B" bold="true" box="[151,399,151,178]" pageId="14" pageNumber="77">
Male of
<taxonomicName id="E8B74D7B8B7FFFC9DD92C6EFFE04FF0B" ID-CoL="6PBX7" box="[253,399,151,177]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B7FFFC9DD92C6EFFE04FF0B" bold="true" box="[253,399,151,177]" italics="true" pageId="14" pageNumber="77">L. flavibasis</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="67AD65738B7FFFC9DDF8C6A6FE43FEA3" pageId="14" pageNumber="77" type="reference_group">
<paragraph id="2F0836F88B7FFFC9DDF8C6A6FBFFFF4D" blockId="14.[151,1140,221,282]" box="[151,1140,221,248]" pageId="14" pageNumber="77">
<taxonomicName id="E8B74D7B8B7FFFC9DDF8C6A6FD1CFF42" ID-CoL="6PBX7" authority="(Cockerell, 1929)" authorityName="Cockerell" authorityYear="1929" baseAuthorityName="Cockerell" baseAuthorityYear="1929" box="[151,663,222,248]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B7FFFC9DDF8C6A6FD1CFF42" bold="true" box="[151,663,222,248]" pageId="14" pageNumber="77">
<emphasis id="1DC3EAEA8B7FFFC9DDF8C6A6FE30FF42" bold="true" box="[151,443,222,248]" italics="true" pageId="14" pageNumber="77">Lepidotrigona flavibasis</emphasis>
(
<bibRefCitation id="4B264B098B7FFFC9DCA5C6A6FD04FF42" box="[458,655,222,248]" pageId="14" pageNumber="77" refId="ref12237" refString="Cockerell, T. D. A. (1929) Descriptions and records of bees. - CXX. Annals and Magazine of Natural History, Series 10, 4, 584 - 594." type="journal article">Cockerell, 1929</bibRefCitation>
)
</emphasis>
</taxonomicName>
, measured as range (Average±SD, n=4).
</paragraph>
<paragraph id="2F0836F88B7FFFC9DDF8C779FE43FEA3" blockId="14.[151,1140,221,282]" box="[151,456,257,282]" pageId="14" pageNumber="77">
<figureCitation id="B78C2A7D8B7FFFC9DDF8C779FF6DFEA3" box="[151,230,257,282]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="14" pageNumber="77">Figs. 2</figureCitation>
: B1–4, 3: B1–4 &amp; 8
</paragraph>
</subSubSection>
<subSubSection id="67AD65738B7FFFC9DDF8C731FB96FD7D" pageId="14" pageNumber="77" type="diagnosis">
<paragraph id="2F0836F88B7FFFC9DDF8C731FB96FD7D" blockId="14.[151,1437,329,2036]" pageId="14" pageNumber="77">
<emphasis id="1DC3EAEA8B7FFFC9DDF8C731FE9AFED8" bold="true" box="[151,273,329,354]" pageId="14" pageNumber="77">Diagnosis.</emphasis>
<taxonomicName id="E8B74D7B8B7FFFC9DC77C731FDA0FED8" box="[280,555,329,354]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B7FFFC9DC77C731FDA0FED8" box="[280,555,329,354]" italics="true" pageId="14" pageNumber="77">Lepidotrigona flavibasis</emphasis>
</taxonomicName>
differs from the other two species as follows: S4 apicolaterally bisinuate with medial convexity (
<figureCitation id="B78C2A7D8B7FFFC9DC08C714FE27FE3F" box="[359,428,364,389]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="14" pageNumber="77">Fig. 2</figureCitation>
: B1) (angularly emarginate in
<taxonomicName id="E8B74D7B8B7FFFC9DE6FC716FCD2FE3F" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[768,857,366,389]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B7FFFC9DE6FC716FCD2FE3F" box="[768,857,366,389]" italics="true" pageId="14" pageNumber="77">L. satun</emphasis>
</taxonomicName>
, weakly concave in
<taxonomicName id="E8B74D7B8B7FFFC9D952C716FB59FE3F" baseAuthorityName="Schwarz" baseAuthorityYear="1939" box="[1085,1234,364,389]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B7FFFC9D952C716FB59FE3F" box="[1085,1234,364,389]" italics="true" pageId="14" pageNumber="77">L. doipaensis</emphasis>
</taxonomicName>
(
<figureCitation id="B78C2A7D8B7FFFC9D98EC714FAADFE3F" box="[1249,1318,364,389]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="14" pageNumber="77">Fig. 2</figureCitation>
: A1, C1)); S5 with apicosubmedial lobes pointed apically (as in
<taxonomicName id="E8B74D7B8B7FFFC9DF88C7EAFCF7FE13" baseAuthorityName="Schwarz" baseAuthorityYear="1939" box="[743,892,400,425]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B7FFFC9DF88C7EAFCF7FE13" box="[743,892,400,425]" italics="true" pageId="14" pageNumber="77">L. doipaensis</emphasis>
</taxonomicName>
, rounded in
<taxonomicName id="E8B74D7B8B7FFFC9D968C7EAFBEAFE13" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[1031,1121,402,425]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B7FFFC9D968C7EAFBEAFE13" box="[1031,1121,402,425]" italics="true" pageId="14" pageNumber="77">L. satun</emphasis>
</taxonomicName>
), with one or two very short setae (generally only one very long seta and a much smaller seta in
<taxonomicName id="E8B74D7B8B7FFFC9DEC3C7CEFBCFFE77" baseAuthorityName="Schwarz" baseAuthorityYear="1939" box="[940,1092,436,461]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B7FFFC9DEC3C7CEFBCFFE77" box="[940,1092,436,461]" italics="true" pageId="14" pageNumber="77">L. doipaensis</emphasis>
</taxonomicName>
but 4–7 setae of intermediate length in
<taxonomicName id="E8B74D7B8B7FFFC9DC63C7A1FEE6FE4A" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[268,365,473,496]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B7FFFC9DC63C7A1FEE6FE4A" box="[268,365,473,496]" italics="true" pageId="14" pageNumber="77">L. satun</emphasis>
</taxonomicName>
); and S5 gradulus mostly diagonal except short transverse section touching the antecosta medially (
<figureCitation id="B78C2A7D8B7FFFC9DC65C784FEDAFDAF" box="[266,337,508,533]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="14" pageNumber="77">Fig. 2</figureCitation>
: B2) (gradulus parallel to and in touch with most of antecosta medially in
<taxonomicName id="E8B74D7B8B7FFFC9D9C3C785FACFFDAF" baseAuthorityName="Schwarz" baseAuthorityYear="1939" box="[1196,1348,508,533]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B7FFFC9D9C3C785FACFFDAF" box="[1196,1348,508,533]" italics="true" pageId="14" pageNumber="77">L. doipaensis</emphasis>
</taxonomicName>
, but no contact in
<taxonomicName id="E8B74D7B8B7FFFC9DC7EC459FEE5FD82" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[273,366,545,568]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B7FFFC9DC7EC459FEE5FD82" box="[273,366,545,568]" italics="true" pageId="14" pageNumber="77">L. satun</emphasis>
</taxonomicName>
(
<figureCitation id="B78C2A7D8B7FFFC9DCEFC467FE43FD82" box="[384,456,543,568]" captionStart="FIGURE 2" captionStartId="4.[151,250,1606,1628]" captionTargetBox="[160,1427,354,1563]" captionTargetId="figure@4.[151,1436,337,1584]" captionTargetPageId="4" captionText="FIGURE 2. Male metasomal sterna, ventral aspect labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.- GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to S4, S5, S6, and S7 respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the characters treated in the diagnoses." httpUri="https://zenodo.org/record/1301394/files/figure.png" pageId="14" pageNumber="77">Fig. 2</figureCitation>
: A2, C2)).
<taxonomicName id="E8B74D7B8B7FFFC9DF22C459FD52FD82" box="[589,729,543,568]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B7FFFC9DF22C459FD52FD82" box="[589,729,543,568]" italics="true" pageId="14" pageNumber="77">L. flavibasis</emphasis>
</taxonomicName>
also differs in having only greyish yellow hairs on metatibia dorsally (
<figureCitation id="B78C2A7D8B7FFFC9DC6DC43BFEC2FDE6" box="[258,329,579,604]" captionStart="FIGURE 8" captionStartId="15.[151,250,1971,1993]" captionTargetBox="[333,1260,195,1951]" captionTargetId="figure@15.[333,1260,195,1951]" captionTargetPageId="15" captionText="FIGURE 8. Male of L. flavibasis from the type locality (Doi Suthep, Chang Khian, Chiang Mai, Thailand). (A) dorsal habitus, (B) lateral habitus, and (C) head, frontal view." httpUri="https://zenodo.org/record/1301408/files/figure.png" pageId="14" pageNumber="77">Fig. 8</figureCitation>
: B) (dark brown intermixed with greyish yellow hairs in
<taxonomicName id="E8B74D7B8B7FFFC9DEB4C43DFBF9FDE6" baseAuthorityName="Schwarz" baseAuthorityYear="1939" box="[987,1138,579,604]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="doipaensis">
<emphasis id="1DC3EAEA8B7FFFC9DEB4C43DFBF9FDE6" box="[987,1138,579,604]" italics="true" pageId="14" pageNumber="77">L. doipaensis</emphasis>
</taxonomicName>
, but variably brown, dark brown or greyish yellow hairs in
<taxonomicName id="E8B74D7B8B7FFFC9DF64C410FDEEFDC5" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[523,613,616,639]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B7FFFC9DF64C410FDEEFDC5" box="[523,613,616,639]" italics="true" pageId="14" pageNumber="77">L. satun</emphasis>
</taxonomicName>
). The ratio of the forewing diagonal to the length of the 2nd submarginal cell separates
<taxonomicName id="E8B74D7B8B7FFFC9DC57C4F4FE49FD1E" box="[312,450,651,676]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B7FFFC9DC57C4F4FE49FD1E" box="[312,450,651,676]" italics="true" pageId="14" pageNumber="77">L. flavibasis</emphasis>
</taxonomicName>
from
<taxonomicName id="E8B74D7B8B7FFFC9DF64C4F4FDE3FD19" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[523,616,652,675]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B7FFFC9DF64C4F4FDE3FD19" box="[523,616,652,675]" italics="true" pageId="14" pageNumber="77">L. satun</emphasis>
</taxonomicName>
(see diagnosis of
<taxonomicName id="E8B74D7B8B7FFFC9DE56C4F4FC1DFD19" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[825,918,652,675]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="satun">
<emphasis id="1DC3EAEA8B7FFFC9DE56C4F4FC1DFD19" box="[825,918,652,675]" italics="true" pageId="14" pageNumber="77">L. satun</emphasis>
</taxonomicName>
above and
<tableCitation id="623503438B7FFFC9D973C4F3FBE4FD1E" box="[1052,1135,651,676]" captionStart="TABLE 1" captionStartId="16.[151,239,1012,1035]" captionText="TABLE 1. Measurement of malar space length (MSL), F1 length (F1L), metatibia width (MTW), forewing diagonal (FWD), and 2nd submarginal cell (SM2L) ratios that separate L. satun n. sp., L. flavibasis, and L. doipaensis, expressed as range (Average±SD). Holotype ratios in bold." pageId="14" pageNumber="77">Table 1</tableCitation>
). The gonostyli and penis valves are also clearly different in the three species (see descriptions and
<figureCitation id="B78C2A7D8B7FFFC9DEA4C4D7FB85FD7D" box="[971,1038,687,712]" captionStart="FIGURE 3" captionStartId="6.[151,250,1918,1940]" captionTargetBox="[229,1364,221,1877]" captionTargetId="figure@6.[201,1389,193,1903]" captionTargetPageId="6" captionText="FIGURE 3. Male genital capsules, labelled as: (A), (B), and (C) refer to L. satun n. sp. (Paratype, H.B.-GP3311), L. flavibasis (H.B.-GP3322), and L. doipaensis (H.B.-GP3183) respectively, with the number following as (1), (2), (3), and (4) refer to dorsal, ventral, lateral, and caudal aspect respectively. All subfigures are shown at the same scale and magnification. Diagonal lines indicate the right penis valve." httpUri="https://zenodo.org/record/1301396/files/figure.png" pageId="14" pageNumber="77">Fig. 3</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="67AD65738B7FFFD7DDA8C4ABFE80FE10" lastPageId="16" lastPageNumber="79" pageId="14" pageNumber="77" type="description">
<paragraph id="2F0836F88B7FFFC9DDA8C4ABFC44FD56" blockId="14.[151,1437,329,2036]" box="[199,975,723,748]" pageId="14" pageNumber="77">
<emphasis id="1DC3EAEA8B7FFFC9DDA8C4ABFED2FD56" bold="true" box="[199,345,723,748]" pageId="14" pageNumber="77">Description.</emphasis>
As for the
<typeStatus id="F00C885A8B7FFFC9DCB9C4ABFDB3FD56" box="[470,568,723,748]" pageId="14" pageNumber="77" type="holotype">holotype</typeStatus>
of
<taxonomicName id="E8B74D7B8B7FFFC9DF32C4ACFD33FD51" authority="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer, 2018" authorityName="Attasopa &amp; Bänziger &amp; Disayathanoowat &amp; Packer" authorityYear="2018" box="[605,696,724,747]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="77" phylum="Arthropoda" rank="species" species="satun" status="sp. nov.">
<emphasis id="1DC3EAEA8B7FFFC9DF32C4ACFD33FD51" box="[605,696,724,747]" italics="true" pageId="14" pageNumber="77">L. satun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="06F057918B7FFFC9DFD0C4ABFD76FD56" box="[703,765,723,748]" pageId="14" pageNumber="77" rank="species">
<emphasis id="1DC3EAEA8B7FFFC9DFD0C4ABFD76FD56" bold="true" box="[703,765,723,748]" pageId="14" pageNumber="77">n. sp.</emphasis>
</taxonomicNameLabel>
except as follows:
</paragraph>
<paragraph id="2F0836F88B7FFFC9DDA8C48FFF64F9FC" blockId="14.[151,1437,329,2036]" pageId="14" pageNumber="77">
<emphasis id="1DC3EAEA8B7FFFC9DDA8C48FFEBDFCB4" box="[199,310,759,782]" italics="true" pageId="14" pageNumber="77">Structure:</emphasis>
Body length 4.8–4.94 (4.91±0.07) mm. Head width 1.76–1.78 (1.78±0.01) mm; head length 1.34– 1.37 (1.35±0.01) mm; eye width 0.56–0.57 (0.56±0.005) mm; gena width 0.18–0.2 (0.19±0.01) mm; interocellar distance 0.34–0.35 (0.35±0.01) mm; ocellocular distance 0.2–0.22 (0.21±0.01) mm; upper interorbital distance 1.1–1.11 (1.1±0.01) mm, lower interorbital distance 0.75–0.78 (0.77±0.01) mm; scape length excluding basal bulb 0.51–0.52 (0.51±0.005) mm, with basal bulb 0.57–0.59 (0.58±0.01) mm; 1
<taxonomicName id="E8B74D7B8B7FFFC9D977C5FEFB41FC24" box="[1048,1226,901,926]" pageId="14" pageNumber="77" rank="subSpecies" subSpecies="flagellomere">st flagellomere</taxonomicName>
length 0.08–0.13 (0.11±0.02) mm; 2nd flagellomere length 0.17–0.18 (0.18±0.005) mm, width 0.14–0.15 (0.14±0.004) mm; 3rd slightly shorter than 2nd; 4th longer than broad; malar space length 0.03–0.04 (0.03±0.005) mm. Forewing length 4.39–4.48 (4.42±0.04) mm, width 1.6–1.61 (1.6±0.01) mm; forewing diagonal 1.23–1.29 (1.26±0.03) mm, length of 1
<taxonomicName id="E8B74D7B8B7FFFC9DDD0C26DFEEDFB94" box="[191,358,1045,1070]" pageId="14" pageNumber="77" rank="subSpecies" subSpecies="submarginal">st submarginal</taxonomicName>
cell 0.3–0.33 (0.31±0.01) mm, 2nd submarginal cell length 0.58–0.61 (0.6±0.01) mm. Metatibia length 1.4–1.42 (1.41±0.01) mm, width 0.54–0.56 (0.55±0.01) mm; metabasitarsus length 0.58–0.59 (0.59±0.01) mm, width 0.37–0.39 (0.38±0.01) mm. S4 convex apicomedially, gradulus of S4 weakly concave medially, with small apicolateral angulation; S5 apical margin with two long, narrow processes bearing one or two very short thick setae, processes delimit an emargination ~0.54 times as deep as length of sternum and as long as wide, apicolateral lobe well developed, bearing long curved setae, gradulus touching antecosta medially; S6 with acute triangular apicomedial process; S7 approximate twice as wide as long, apical margin weakly emarginate. Genital capsule dark brown to black except base of penis valve, apical half to 2/3 of gonostylus and gonocoxa anteriorly pale brown to brown; the right penis valve overarches the left one and has two right angles directed to the right, so that the tip of the valve ends to the right of the midline of the bee, and generally posterior and often dorsal to the tip of the left penis valve. In the left penis valve the first angle is less acute and the second angle is missing but replaced by a curvature directed to the right, hence also bringing the tip of the left valve to the right of the midline, the tip is anterior and often ventral to the tip of the right penis valve. In addition, the final fourth or fifth of the valves is sinuous, the tips diverging from each other. Gonostylus sclerotized basally, approximately 12 times longer than wide (length
<quantity id="E84F9B1D8B7FFFC9DC06C072FD9BF998" box="[361,528,1546,1571]" metricMagnitude="-4" metricUnit="m" metricValue="9.35" metricValueMax="9.6" metricValueMin="9.1" pageId="14" pageNumber="77" unit="mm" value="0.935" valueMax="0.96" valueMin="0.91">0.91–0.96 mm</quantity>
, width
<quantity id="E84F9B1D8B7FFFC9DF08C072FCA2F998" box="[615,809,1546,1570]" metricMagnitude="-5" metricUnit="m" metricValue="7.65" metricValueMax="8.0" metricValueMin="7.3" pageId="14" pageNumber="77" unit="mm" value="0.0765" valueMax="0.08" valueMin="0.073">0.073–0.080 mm</quantity>
) hairs equally dense throughout the apically expanded portion.
</paragraph>
<paragraph id="2F0836F88B7FFFC9DDA8C029FA1CF8FB" blockId="14.[151,1437,329,2036]" pageId="14" pageNumber="77">
<emphasis id="1DC3EAEA8B7FFFC9DDA8C029FE63F9D0" box="[199,488,1617,1642]" italics="true" pageId="14" pageNumber="77">Coloration of integument:</emphasis>
Head black except as follows: yellowish brown on basal bulb and extreme apex of scape, labrum, and apical half of mandible; F2–F11 dark brown. Mesosoma black except pronotal lobe and tegula dark brown; fore and mid legs dark brown except medio- and distitarsi yellowish brown; hind leg black except tarsus yellowish brown (sometimes metabasitarsus dark brown). Metasomal terga black except basal depression of T1 transparent brown and apex of T6 and T7 brown. S1–S3 dark brown except posterior half of S1, posterior margin of S2, and apex of S3 whitish translucent; S4 brown except apical margin and most of disc medially translucent; S5 with apical and apicolateral lobes brown; S6 with anterolateral brown spot; S7 translucent except gradulus narrowly brown.
</paragraph>
<paragraph id="2F0836F88B7FFFD7DDA8C134FC8DFE85" blockId="14.[151,1437,329,2036]" lastBlockId="16.[151,1437,151,964]" lastPageId="16" lastPageNumber="79" pageId="14" pageNumber="77">
<emphasis id="1DC3EAEA8B7FFFC9DDA8C134FEDFF8DF" box="[199,340,1868,1893]" italics="true" pageId="14" pageNumber="77">Pubescence:</emphasis>
Face with short plumose dense appressed white hairs; vertex with erect long pale whitish brown hairs. Mesoscutum with long and very short pale whitish brown hairs; mesoscutum, pronotal lobe and mesepisternum lacking distinct band of short plumose scale-like yellow hairs on margins; mesoscutellum with erect long pale whitish brown hairs longest on posterior margin; mesepisternum, metepisternum and metanotum with brownish white hairs; coxa and trochanter of all legs and ventral surface of all femora with long pale brownish white hairs, rest of femora with brownish white hairs intermixed with pale brown hairs; pro- and mesotibiae with dark brown hairs intermixed with brownish white hairs; metatibia with short greyish yellow hairs, coloration and density approximately the same on dorsal and ventral margins; all tarsi with whitish brown hairs intermixed with yellowish brown hairs, metatibia with short sparse brown keirotrichia. Posterolateral margin of T3–T7 with fine brownish white hairs progressively longer from T3–T7.
</paragraph>
<caption id="7BC866708B7EFFC8DDF8C1CBFDE1F852" httpUri="https://zenodo.org/record/1301408/files/figure.png" pageId="15" pageNumber="78" startId="15.[151,250,1971,1993]" targetBox="[333,1260,195,1951]" targetPageId="15">
<paragraph id="2F0836F88B7EFFC8DDF8C1CBFDE1F852" blockId="15.[151,1436,1971,2024]" pageId="15" pageNumber="78">
<emphasis id="1DC3EAEA8B7EFFC8DDF8C1CBFE99F872" bold="true" box="[151,274,1971,1993]" pageId="15" pageNumber="78">FIGURE 8.</emphasis>
Male of
<taxonomicName id="E8B74D7B8B7EFFC8DC01C1CCFE61F873" box="[366,490,1971,1993]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="flavibasis">
<emphasis id="1DC3EAEA8B7EFFC8DC01C1CCFE61F873" box="[366,490,1971,1993]" italics="true" pageId="15" pageNumber="78">L. flavibasis</emphasis>
</taxonomicName>
from the type locality (Doi Suthep, Chang Khian, Chiang Mai, Thailand). (A) dorsal habitus, (B) lateral habitus, and (C) head, frontal view.
</paragraph>
</caption>
<paragraph id="2F0836F88B61FFD7DDA8C733FE80FE10" blockId="16.[151,1437,151,964]" pageId="16" pageNumber="79">
<emphasis id="1DC3EAEA8B61FFD7DDA8C733FE18FED9" box="[199,403,330,355]" italics="true" pageId="16" pageNumber="79">Surface sculpture:</emphasis>
All legs with fine dense punctures. T1 with sparse fine shallow punctures except impunctate on basal depression; metasomal sterna with coarse shallow crowded punctures intermixed with sparse coarse punctures.
</paragraph>
</subSubSection>
<subSubSection id="67AD65738B61FFD7DDA8C7CEFE8DFC7E" pageId="16" pageNumber="79" type="materials_examined">
<paragraph id="2F0836F88B61FFD7DDA8C7CEFDDCFDAC" blockId="16.[151,1437,151,964]" pageId="16" pageNumber="79">
<emphasis id="1DC3EAEA8B61FFD7DDA8C7CEFE3DFE75" bold="true" box="[199,438,438,463]" pageId="16" pageNumber="79">Material examined.</emphasis>
Type material:
<emphasis id="1DC3EAEA8B61FFD7DF19C7CEFD56FE75" box="[630,733,438,463]" italics="true" pageId="16" pageNumber="79">Holotype</emphasis>
(worker, AMNH); Type locality:
<emphasis id="1DC3EAEA8B61FFD7D91BC7CEFA80FE75" bold="true" box="[1140,1291,438,463]" pageId="16" pageNumber="79">THAILAND</emphasis>
; labelled as follows: “Doi Sutep./Siam/
<date id="5B0910388B61FFD7DCB2C7A2FDACFE48" box="[477,551,474,499]" pageId="16" pageNumber="79">Feb. 9</date>
/ Alice Mackie”, “
<emphasis id="1DC3EAEA8B61FFD7DE6BC7A3FC57FE49" box="[772,988,474,499]" italics="true" pageId="16" pageNumber="79">
<taxonomicName id="E8B74D7B8B61FFD7DE6BC7A3FCD3FE48" box="[772,856,475,498]" class="Insecta" family="Apidae" genus="Trigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="genus">Trigona</taxonomicName>
/
<taxonomicName id="E8B74D7B8B61FFD7DE1EC7A2FC57FE49" box="[881,988,474,499]" class="Insecta" family="Apidae" genus="Lepidotrigona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="79" phylum="Arthropoda" rank="species" species="flavibasis">flavibasis</taxonomicName>
</emphasis>
/ TYPE. CKll”, “acc 35740”. [Images examined (provided by AMNH, 2017)].
</paragraph>
<paragraph id="2F0836F88B61FFD7DDA8C459FDD3FCC6" blockId="16.[151,1437,151,964]" pageId="16" pageNumber="79">
<materialsCitation id="9FDF3CA58B61FFD7DDA8C459FE91FD1C" collectingDate="2017-04-06" collectorName="H. Banziger &amp; Nest &amp; Measured &amp; Doi Suthep" country="Thailand" county="Measured" location="Doi Suthep" municipality="Chiang Mai Prov." pageId="16" pageNumber="79" specimenCount="4" specimenCount-male="4">
Additional material:
<emphasis id="1DC3EAEA8B61FFD7DCC0C459FA17FD83" box="[431,1436,545,570]" italics="true" pageId="16" pageNumber="79">
<collectingCounty id="C6694E748B61FFD7DCC0C459FD95FD80" box="[431,542,545,570]" pageId="16" pageNumber="79">Measured</collectingCounty>
male specimens included in description and
<tableCitation id="623503438B61FFD7D97CC459FBEEFD80" box="[1043,1125,545,570]" captionStart="TABLE 1" captionStartId="16.[151,239,1012,1035]" captionText="TABLE 1. Measurement of malar space length (MSL), F1 length (F1L), metatibia width (MTW), forewing diagonal (FWD), and 2nd submarginal cell (SM2L) ratios that separate L. satun n. sp., L. flavibasis, and L. doipaensis, expressed as range (Average±SD). Holotype ratios in bold." pageId="16" pageNumber="79">Table 1</tableCitation>
(collected from wild nests):
</emphasis>
<collectingCountry id="57A076688B61FFD7DDF8C43DFEA5FDE4" box="[151,302,581,606]" name="Thailand" pageId="16" pageNumber="79">
<emphasis id="1DC3EAEA8B61FFD7DDF8C43DFEA5FDE4" bold="true" box="[151,302,581,606]" pageId="16" pageNumber="79">THAILAND</emphasis>
</collectingCountry>
,
<collectingMunicipality id="CF6CAC828B61FFD7DC51C43DFD86FDE7" box="[318,525,581,606]" pageId="16" pageNumber="79">Chiang Mai Prov.</collectingMunicipality>
,
<location id="2A6860238B61FFD7DF75C43DFD14FDE4" LSID="urn:lsid:plazi:treatment:A71E87EE8B7FFFD7DDF8C6E0FE8DFC7E:2A6860238B61FFD7DF75C43DFD14FDE4" box="[538,671,581,606]" country="Thailand" county="Measured" municipality="Chiang Mai Prov." name="Doi Suthep" pageId="16" pageNumber="79">Doi Suthep</location>
:
<collectingDate id="4B4DE9D08B61FFD7DFC3C43DFC9CFDE4" box="[684,791,581,606]" pageId="16" pageNumber="79" value="2017-04-06">
<date id="5B0910388B61FFD7DFC3C43DFC9CFDE4" box="[684,791,581,606]" pageId="16" pageNumber="79" value="2017-04-06">6.iv.2017</date>
</collectingDate>
,
<collectorName id="8242532E8B61FFD7DE4AC43EFC24FDE4" box="[805,943,581,606]" pageId="16" pageNumber="79">H. Bänziger</collectorName>
(
<collectorName id="8242532E8B61FFD7DEAFC43EFC7EFDE4" box="[960,1013,582,606]" pageId="16" pageNumber="79">Nest</collectorName>
SHQ2,
<specimenCount id="39B1FD718B61FFD7D93AC43EFB3AFDE4" box="[1109,1201,581,606]" pageId="16" pageNumber="79" type="male">4 males</specimenCount>
, caught leaving nest [not swarming]).
<emphasis id="1DC3EAEA8B61FFD7DC32C411FB48FD3B" box="[349,1219,617,642]" italics="true" pageId="16" pageNumber="79">
<collectorName id="8242532E8B61FFD7DC32C411FE47FD38" box="[349,460,617,642]" pageId="16" pageNumber="79">Measured</collectorName>
worker specimens included in
<tableCitation id="623503438B61FFD7DE40C411FC08FD3B" box="[815,899,617,642]" captionStart="TABLE 1" captionStartId="16.[151,239,1012,1035]" captionText="TABLE 1. Measurement of malar space length (MSL), F1 length (F1L), metatibia width (MTW), forewing diagonal (FWD), and 2nd submarginal cell (SM2L) ratios that separate L. satun n. sp., L. flavibasis, and L. doipaensis, expressed as range (Average±SD). Holotype ratios in bold." pageId="16" pageNumber="79">Table 1</tableCitation>
(collected from wild nests):
</emphasis>
<location id="2A6860238B61FFD7D9A4C411FA12FD3B" LSID="urn:lsid:plazi:treatment:A71E87EE8B7FFFD7DDF8C6E0FE8DFC7E:2A6860238B61FFD7D9A4C411FA12FD3B" box="[1227,1433,617,642]" country="Thailand" county="Measured" municipality="Chiang Mai Prov." name="Chiang Mai Prov." pageId="16" pageNumber="79">Chiang Mai Prov.</location>
:
<collectorName id="8242532E8B61FFD7DDF8C4F5FE91FD1C" box="[151,282,653,678]" pageId="16" pageNumber="79">Doi Suthep</collectorName>
</materialsCitation>
,
<date id="5B0910388B61FFD7DC48C4F5FE22FD1C" box="[295,425,653,678]" pageId="16" pageNumber="79" value="2015-12-26">26.xii.2015</date>
, K. Attasopa (Nest PHPL1, 2 workers); Doi Suthep-Pui,
<quantity id="E84F9B1D8B61FFD7D959C4F5FB05FD1F" box="[1078,1166,653,677]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="16" pageNumber="79" unit="m" value="1080.0">1080 m</quantity>
,
<date id="5B0910388B61FFD7D9F3C4F5FA8DFD1C" box="[1180,1286,653,678]" pageId="16" pageNumber="79" value="2017-04-06">6.iv.2017</date>
, H. Bänziger (Nest SHQ2, 2 workers).
<materialsCitation id="9FDF3CA58B61FFD7DCD3C4C8FC25FD57" collectingDate="2016-01-17" collectionCode="MYAL, MKLL" collectorName="K. Attasopa &amp; Nest" country="Thailand" location="Doi Inthanon" municipality="Non-measured" pageId="16" pageNumber="79" specimenCode="MYAL1, MKLL3" specimenCount="6" specimenCount-worker="6" stateProvince="Chiang Mai">
<emphasis id="1DC3EAEA8B61FFD7DCD3C4C8FB97FD72" box="[444,1052,688,713]" italics="true" pageId="16" pageNumber="79">
<collectingMunicipality id="CF6CAC828B61FFD7DCD3C4C8FDEBFD73" box="[444,608,688,713]" pageId="16" pageNumber="79">Non-measured</collectingMunicipality>
specimens (collected from wild nests):
</emphasis>
<collectingRegion id="ED73F81A8B61FFD7D94BC4C8FB20FD73" box="[1060,1195,688,713]" country="Thailand" name="Chiang Mai" pageId="16" pageNumber="79">Chiang Mai</collectingRegion>
Prov.:
<location id="2A6860238B61FFD7D991C4C8FA13FD73" LSID="urn:lsid:plazi:treatment:A71E87EE8B7FFFD7DDF8C6E0FE8DFC7E:2A6860238B61FFD7D991C4C8FA13FD73" box="[1278,1432,688,713]" country="Thailand" municipality="Non-measured" name="Doi Inthanon" pageId="16" pageNumber="79" stateProvince="Chiang Mai">Doi Inthanon</location>
,
<collectingDate id="4B4DE9D08B61FFD7DDF8C4ACFE88FD57" box="[151,259,724,749]" pageId="16" pageNumber="79" value="2016-01-17">
<date id="5B0910388B61FFD7DDF8C4ACFE88FD57" box="[151,259,724,749]" pageId="16" pageNumber="79" value="2016-01-17">17.i.2016</date>
</collectingDate>
,
<collectorName id="8242532E8B61FFD7DC61C4ADFE1FFD57" box="[270,404,725,749]" pageId="16" pageNumber="79">K. Attasopa</collectorName>
(
<collectorName id="8242532E8B61FFD7DCCDC4ADFE5CFD57" box="[418,471,725,749]" pageId="16" pageNumber="79">Nest</collectorName>
<specimenCode id="7F119E838B61FFD7DCB1C4ADFDB4FD57" box="[478,575,725,749]" collectionCode="MYAL" pageId="16" pageNumber="79">MYAL1</specimenCode>
,
<specimenCount id="39B1FD718B61FFD7DF26C4ADFD37FD57" box="[585,700,724,749]" pageId="16" pageNumber="79" type="worker">3 workers</specimenCount>
;
<specimenCode id="7F119E838B61FFD7DFA8C4ADFCACFD57" box="[711,807,725,749]" collectionCode="MKLL" pageId="16" pageNumber="79">MKLL3</specimenCode>
,
<specimenCount id="39B1FD718B61FFD7DE5DC4ADFC2DFD57" box="[818,934,724,749]" pageId="16" pageNumber="79" type="worker">3 workers</specimenCount>
)
</materialsCitation>
;
<materialsCitation id="9FDF3CA58B61FFD7DED6C4ACFC45FCAA" collectingDate="2015-12-26" collectionCode="DSTL, PHPL" collectorName="K. Attasopa &amp; Nest &amp; Doi Suthep-Pui" country="Thailand" elevation="1080" location="Doi Suthep" pageId="16" pageNumber="79" specimenCode="DSTL1, PHPL1" specimenCount="5" specimenCount-worker="5" stateProvince="Chiang Mai">
<location id="2A6860238B61FFD7DED6C4ACFBB0FD57" LSID="urn:lsid:plazi:treatment:A71E87EE8B7FFFD7DDF8C6E0FE8DFC7E:2A6860238B61FFD7DED6C4ACFBB0FD57" box="[953,1083,724,749]" country="Thailand" name="Doi Suthep" pageId="16" pageNumber="79" stateProvince="Chiang Mai">Doi Suthep</location>
,
<collectingDate id="4B4DE9D08B61FFD7D929C4ACFB43FD57" box="[1094,1224,724,749]" pageId="16" pageNumber="79" value="2015-12-26">
<date id="5B0910388B61FFD7D929C4ACFB43FD57" box="[1094,1224,724,749]" pageId="16" pageNumber="79" value="2015-12-26">26.xii.2015</date>
</collectingDate>
,
<collectorName id="8242532E8B61FFD7D9BCC4ADFAD2FD57" box="[1235,1369,725,749]" pageId="16" pageNumber="79">K. Attasopa</collectorName>
(
<collectorName id="8242532E8B61FFD7D808C4ADFA17FD57" box="[1383,1436,725,749]" pageId="16" pageNumber="79">Nest</collectorName>
<specimenCode id="7F119E838B61FFD7DDF8C481FF66FCAB" box="[151,237,761,785]" collectionCode="DSTL" pageId="16" pageNumber="79">DSTL1</specimenCode>
,
<specimenCount id="39B1FD718B61FFD7DD92C481FEFFFCAB" box="[253,372,760,785]" pageId="16" pageNumber="79" type="worker">2 workers</specimenCount>
; nest
<specimenCode id="7F119E838B61FFD7DCD4C481FD9BFCAB" box="[443,528,761,785]" collectionCode="PHPL" pageId="16" pageNumber="79">PHPL1</specimenCode>
,
<specimenCount id="39B1FD718B61FFD7DF4FC481FD12FCAB" box="[544,665,760,785]" pageId="16" pageNumber="79" type="worker">3 workers</specimenCount>
);
<collectorName id="8242532E8B61FFD7DFC0C480FCE8FCAB" box="[687,867,760,785]" pageId="16" pageNumber="79">Doi Suthep-Pui</collectorName>
,
<elevation id="A49AD1CB8B61FFD7DE1CC481FC45FCAA" box="[883,974,761,785]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="16" pageNumber="79" unit="m" value="1080.0">
<quantity id="E84F9B1D8B61FFD7DE1CC481FC45FCAA" box="[883,974,761,785]" metricMagnitude="3" metricUnit="m" metricValue="1.08" pageId="16" pageNumber="79" unit="m" value="1080.0">1080 m</quantity>
</elevation>
</materialsCitation>
,
<date id="5B0910388B61FFD7DEB1C480FBC3FCAB" box="[990,1096,760,785]" pageId="16" pageNumber="79" value="2017-04-01">1.iv.2017</date>
, H. Bänziger (Nest SHQ2, 6 workers),
<date id="5B0910388B61FFD7DC64C564FEFDFC8F" box="[267,374,796,821]" pageId="16" pageNumber="79" value="2017-04-06">6.iv.2017</date>
, H. Bänziger (Nest SHQ2, 16 males, 9 workers, caught leaving nest [not swarming]).
<emphasis id="1DC3EAEA8B61FFD7D80BC565FC14FCED" italics="true" pageId="16" pageNumber="79">Nonmeasured specimens (collected by H.B. while they sucked his sweat):</emphasis>
Chiang Mai Prov., Doi Inthanon, above Mae Ya Noi,
<quantity id="E84F9B1D8B61FFD7DD9BC51CFEC0FCC1" box="[244,331,868,892]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="79" unit="m" value="1700.0">1700 m</quantity>
,
<date id="5B0910388B61FFD7DC39C51BFE43FCC6" box="[342,456,867,892]" pageId="16" pageNumber="79" value="1993-02-22">22.ii.1993</date>
(4 workers).
</paragraph>
<paragraph id="2F0836F88B61FFD7DDA8C5FFFE8DFC7E" blockId="16.[151,1437,151,964]" pageId="16" pageNumber="79">
Genitalia examined: GP 3296, 3297, 3298, 3299, 3300, 3301, 3302, nest SHQ2 of type locality, Doi Suthep,
<date id="5B0910388B61FFD7DDF8C5D3FE89FC7E" box="[151,258,939,964]" pageId="16" pageNumber="79" value="2017-04-06">6.iv.2017</date>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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