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<document id="00850A318BEDFFDFA7167F7EBD1AA53E" ID-DOI="10.5281/zenodo.195791" ID-GBIF-Dataset="c3cabf80-b53f-42f3-8580-43aac34bc1d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="195791" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460234708641" checkinUser="plazi" docAuthor="Naderloo, Reza, Türkay, Michael &amp; Chen, Hui-Lian" docDate="2010" docId="1C0430193229330C4BB565C3FBBBF80A" docLanguage="en" docName="zt02500p038.pdf" docOrigin="Zootaxa 2500" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Uca (Austruca) mjoebergi Rathbun 1924" docType="treatment" docVersion="12" lastPageNumber="22" masterDocId="E03D4861323C331A4B226454FFD1F928" masterDocTitle="Taxonomic revision of the wide-front fiddler crabs of the Uca lactea group (Crustacea: Decapoda: Brachyura: Ocypodidae) in the Indo-West Pacific" masterLastPageNumber="38" masterPageNumber="1" pageNumber="22" updateTime="1698239279183" updateUser="plazi">
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<mods:title id="59B1BB8E7033B6DBCA7F14B3E3A3F1FF">Taxonomic revision of the wide-front fiddler crabs of the Uca lactea group (Crustacea: Decapoda: Brachyura: Ocypodidae) in the Indo-West Pacific</mods:title>
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<mods:namePart id="E7A76C1AB84FBB78C9BAD99F467EBF33">Naderloo, Reza</mods:namePart>
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<mods:namePart id="9330AD18D99BE6B919AED76BDC96E4C2">Türkay, Michael</mods:namePart>
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<mods:namePart id="572907FC6411EFB1FD181106F68166D2">Chen, Hui-Lian</mods:namePart>
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<mods:date id="7FB975EC5855F11275FC4A0E83ECF17D">2010</mods:date>
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<treatment id="1C0430193229330C4BB565C3FBBBF80A" ID-DOI="http://doi.org/10.5281/zenodo.6209449" ID-GBIF-Taxon="119387813" ID-Zenodo-Dep="6209449" LSID="urn:lsid:plazi:treatment:1C0430193229330C4BB565C3FBBBF80A" httpUri="http://treatment.plazi.org/id/1C0430193229330C4BB565C3FBBBF80A" lastPageId="22" lastPageNumber="22" pageId="21" pageNumber="22">
<subSubSection id="DCB7D2843229330F4BB565C3FE24F8FC" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="9412810F3229330F4BB565C3FD41F899" blockId="21.[151,656,407,468]" box="[151,656,407,433]" pageId="21" pageNumber="22">
<heading id="CF5A36633229330F4BB565C3FD41F899" bold="true" box="[151,656,407,433]" fontSize="11" level="1" pageId="21" pageNumber="22" reason="1">
<taxonomicName id="53ADFA8C3229330F4BB565C3FD41F899" authority="Rathbun, 1924" authorityName="Rathbun" authorityYear="1924" box="[151,656,407,433]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mjoebergi" subGenus="Austruca">
<emphasis id="A6D95D1D3229330F4BB565C3FD41F899" bold="true" box="[151,656,407,433]" pageId="21" pageNumber="22">
<emphasis id="A6D95D1D3229330F4BB565C3FF19F899" bold="true" box="[151,200,407,433]" italics="true" pageId="21" pageNumber="22">Uca</emphasis>
(
<emphasis id="A6D95D1D3229330F4BFB65C3FE99F899" bold="true" box="[217,328,407,433]" italics="true" pageId="21" pageNumber="22">Austruca</emphasis>
)
<emphasis id="A6D95D1D3229330F4A7A65C3FE00F899" bold="true" box="[344,465,407,433]" italics="true" pageId="21" pageNumber="22">mjoebergi</emphasis>
<bibRefCitation id="F03CFCFE3229330F4AFA65C3FD41F899" author="Rathbun" box="[472,656,407,433]" lastPageId="21" lastPageNumber="37" pageId="21" pageNumber="36" refString="Rathbun M. G. (1924) Results of Dr. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. 37. Brachyura, Albuneidae and Porcellanidae. Arkiv for Zoologi, 16 (23), 1 - 33." type="journal article" year="1924">Rathbun, 1924</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="9412810F3229330F4BB565EEFE24F8FC" blockId="21.[151,656,407,468]" box="[151,501,442,468]" pageId="21" pageNumber="22">
(
<figureCitation id="0C969D8A3229330F4BBD65EEFF2FF8FC" box="[159,254,442,468]" captionStart="FIGURE 12" captionStartId="22.[151,255,1823,1847]" captionTargetBox="[187,1403,353,1788]" captionTargetId="figure@22.[151,1432,320,1827]" captionTargetPageId="22" captionText="FIGURE 12. Uca annulipes (H. Milne Edwards, 1837): a, b, Mozambique, Inhaca Island; c, Thailand, Phuket, Ao Tang Kheng. d – f, Uca mjoebergi Rathbun, 1924: d, Indonesia, Sulawesi Tenggara; e, f, Australia, Darwin Island;. Photos credits: a, b, e, f by T. Detto (The Australian National University); c by S. Komai (Natural History Museum and Institute, Chiba, Japan); d by R. S. K. Barnes (Cambridge University)." httpUri="https://zenodo.org/record/195803/files/figure.png" pageId="21" pageNumber="22">Figs. 12</figureCitation>
d–f, 13a–j, 14b, 15b)
</paragraph>
</subSubSection>
<subSubSection id="DCB7D2843229330F4BB56651FA4DFB54" pageId="21" pageNumber="22" type="reference_group">
<paragraph id="9412810F3229330F4BB56651FE39FB34" blockId="21.[151,1436,517,636]" box="[151,488,517,540]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="B4BDA6213229330F4BB56651FE39FB34" box="[151,488,517,540]" pageId="21" pageNumber="22">
<taxonomicName id="53ADFA8C3229330F4BB56651FE35FB34" ID-CoL="84HGR" authority="Rathbun 1924: 9" authorityName="Rathbun" authorityPageNumber="9" authorityYear="1924" box="[151,484,517,540]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mjobergi">
<emphasis id="A6D95D1D3229330F4BB56651FEF8FB34" box="[151,297,517,540]" italics="true" pageId="21" pageNumber="22">Uca mjobergi</emphasis>
<treatmentCitation id="150CA71E3229330F4A126651FE35FB34" author="Rathbun" box="[304,484,517,540]" page="9" pageId="21" pageNumber="22" year="1924">
<bibRefCitation id="F03CFCFE3229330F4A126651FE1BFB34" author="Rathbun" box="[304,458,517,540]" lastPageId="21" lastPageNumber="37" pageId="21" pageNumber="36" refString="Rathbun M. G. (1924) Results of Dr. E. Mjoberg's Swedish scientific expeditions to Australia 1910 - 1913. 37. Brachyura, Albuneidae and Porcellanidae. Arkiv for Zoologi, 16 (23), 1 - 33." type="journal article" year="1924">Rathbun 1924</bibRefCitation>
: 9
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9412810F3229330F4BB56671FC67FB14" blockId="21.[151,1436,517,636]" box="[151,950,549,572]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="B4BDA6213229330F4BB56671FC67FB14" box="[151,950,549,572]" pageId="21" pageNumber="22">
<taxonomicName id="53ADFA8C3229330F4BB56671FEACFB14" box="[151,381,549,572]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="lactea" subSpecies="mjoebergi">
<emphasis id="A6D95D1D3229330F4BB56671FEACFB14" box="[151,381,549,572]" italics="true" pageId="21" pageNumber="22">Uca lactea mjoebergi</emphasis>
</taxonomicName>
—
<treatmentCitation id="150CA71E3229330F4A876671FD8BFB14" author="Crane" box="[421,602,549,572]" page="299" pageId="21" pageNumber="22" year="1975">
<bibRefCitation id="F03CFCFE3229330F4A876671FDF5FB14" author="Crane" box="[421,548,549,572]" pageId="21" pageNumber="33" refString="Crane J. (1975) Fiddler Crabs of the World. Ocypodidae: Genus Uca. Princeton University Press, Princeton, New Jersey, i - xxiii + 736 pp." type="book" year="1975">Crane 1975</bibRefCitation>
: 299
</treatmentCitation>
, figs. 17A–C, 19O–P, 54H, 69B.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9412810F3229330F4BB56611FA4DFB54" blockId="21.[151,1436,517,636]" pageId="21" pageNumber="22">
<treatmentCitationGroup id="B4BDA6213229330F4BB56611FA4DFB54" pageId="21" pageNumber="22">
<taxonomicName id="53ADFA8C3229330F4BB56611FEE4FB74" box="[151,309,581,604]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="A6D95D1D3229330F4BB56611FEE4FB74" box="[151,309,581,604]" italics="true" pageId="21" pageNumber="22">Uca mjoebergi</emphasis>
</taxonomicName>
—
<treatmentCitation id="150CA71E3229330F4A7E6611FDBAFB74" author="George" box="[348,619,581,604]" page="10" pageId="21" pageNumber="22" year="1982">
<bibRefCitation id="F03CFCFE3229330F4A7E6611FD97FB74" author="George" box="[348,582,581,604]" pageId="21" pageNumber="33" refString="George R. W. &amp; Jones D. S. (1982) A revision of the fiddler crabs of Australia (Ocypodidae: Uca). Record of the Western Australian Museum, Suppl. 14, 99 pp." type="book" year="1982">George &amp; Jones 1982</bibRefCitation>
: 10
</treatmentCitation>
(in key), 80–83, figs. 4b, 7a, 53a–f, 58d. — von Hagen &amp;
<treatmentCitation id="150CA71E3229330F4FFB6611FAA9FB74" author="Jones" box="[1241,1400,581,604]" page="57" pageId="21" pageNumber="22" year="1986">
<bibRefCitation id="F03CFCFE3229330F4FFB6611FA82FB74" author="Jones" box="[1241,1363,581,604]" pageId="21" pageNumber="34" refString="Jones D. A. (1986) A Field Guide to the Sea Shores of Kuwait and the Arabian Gulf. University of Kuwait Blandford Press, Poole, Kuwait (1986), 192 pp." type="book" year="1986">Jones 1986</bibRefCitation>
: 57
</treatmentCitation>
(in key), 66. —
<treatmentCitation id="150CA71E3229330F4A6E6631FDD7FB54" author="Morgan" box="[332,518,613,636]" page="62" pageId="21" pageNumber="22" year="1990">
<bibRefCitation id="F03CFCFE3229330F4A6E6631FE0EFB54" author="Morgan" box="[332,479,613,636]" pageId="21" pageNumber="36" refString="Morgan G. J. (1990) A collection of Thalassinidea, Anomura and Brachyura (Crustacea: Decapoda) from the Kimberley Region of northwestern Australia. Zoologische Verhandelingen, Leiden, 265, 90 pp, figs. 1 - 6." type="book chapter" year="1990">Morgan 1990</bibRefCitation>
: 62
</treatmentCitation>
, 69 (in list). — Davie 2002: 362. —
<treatmentCitation id="150CA71E3229330F48A96631FBBAFB54" author="Shih" box="[907,1131,613,636]" page="377" pageId="21" pageNumber="22" year="2009">
<bibRefCitation id="F03CFCFE3229330F48A96631FBE7FB54" author="Shih" box="[907,1078,613,636]" pageId="21" pageNumber="37" refString="Shih H. - T, Kamrani E., Davie P. J. F &amp; Liu M. Y. (2009) Genetic evidence for the recognition of two fiddler crabs, Uca iranica and U. albimana (Crustacea: Brachyura: Ocypodidae), from the northwestern Indian Ocean, with notes on the U. lactea species-complex. Hydrobiologia, 635, 373 - 382." type="journal article" year="2009">
Shih
<emphasis id="A6D95D1D3229330F48E06631FC3EFB54" box="[962,1007,613,636]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
. 2009
</bibRefCitation>
: 377
</treatmentCitation>
. —
<treatmentCitation id="150CA71E3229330F4FB76631FA82FB54" author="Barnes" box="[1173,1363,613,636]" page="250" pageId="21" pageNumber="22" year="2010">
<bibRefCitation id="F03CFCFE3229330F4FB76631FACEFB54" author="Barnes" box="[1173,1311,613,636]" pageId="21" pageNumber="32" refString="Barnes R. S. K. (2010) A remarkable case of fiddler crab, Uca spp., alpha diversity of Wallacea. Hydrobiologia, 637, 249 - 253." type="journal article" year="2010">Barnes 2010</bibRefCitation>
: 250
</treatmentCitation>
, fig. 1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DCB7D2843229330F4BB566F1FE57FD72" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="9412810F3229330F4BB566F1FEF3FA7C" blockId="21.[151,1437,677,2010]" pageId="21" pageNumber="22">
<emphasis id="A6D95D1D3229330F4BB566F1FE48FB97" bold="true" box="[151,409,677,703]" pageId="21" pageNumber="22">Material examined.</emphasis>
<emphasis id="A6D95D1D3229330F4A8066F1FDCCFB96" box="[418,541,677,702]" italics="true" pageId="21" pageNumber="22">
<collectingCountry id="ECBAC19F3229330F4A8066F1FDCCFB96" box="[418,541,677,702]" name="Indonesia" pageId="21" pageNumber="22">Indonesia</collectingCountry>
</emphasis>
:
<specimenCount id="82AB4A863229330F490D66F1FD52FB97" box="[559,643,677,703]" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
(
<collectionCode id="F2BC19CA3229330F49B666F1FD0CFB97" box="[660,733,677,703]" httpUri="http://grbio.org/cool/wkmc-b7hm" name="National Museum of Natural History, Naturalis" pageId="21" pageNumber="22">NNM</collectionCode>
32589),
<collectingCountry id="ECBAC19F3229330F486C66F1FC1BFB97" box="[846,970,677,703]" name="Indonesia" pageId="21" pageNumber="22">Indonesia</collectingCountry>
, Irian Jaya (= Western New
<collectingCountry id="ECBAC19F3229330F4E1266F1FA5EFB97" box="[1328,1423,677,703]" name="Guinea-Bissau" pageId="21" pageNumber="22">Guinea</collectingCountry>
), mangroves near Kampong Madong, between Mokmer and Bosnek, SE Biak,
<date id="E013A7CF3229330F4F09669EFB63FBCC" box="[1067,1202,714,740]" pageId="21" pageNumber="22" value="1955-02-20">20.02.1955</date>
, leg. L.B. Holthuis;
<specimenCount id="82AB4A863229330F4BB566A4FED5FA22" box="[151,260,752,778]" pageId="21" pageNumber="22" type="male">34 males</specimenCount>
,
<specimenCount id="82AB4A863229330F4A3366A4FE42FA22" box="[273,403,752,778]" pageId="21" pageNumber="22" type="female">14 females</specimenCount>
(UZM),
<specimenCount id="82AB4A863229330F492266A4FD81FA22" box="[512,592,752,778]" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
(
<collectionCode id="F2BC19CA3229330F494366A4FD4FFA22" LSID="urn:lsid:biocol.org:col:34838" box="[609,670,752,778]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="22">SMF</collectionCode>
17163), Banda-Isles, Lontor, Coast near Lavvande, 0 6.06.1922, &quot;Dan. Expedition to the Kei Islands, 1922&quot;;
<specimenCount id="82AB4A863229330F49ED6741FCE4FA07" box="[719,821,789,815]" pageId="21" pageNumber="22" type="male">2 males</specimenCount>
,
<specimenCount id="82AB4A863229330F48616741FC63FA07" box="[835,946,789,815]" pageId="21" pageNumber="22" type="female">1 female</specimenCount>
(UZM), same locality and expedition, 0 7.06.1922.
</paragraph>
<paragraph id="9412810F3229330F4BE66734FE57FD72" blockId="21.[151,1437,677,2010]" pageId="21" pageNumber="22">
<emphasis id="A6D95D1D3229330F4BE66734FEE9FA51" box="[196,312,864,889]" italics="true" pageId="21" pageNumber="22">
<collectingCountry id="ECBAC19F3229330F4BE66734FEE9FA51" box="[196,312,864,889]" name="Australia" pageId="21" pageNumber="22">Australia</collectingCountry>
</emphasis>
: Northern Territory:
<specimenCount id="82AB4A863229330F49616734FD72FA52" box="[579,675,864,890]" pageId="21" pageNumber="22" type="male">2 males</specimenCount>
(
<collectionCode id="F2BC19CA3229330F49966734FCC0FA52" box="[692,785,864,890]" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="21" pageNumber="22">NHMB</collectionCode>
574C), Port Darwin, 1eg. E. Handschin. —Western
<collectingCountry id="ECBAC19F3229330F4BB567D1FED9FAB7" box="[151,264,901,927]" name="Australia" pageId="21" pageNumber="22">Australia</collectingCountry>
:
<specimenCount id="82AB4A863229330F4A3167D1FE50FAB7" box="[275,385,901,927]" pageId="21" pageNumber="22" type="male">10 males</specimenCount>
,
<specimenCount id="82AB4A863229330F4AAF67D1FDDEFAB7" box="[397,527,901,927]" pageId="21" pageNumber="22" type="female">13 females</specimenCount>
(
<collectionCode id="F2BC19CA3229330F493D67D1FDB7FAB7" box="[543,614,901,927]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="21" pageNumber="22">ZMH</collectionCode>
),
<specimenCount id="82AB4A863229330F495E67D1FD0BFAB7" box="[636,730,901,927]" pageId="21" pageNumber="22" type="male">2 males</specimenCount>
,
<specimenCount id="82AB4A863229330F49C567D1FC88FAB7" box="[743,857,901,927]" pageId="21" pageNumber="22" type="female">2 females</specimenCount>
(
<collectionCode id="F2BC19CA3229330F484867D1FC76FAB7" LSID="urn:lsid:biocol.org:col:34838" box="[874,935,901,927]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="22">SMF</collectionCode>
17155), Broome, mangroves,
<date id="E013A7CF3229330F4E3367D1FA49FAB7" box="[1297,1432,901,927]" pageId="21" pageNumber="22" value="1975-11-24">24.11.1975</date>
, leg. G. Hartmann &amp; G. Hartmann-Schröder;
<specimenCount id="82AB4A863229330F498B67FEFCC6FAEC" box="[681,791,938,964]" pageId="21" pageNumber="22" type="male">10 males</specimenCount>
,
<specimenCount id="82AB4A863229330F480667FEFC46FAEC" box="[804,919,938,964]" pageId="21" pageNumber="22" type="female">2 females</specimenCount>
(
<collectionCode id="F2BC19CA3229330F488A67FEFC21FAEC" box="[936,1008,938,964]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="21" pageNumber="22">ZMH</collectionCode>
);
<specimenCount id="82AB4A863229330F4F2467FEFBB3FAEC" box="[1030,1122,938,964]" pageId="21" pageNumber="22" type="male">2 males</specimenCount>
(
<collectionCode id="F2BC19CA3229330F4F5067FEFB7EFAEC" LSID="urn:lsid:biocol.org:col:34838" box="[1138,1199,938,964]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="21" pageNumber="22">SMF</collectionCode>
17156), Harbour,
<quantity id="53552CEA3229330F4EAF67FEFF6CFAC2" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="21" pageNumber="22" unit="km" value="1.0">1 km</quantity>
E. of Onslow, about
<specimenCount id="82AB4A863229330F4A906784FDD5FAC2" box="[434,516,976,1002]" pageId="21" pageNumber="22" type="generic">
<quantity id="53552CEA3229330F4A906784FDD1FAC2" box="[434,512,976,1002]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="21" pageNumber="22" unit="m" value="500.0">500 m</quantity>
.
</specimenCount>
from mouth of Beadon Creek, 0 7.06.1975, leg. G. Hartmannn &amp; G. Hartmann- Schröder;
<specimenCount id="82AB4A863229330F4A3767A1FE54FD27" box="[277,389,1013,1039]" pageId="21" pageNumber="22" type="male">12 males</specimenCount>
,
<specimenCount id="82AB4A863229330F4AB067A1FDD9FD27" box="[402,520,1013,1039]" pageId="21" pageNumber="22" type="female">4 females</specimenCount>
(
<collectionCode id="F2BC19CA3229330F493A67A1FDB0FD27" box="[536,609,1013,1039]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="21" pageNumber="22">ZMH</collectionCode>
), Exmouth, external side of peninsula, Reef near Tantabiddy Creek,
<date id="E013A7CF3229330F4BB5604EFECEFD1C" box="[151,287,1050,1076]" pageId="21" pageNumber="22" value="1975-11-10">11.10.1975</date>
or Carnavon, Pelican Point, inner beach with mangroves,
<date id="E013A7CF3229330F48CC604EFBAAFD1C" box="[1006,1147,1050,1076]" pageId="21" pageNumber="22" value="1975-10-13">13.10.1975</date>
, leg. G. Hartmann &amp; G. Hartmann-Schröder.
</paragraph>
</subSubSection>
<subSubSection id="DCB7D2843229330F4BE66031FAB7FD57" box="[196,1382,1125,1151]" pageId="21" pageNumber="22" type="description">
<paragraph id="9412810F3229330F4BE66031FAB7FD57" blockId="21.[151,1437,677,2010]" box="[196,1382,1125,1151]" pageId="21" pageNumber="22">
<emphasis id="A6D95D1D3229330F4BE66031FEA9FD56" box="[196,376,1125,1150]" italics="true" pageId="21" pageNumber="22">
<collectingCountry id="ECBAC19F3229330F4BE66031FEA9FD56" box="[196,376,1125,1150]" name="Samoa" pageId="21" pageNumber="22">Western Samoa</collectingCountry>
</emphasis>
:
<specimenCount id="82AB4A863229330F4AAA6031FE06FD57" box="[392,471,1125,1151]" pageId="21" pageNumber="22" type="male">1 male</specimenCount>
(SMF 5676), Upolu, vend. Mus. Goddefroy [this locality is probably wrong].
</paragraph>
</subSubSection>
<subSubSection id="DCB7D2843229330F4BE660DEFB4AFC3C" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="9412810F3229330F4BE660DEFB4AFC3C" blockId="21.[151,1437,677,2010]" pageId="21" pageNumber="22">
<emphasis id="A6D95D1D3229330F4BE660DEFE95FD8C" bold="true" box="[196,324,1162,1188]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
Front broad. Anterolateral borders of carapace slightly convergent. Palm of major male chela (
<figureCitation id="0C969D8A3229330F4BBD60E4FED1FDE2" box="[159,256,1200,1226]" captionStart="FIGURE 13" captionStartId="23.[151,256,1611,1635]" captionTargetBox="[302,1307,191,1610]" captionTargetId="figure@23.[279,1321,181,1617]" captionTargetPageId="23" captionText="FIGURE 13. Uca mjoebergi Rathbun, 1924. a – e, male (SMF 17156), Western Australia, 1 km E. Onslow; f – i, males (ZUM), Banda Islands; j, female (SMF 17155), Western Australia, Broome. a, merus of major chela, outer surface; b, merus of major chela, inner surface; c, major chela, outer surface; d, major chela, inner surface; e, G 1, lateral surface. f, g, h, i, apical part of G 1, lateral surface; j, genital opening of female." httpUri="https://zenodo.org/record/195804/files/figure.png" pageId="21" pageNumber="22">Figs. 13</figureCitation>
c, d) without supramarginal groove adjacent to lower margin; oblique ridge at inner surface low. G1 generally as in
<taxonomicName id="53ADFA8C3229330F4A6B6081FE0EFDC6" box="[329,479,1237,1263]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="A6D95D1D3229330F4A6B6081FE8FFDC6" box="[329,350,1237,1262]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4A4F6081FE0EFDC6" box="[365,479,1237,1262]" italics="true" pageId="21" pageNumber="22">annulipes</emphasis>
</taxonomicName>
, but horny endpiece comparatively broader, directed remarkably in a dorsolateral direction (
<figureCitation id="0C969D8A3229330F4A2D60AEFEB5FC3C" box="[271,356,1274,1300]" captionStart="FIGURE 13" captionStartId="23.[151,256,1611,1635]" captionTargetBox="[302,1307,191,1610]" captionTargetId="figure@23.[279,1321,181,1617]" captionTargetPageId="23" captionText="FIGURE 13. Uca mjoebergi Rathbun, 1924. a – e, male (SMF 17156), Western Australia, 1 km E. Onslow; f – i, males (ZUM), Banda Islands; j, female (SMF 17155), Western Australia, Broome. a, merus of major chela, outer surface; b, merus of major chela, inner surface; c, major chela, outer surface; d, major chela, inner surface; e, G 1, lateral surface. f, g, h, i, apical part of G 1, lateral surface; j, genital opening of female." httpUri="https://zenodo.org/record/195804/files/figure.png" pageId="21" pageNumber="22">Fig. 13</figureCitation>
e), mesial border of stem between palp and endpiece bulging outward.
</paragraph>
</subSubSection>
<subSubSection id="DCB7D2843229330F4BE6614BFE62FCAC" pageId="21" pageNumber="22" type="distribution">
<paragraph id="9412810F3229330F4BE6614BFE62FCAC" blockId="21.[151,1437,677,2010]" pageId="21" pageNumber="22">
<emphasis id="A6D95D1D3229330F4BE6614BFEBBFC11" bold="true" box="[196,362,1311,1337]" pageId="21" pageNumber="22">Distribution.</emphasis>
<collectingCountry id="ECBAC19F3229330F4A516174FE36FC12" box="[371,487,1312,1338]" name="Australia" pageId="21" pageNumber="22">Australia</collectingCountry>
: Northern Territory and Western
<collectingCountry id="ECBAC19F3229330F485E6174FC3EFC12" box="[892,1007,1312,1338]" name="Australia" pageId="21" pageNumber="22">Australia</collectingCountry>
, New
<collectingCountry id="ECBAC19F3229330F4F1F6174FB45FC12" box="[1085,1172,1312,1338]" name="Guinea-Bissau" pageId="21" pageNumber="22">Guinea</collectingCountry>
(where it probably is sympatric with
<taxonomicName id="53ADFA8C3229330F4A746111FE22FC76" box="[342,499,1349,1375]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="A6D95D1D3229330F4A746111FEBAFC76" box="[342,363,1349,1374]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4A5E6111FE22FC76" box="[380,499,1349,1374]" italics="true" pageId="21" pageNumber="22">annulipes</emphasis>
</taxonomicName>
), central
<collectingCountry id="ECBAC19F3229330F494B6111FD31FC77" box="[617,736,1349,1375]" name="Indonesia" pageId="21" pageNumber="22">Indonesia</collectingCountry>
(Sulawesi Tenggara,
<collectingCountry id="ECBAC19F3229330F48C86111FBFEFC77" box="[1002,1071,1349,1375]" name="Iceland" pageId="21" pageNumber="22">Pulau</collectingCountry>
Kaledupa) sympatric with
<taxonomicName id="53ADFA8C3229330F4E5D6111FE7EFCAC" authority="Barnes 2010" authorityName="Barnes" authorityYear="2010" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="A6D95D1D3229330F4E5D6111FA45FC76" box="[1407,1428,1349,1374]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4BB5613FFF2FFCAC" box="[151,254,1387,1412]" italics="true" pageId="21" pageNumber="22">perplexa</emphasis>
(
<bibRefCitation id="F03CFCFE3229330F4A2E613EFE76FCAC" author="Barnes" box="[268,423,1386,1412]" pageId="21" pageNumber="32" refString="Barnes R. S. K. (2010) A remarkable case of fiddler crab, Uca spp., alpha diversity of Wallacea. Hydrobiologia, 637, 249 - 253." type="journal article" year="2010">Barnes 2010</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="DCB7D2843229330C4BE661DBFBBBF80A" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="discussion">
<paragraph id="9412810F3229330C4BE661DBFBBBF80A" blockId="21.[151,1437,677,2010]" lastBlockId="22.[151,1438,152,290]" lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
<emphasis id="A6D95D1D3229330F4BE661DBFEECFC81" bold="true" box="[196,317,1423,1449]" pageId="21" pageNumber="22">Remarks.</emphasis>
The most reliable characters to recognise this species is the shape of the G1, especially the curb in the distal part of stem adjacent to the horny endpiece as well as its convexity, of course with a certain amount of variability (see
<figureCitation id="0C969D8A3229330F4ACE618EFD83FCDC" box="[492,594,1498,1524]" captionStart="FIGURE 11" captionStartId="20.[151,255,1494,1518]" captionTargetBox="[237,1360,850,1474]" captionTargetId="figure@20.[215,1372,831,1486]" captionTargetPageId="20" captionText="FIGURE 11. Uca lactea (De Haan, 1835): a, female (SMF 17159) Taiwan; b, c, paralectotype male (ZMG 105), Japan; a, genital opening; b, apical part of G 1, lateral surface; c, apical part of G 1, mesial surface." httpUri="https://zenodo.org/record/195802/files/figure.png" pageId="21" pageNumber="22">Figs. 11</figureCitation>
f–i). It is easily distinguished from
<taxonomicName id="53ADFA8C3229330F4F3E618FFB63FCDC" box="[1052,1202,1498,1524]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="A6D95D1D3229330F4F3E618FFBE0FCDC" box="[1052,1073,1499,1524]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4F61618FFB63FCDC" box="[1091,1202,1499,1524]" italics="true" pageId="21" pageNumber="22">perplexa</emphasis>
</taxonomicName>
by the lack of the supramarginal groove at the outer surface of the palm of the large male chela. From
<taxonomicName id="53ADFA8C3229330F4F4A6254FB2CFF31" box="[1128,1277,1536,1562]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="A6D95D1D3229330F4F4A6254FBACFF31" box="[1128,1149,1536,1561]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4FA96254FB2CFF31" box="[1163,1277,1536,1561]" italics="true" pageId="21" pageNumber="22">annulipes</emphasis>
</taxonomicName>
it can be also easily distinguished by this last character. All Australian specimens examined are typical both in the shape of the G1 and the palm of the male chela. In the examined specimens also the oblique ridge on the inner surface of large male chela palm is typically low as described by
<bibRefCitation id="F03CFCFE3229330F48776224FC24FFA2" author="Crane" box="[853,1013,1648,1674]" pageId="21" pageNumber="33" refString="Crane J. (1975) Fiddler Crabs of the World. Ocypodidae: Genus Uca. Princeton University Press, Princeton, New Jersey, i - xxiii + 736 pp." type="book" year="1975">Crane (1975)</bibRefCitation>
and
<bibRefCitation id="F03CFCFE3229330F4F136224FA80FFA2" author="George" box="[1073,1361,1648,1674]" pageId="21" pageNumber="33" refString="George R. W. &amp; Jones D. S. (1982) A revision of the fiddler crabs of Australia (Ocypodidae: Uca). Record of the Western Australian Museum, Suppl. 14, 99 pp." type="book" year="1982">George &amp; Jones (1982)</bibRefCitation>
. In
<taxonomicName id="53ADFA8C3229330F4E5D6224FEDDFF86" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="A6D95D1D3229330F4E5D6224FA45FFA1" box="[1407,1428,1648,1673]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4BB562C1FEDDFF86" box="[151,268,1685,1710]" italics="true" pageId="21" pageNumber="22">annulipes</emphasis>
</taxonomicName>
this crest is typically high, but there is a certain amount of variability in this character, so that it cannot be used as good discrimination character. In the specimens from the Banda Archipelago different forms of G1 were found, exhibiting some variations from typical
<taxonomicName id="53ADFA8C3229330F488E62B4FB94FFD1" box="[940,1093,1760,1786]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="A6D95D1D3229330F488E62B4FC10FFD1" box="[940,961,1760,1785]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F48F362B4FB94FFD1" box="[977,1093,1760,1785]" italics="true" pageId="21" pageNumber="22">annulipes</emphasis>
</taxonomicName>
to typical
<taxonomicName id="53ADFA8C3229330F4FE562B4FAB5FFD1" box="[1223,1380,1760,1786]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="A6D95D1D3229330F4FE562B4FB0DFFD1" box="[1223,1244,1760,1785]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4FCE62B4FAB5FFD1" box="[1260,1380,1760,1785]" italics="true" pageId="21" pageNumber="22">mjoebergi</emphasis>
</taxonomicName>
(see
<figureCitation id="0C969D8A3229330F4BB56351FF39FE37" box="[151,232,1797,1823]" captionStart="FIGURE 11" captionStartId="20.[151,255,1494,1518]" captionTargetBox="[237,1360,850,1474]" captionTargetId="figure@20.[215,1372,831,1486]" captionTargetPageId="20" captionText="FIGURE 11. Uca lactea (De Haan, 1835): a, female (SMF 17159) Taiwan; b, c, paralectotype male (ZMG 105), Japan; a, genital opening; b, apical part of G 1, lateral surface; c, apical part of G 1, mesial surface." httpUri="https://zenodo.org/record/195802/files/figure.png" pageId="21" pageNumber="22">Fig. 11</figureCitation>
f–i). The specimen from New
<collectingCountry id="ECBAC19F3229330F49696351FD4EFE37" box="[587,671,1797,1823]" name="Guinea-Bissau" pageId="21" pageNumber="22">Guinea</collectingCountry>
identified by
<bibRefCitation id="F03CFCFE3229330F48616351FC0EFE37" author="Crane" box="[835,991,1797,1823]" pageId="21" pageNumber="33" refString="Crane J. (1975) Fiddler Crabs of the World. Ocypodidae: Genus Uca. Princeton University Press, Princeton, New Jersey, i - xxiii + 736 pp." type="book" year="1975">Crane (1975)</bibRefCitation>
as
<taxonomicName id="53ADFA8C3229330F4F276351FB4EFE36" box="[1029,1183,1797,1823]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="A6D95D1D3229330F4F276351FBCBFE36" box="[1029,1050,1797,1822]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4F0B6351FB4EFE36" box="[1065,1183,1797,1822]" italics="true" pageId="21" pageNumber="22">mjoebergi</emphasis>
</taxonomicName>
is quite typical in the major chela. Regarding the features of median and lateral teeth of the gastric mill, this species is readily separated from all other members of this group. The median tooth plate (
<figureCitation id="0C969D8A3229330F48CE6304FB91FE42" box="[1004,1088,1872,1898]" captionStart="FIGURE 14" captionStartId="24.[151,255,1721,1745]" captionTargetBox="[219,1372,595,1720]" captionTargetId="figure@24.[189,1397,563,1735]" captionTargetPageId="24" captionText="FIGURE 14. Median tooth of the gastric mill, ventral face. a, Uca lactea (De Haan, 1835), male (SMF 17161); b, Uca mjoebergi Rathbun, 1924, male (SMF 17163)." httpUri="https://zenodo.org/record/195805/files/figure.png" pageId="21" pageNumber="22">Fig. 14</figureCitation>
b) has four teeth, the two first ones being variable in size, the third and fourth ones nearly equal, one very small tooth attached dorsally to the basal plate. The lateral tooth plate (
<figureCitation id="0C969D8A3229330F495B63CEFD03FE9C" box="[633,722,1946,1972]" captionStart="FIGURE 15" captionStartId="25.[151,255,1362,1386]" captionTargetBox="[183,1402,432,1326]" captionTargetId="figure@25.[174,1412,421,1339]" captionTargetPageId="25" captionText="FIGURE 15. Lateral tooth of the gastric mill, mesial face. a, Uca lactea (De Haan, 1835), male (SMF 17161); b, Uca mjoebergi Rathbun, 1924, male (SMF 17163)." httpUri="https://zenodo.org/record/195806/files/figure.png" pageId="21" pageNumber="22">Fig. 15</figureCitation>
b) is typical in having a shoe-shaped form, with 17 combshaped teeth.
<taxonomicName id="53ADFA8C3229330F4A1A6394FE03FEF1" box="[312,466,1984,2010]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="mjoebergi">
<emphasis id="A6D95D1D3229330F4A1A6394FE9CFEF1" box="[312,333,1984,2009]" italics="true" pageId="21" pageNumber="22">U</emphasis>
.
<emphasis id="A6D95D1D3229330F4A7E6394FE03FEF1" box="[348,466,1984,2009]" italics="true" pageId="21" pageNumber="22">mjoebergi</emphasis>
</taxonomicName>
has a local distribution in northern and Western
<collectingCountry id="ECBAC19F3229330F4F2C6394FBA8FEF2" box="[1038,1145,1984,2010]" name="Australia" pageId="21" pageNumber="22">Australia</collectingCountry>
(Davie 2002). There are also specimens from western New
<collectingCountry id="ECBAC19F322A330C496064CCFD4CF99A" box="[578,669,152,178]" name="Guinea-Bissau" pageId="22" pageNumber="23">Guinea</collectingCountry>
, where it is likely sympatric with
<taxonomicName id="53ADFA8C322A330C4F1D64CCFB0BF999" box="[1087,1242,152,178]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="annulipes">
<emphasis id="A6D95D1D322A330C4F1D64CCFB85F999" box="[1087,1108,152,177]" italics="true" pageId="22" pageNumber="23">U</emphasis>
.
<emphasis id="A6D95D1D322A330C4F4664CCFB0BF999" box="[1124,1242,152,177]" italics="true" pageId="22" pageNumber="23">annulipes</emphasis>
</taxonomicName>
. More recently,
<bibRefCitation id="F03CFCFE322A330C4BB564E9FE96F9FF" author="Barnes" box="[151,327,189,215]" pageId="22" pageNumber="32" refString="Barnes R. S. K. (2010) A remarkable case of fiddler crab, Uca spp., alpha diversity of Wallacea. Hydrobiologia, 637, 249 - 253." type="journal article" year="2010">Barnes (2010)</bibRefCitation>
recorded this species from Sulawesi Tenggara in central
<collectingCountry id="ECBAC19F322A330C4F2C64E9FB57F9FF" box="[1038,1158,189,215]" name="Indonesia" pageId="22" pageNumber="23">Indonesia</collectingCountry>
in sympatry with four different species of fiddler crabs, including
<taxonomicName id="53ADFA8C322A330C498364B6FCFFF9D3" box="[673,814,226,252]" class="Malacostraca" family="Ocypodidae" genus="Uca" kingdom="Animalia" order="Decapoda" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="perplexa">
<emphasis id="A6D95D1D322A330C498364B6FD67F9D3" box="[673,694,226,251]" italics="true" pageId="22" pageNumber="23">U</emphasis>
.
<emphasis id="A6D95D1D322A330C49E464B6FCFFF9D3" box="[710,814,226,251]" italics="true" pageId="22" pageNumber="23">perplexa</emphasis>
</taxonomicName>
from the present group. These records from outside
<collectingCountry id="ECBAC19F322A330C4BB5655CFED2F80A" box="[151,259,264,290]" name="Australia" pageId="22" pageNumber="23">Australia</collectingCountry>
extend its distiribution north to South
<collectingCountry id="ECBAC19F322A330C49EA655CFCEBF80A" box="[712,826,264,290]" name="China" pageId="22" pageNumber="23">East Asia</collectingCountry>
, but still within Wallacea.
</paragraph>
</subSubSection>
</treatment>
</document>
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