153 lines
19 KiB
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153 lines
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<document id="F5394BF0EC6D33840CB134C9C3AA4B0F" ID-DOI="10.5281/zenodo.5404276" ID-ISSN="0374-1036" ID-Zenodo-Dep="5404276" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630132078781" checkinUser="felipe" docAuthor="Hromádka, Lubomír" docDate="2009" docId="142187F5FFC4FF8CF5D4FC07FEF6FBD3" docLanguage="en" docName="ActaEntMusNatPra.2009.49.1.161-190.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 49 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Philonthus aminius , Tottenham 1955" docType="treatment" docVersion="3" lastPageNumber="164" masterDocId="E818FF8DFFC6FF8FF464FFBCFFA1FFC7" masterDocTitle="Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)" masterLastPageNumber="190" masterPageNumber="161" pageNumber="163" updateTime="1699162834734" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CE40B18649A94B80925CE787FDA88848">Revision of the Afrotropical species of the Philonthus caffer species group (Coleoptera: Staphylinidae: Philonthina)</mods:title>
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<mods:namePart id="1E200DA4ECC7CA98FFD4C23C394A3C4D">Hromádka, Lubomír</mods:namePart>
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<mods:typeOfResource id="502D46EA3568368B25E08B42A9270207">text</mods:typeOfResource>
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<mods:title id="CB7D2D0D53F4445C51C5259265E7259A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:part id="A0F45513DBEFD405A76F15B11ABF7181">
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<mods:date id="3DBC4ACEDBE4DDD14C7C266E7E9C899D">2009</mods:date>
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<mods:detail id="53B0815B50EEB73228275E4AE4CCCA2D" type="pubDate">
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<mods:number id="2C8CD198808D16D611DB9069975F4319">2009-06-30</mods:number>
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<mods:number id="0F19F53F947C76AC0CF1D31DC171829C">49</mods:number>
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<mods:number id="5D27111F6533B0DEF5C3CC8E9D3CFC3E">1</mods:number>
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<mods:classification id="9705023059985B17410B7489A919382F">journal article</mods:classification>
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<mods:identifier id="DA539612C876009E00CF8B7D8B1FF03F" type="DOI">10.5281/zenodo.5404276</mods:identifier>
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<mods:identifier id="62C0482380B1E20E75A42B3ABBB14056" type="ISSN">0374-1036</mods:identifier>
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<mods:identifier id="A1C62BE867011A103500D142E352A935" type="Zenodo-Dep">5404276</mods:identifier>
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</mods:mods>
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<treatment id="142187F5FFC4FF8CF5D4FC07FEF6FBD3" LSID="urn:lsid:plazi:treatment:142187F5FFC4FF8CF5D4FC07FEF6FBD3" httpUri="http://treatment.plazi.org/id/142187F5FFC4FF8CF5D4FC07FEF6FBD3" lastPageId="3" lastPageNumber="164" pageId="2" pageNumber="163">
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<subSubSection id="D4926568FFC4FF8DF5D4FC07FCECFC14" box="[432,845,955,980]" pageId="2" pageNumber="163" type="nomenclature">
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<paragraph id="9C3736E3FFC4FF8DF5D4FC07FCECFC14" blockId="2.[432,845,955,980]" box="[432,845,955,980]" pageId="2" pageNumber="163">
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<heading id="C77F818FFFC4FF8DF5D4FC07FCECFC14" bold="true" box="[432,845,955,980]" centered="true" fontSize="8" level="2" pageId="2" pageNumber="163" reason="3">
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<taxonomicName id="5B884D60FFC4FF8DF5D4FC07FCECFC14" ID-CoL="76XW6" authority="Tottenham, 1955" authorityName=", Tottenham" authorityYear="1955" box="[432,845,955,980]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="aminius">
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<emphasis id="AEFCEAF1FFC4FF8DF5D4FC07FCECFC14" bold="true" box="[432,845,955,980]" pageId="2" pageNumber="163">
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<emphasis id="AEFCEAF1FFC4FF8DF5D4FC07FD28FC14" bold="true" box="[432,649,955,980]" italics="true" pageId="2" pageNumber="163">Philonthus aminius</emphasis>
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<bibRefCitation id="F8194B12FFC4FF8DF6EBFC00FCECFC14" author="TOTTENHAM C. E." box="[655,845,955,980]" pageId="2" pageNumber="163" pagination="153 - 195" refId="ref16822" refString="TOTTENHAM C. E. 1955: Studies in the genus Philonthus Stephens (Coleoptera: Staphylinidae). Parts II, III, and IV. Transactions of the Royal Entomological Society of London 106: 153 - 195." type="journal article" year="1955">Tottenham, 1955</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFC4FF8DF634FC66FD0FFC2A" box="[592,686,986,1005]" pageId="2" pageNumber="163" type="description">
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<paragraph id="9C3736E3FFC4FF8DF634FC66FD0FFC2A" blockId="2.[151,1126,986,1127]" box="[592,686,986,1005]" pageId="2" pageNumber="163">
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(
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<figureCitation id="04B32A66FFC4FF8DF632FC66FD06FC2A" box="[598,679,986,1005]" captionStart="Figs" captionStartId="4.[151,190,1507,1526]" captionTargetBox="[168,1121,220,1471]" captionTargetId="figure-10@4.[159,1121,210,1474]" captionTargetPageId="4" captionText="Figs. 1–11. 1–2 – Philonthus aminius Tottenham, 1955; 3–7 – P. aonyx sp. nov.; 8–11 – P. biguttulus Fauvel, 1907 (8–9: Angola; 10–11: Ethiopia). 1, 3, 8, 10 – aedeagus, ventral view; 2 – male sternite VIII, ventral view; 4 – aedeagus, lateral view; 5 – apex of paramere with sensory peg setae, ventral view; 6 – apical portion of male sternite VIII, ventral view; 7 – elytra; 9, 11 – paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="2" pageNumber="163">Figs. 1–2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFC4FF8DF4F3FC41FDA2FBC8" box="[151,515,1020,1039]" pageId="2" pageNumber="163" type="reference_group">
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<paragraph id="9C3736E3FFC4FF8DF4F3FC41FDA2FBC8" blockId="2.[151,1126,986,1127]" box="[151,515,1020,1039]" pageId="2" pageNumber="163">
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<treatmentCitationGroup id="BC9811CDFFC4FF8DF4F3FC41FDA2FBC8" box="[151,515,1020,1039]" pageId="2" pageNumber="163">
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<treatmentCitation id="1D2910F2FFC4FF8DF4F3FC41FE5EFBC8" author="TOTTENHAM C. E." box="[151,511,1020,1039]" page="173" pageId="2" pageNumber="163" year="1955">
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<taxonomicName id="5B884D60FFC4FF8DF4F3FC41FE5EFBC8" ID-CoL="76XW6" authority="Tottenham, 1955: 173" authorityName=", Tottenham" authorityPageNumber="173" authorityYear="1955" box="[151,511,1020,1039]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="aminius">
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<emphasis id="AEFCEAF1FFC4FF8DF4F3FC41FE9CFBC8" box="[151,317,1021,1039]" italics="true" pageId="2" pageNumber="163">Philonthus aminius</emphasis>
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<bibRefCitation id="F8194B12FFC4FF8DF525FC40FE5EFBC8" author="TOTTENHAM C. E." box="[321,511,1020,1039]" pageId="2" pageNumber="163" pagination="153 - 195" refId="ref16822" refString="TOTTENHAM C. E. 1955: Studies in the genus Philonthus Stephens (Coleoptera: Staphylinidae). Parts II, III, and IV. Transactions of the Royal Entomological Society of London 106: 153 - 195." type="journal article" year="1955">Tottenham, 1955: 173</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFC4FF8DF4F3FBA3FCE1FBA0" pageId="2" pageNumber="163" type="materials_examined">
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<paragraph id="9C3736E3FFC4FF8DF4F3FBA3FCE1FBA0" blockId="2.[151,1126,986,1127]" pageId="2" pageNumber="163">
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<emphasis id="AEFCEAF1FFC4FF8DF4F3FBA3FEB1FBF5" bold="true" box="[151,272,1054,1074]" pageId="2" pageNumber="163">Type locality.</emphasis>
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<materialsCitation id="2CE03CBEFFC4FF8DF571FBA3FCE1FBA0" collectingDate="1953-01" collectionCode="BMNH" collectorName="V. R. Eastop & C. E. Tottenham & C. H. BOHEMAN" country="Kenya" location="Kenya" pageId="2" pageNumber="163" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="E49F7673FFC4FF8DF571FBA3FEF1FBF5" box="[277,336,1055,1074]" name="Kenya" pageId="2" pageNumber="163">Kenya</collectingCountry>
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: Muguga.
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<emphasis id="AEFCEAF1FFC4FF8DF4F3FB85FEBDFB8B" bold="true" box="[151,284,1080,1100]" pageId="2" pageNumber="163">Type material.</emphasis>
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<typeStatus id="43338841FFC4FF8DF545FB85FED4FB8C" box="[289,373,1081,1100]" pageId="2" pageNumber="163" type="holotype">HOLOTYPE</typeStatus>
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: J, ‛Muguga,
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<date id="E8361023FFC4FF8DF590FB85FD8DFB8B" box="[500,556,1081,1100]" pageId="2" pageNumber="163" value="1953-01">
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<collectingDate id="F872E9CBFFC4FF8DF590FB85FD8DFB8B" box="[500,556,1081,1100]" pageId="2" pageNumber="163" value="1953-01">i.1953</collectingDate>
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</date>
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,
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<collectorName id="317D5335FFC4FF8DF650FB85FD3EFB8B" box="[564,671,1081,1100]" pageId="2" pageNumber="163">V. R. Eastop</collectorName>
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//
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<taxonomicName id="5B884D60FFC4FF8DF6D2FB86FC62FB8B" authority=", Tottenham" authorityName=", Tottenham" authorityYear="1955" box="[694,963,1081,1100]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="163" phylum="Arthropoda" rank="species" species="aminius">
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<emphasis id="AEFCEAF1FFC4FF8DF6D2FB86FCFDFB8B" box="[694,860,1082,1100]" italics="true" pageId="2" pageNumber="163">Philonthus aminius</emphasis>
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, Tottenham
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</taxonomicName>
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, TYPE //
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<collectorName id="317D5335FFC4FF8DF07CFB85FF77FBA0" pageId="2" pageNumber="163">C.E. Tottenham</collectorName>
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collection,
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<collectorName id="317D5335FFC4FF8DF55EFBE8FEC6FBA0" box="[314,359,1108,1127]" name="C. H. BOHEMAN" pageId="2" pageNumber="163">B.M.</collectorName>
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1974-587 [ochre oblong label, handwritten]’ (
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<collectionCode id="FA99AE26FFC4FF8DF695FBE8FC97FBA0" box="[753,822,1108,1127]" pageId="2" pageNumber="163">BMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFC4FF8CF4F3FBC3FEFDFCB4" lastPageId="3" lastPageNumber="164" pageId="2" pageNumber="163" type="description">
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<paragraph id="9C3736E3FFC4FF8DF4F3FBC3FBB2FB50" blockId="2.[151,1128,1151,1623]" box="[151,1043,1151,1175]" pageId="2" pageNumber="163">
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<emphasis id="AEFCEAF1FFC4FF8DF4F3FBC3FE9CFB50" bold="true" box="[151,317,1151,1175]" pageId="2" pageNumber="163">Redescription.</emphasis>
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Body length
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<quantity id="5B709B06FFC4FF8DF5A9FBC3FD80FB50" box="[461,545,1151,1175]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" pageId="2" pageNumber="163" unit="mm" value="8.8">8.8 mm</quantity>
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, length of fore body (to end of elytra)
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<quantity id="5B709B06FFC4FF8DF7DEFBC3FBAEFB50" box="[954,1039,1151,1175]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="2" pageNumber="163" unit="mm" value="4.1">4.1 mm</quantity>
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.
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</paragraph>
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<paragraph id="9C3736E3FFC4FF8DF4D1FB23FC53FAD0" blockId="2.[151,1128,1151,1623]" pageId="2" pageNumber="163">Colouration. Head, pronotum, elytra and abdomen black, maxillary and labial palpi and mandibles brown, maxillary palpomere 3 somewhat paler, antennae black, antennomere 1 and base of antennomere 2 black-brown, all femora and entire protarsomeres brown-yellow, tibiae dark brown, meso- and metatarsomeres dark brown, slightly paler distally.</paragraph>
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<paragraph id="9C3736E3FFC4FF8DF4D1FAA3FC68FA70" blockId="2.[151,1128,1151,1623]" pageId="2" pageNumber="163">Head quadrate, slightly wider than long (ratio 24: 22). Posterior angles of head slightly rounded, each with one long black bristle. Eyes flat, somewhat longer than temples (ratio 10.0: 8.5). Distance between medial interocular punctures about 4 times as long as distance between medial and lateral interocular punctures. Temporal area each with numerous punctures of variable size. Dorsal surface with very fine microsculpture here and there.</paragraph>
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<paragraph id="9C3736E3FFC4FF8DF4D1FA03FD30F9D0" blockId="2.[151,1128,1151,1623]" pageId="2" pageNumber="163">Antennae long, reaching posterior margin of pronotum when reclined. Antennomeres 1–8 and 11 longer than wide, antennomeres 9–10 as long as wide. Relative length of antennomeres: 1 = 7; 2 = 5; 3 = 6; 4–8 = 4; 9–10 = 3.5; 11 = 5.</paragraph>
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<paragraph id="9C3736E3FFC4FF8CF4D1F99CFBB8FED4" blockId="2.[151,1128,1151,1623]" lastBlockId="3.[113,1089,219,1044]" lastPageId="3" lastPageNumber="164" pageId="2" pageNumber="163">Pronotum vaguely longer than wide (ratio 33: 30), parallel-sided, anterior angles almost rectangular, bluntly rounded, posterior angles obtusely rounded. Each dorsal row with 5 coarse punctures, distances between punctures variable. Each sublateral row with 2 very fine punctures, puncture 2 slightly shifted to lateral margin. Surface without microsculpture.</paragraph>
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<paragraph id="9C3736E3FFC5FF8CF4F4FEA7FF74FEB4" blockId="3.[113,1089,219,1044]" pageId="3" pageNumber="164">Entire scutellum very densely and coarsely punctured. Punctures somewhat larger than those on elytra, separated in transverse direction by distance much smaller than puncture diameter.</paragraph>
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<paragraph id="9C3736E3FFC5FF8CF4F4FEC7FE80FE34" blockId="3.[113,1089,219,1044]" pageId="3" pageNumber="164">Elytra combined wider than long (ratio 38: 34), very slightly widened posteriad. Anterior angles with one long black bristle. Punctation somewhat sparser than that on scutellum, punctures somewhat larger than eye-facets. Surface between punctures without microsculpture; setation greyish.</paragraph>
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<paragraph id="9C3736E3FFC5FF8CF4F4FE40FD1FFD94" blockId="3.[113,1089,219,1044]" pageId="3" pageNumber="164">Legs. Metatibia somewhat longer than metatarsus (ratio 27: 25). Metatarsomere 1 as long as metatarsomere 5, metatarsomere 2 somewhat longer than metatarsomere 3. Relative length of metatarsomeres: 1 = 7; 2 = 4.5; 3 = 4; 4 = 3.5; 5 = 7.</paragraph>
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<paragraph id="9C3736E3FFC5FF8CF4F4FDE7FDA2FD34" blockId="3.[113,1089,219,1044]" pageId="3" pageNumber="164">Abdomen narrowing posteriad. First three visible abdominal tergites with two basal lines, elevated area between basal lines densely punctate. Punctation of all visible tergites very fine and dense. Punctures smaller than eye-facets, separated by puncture diameter in transverse direction, punctures of raindrop shape, contiguous here and there. Surface without microsculpture; setation similar to that on elytra.</paragraph>
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<paragraph id="9C3736E3FFC5FF8CF4F4FD47FE91FC93" blockId="3.[113,1089,219,1044]" pageId="3" pageNumber="164">
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Male. Protarsomeres 1–3 markedly dilated, sub-bilobed, covered with modified pale setae ventrally, protarsomere 4 distinctly narrower than preceding ones. Male sternite IX (
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<figureCitation id="04B32A66FFC5FF8CF796FCA7FB92FCF3" box="[1010,1075,795,820]" captionStart="Figs" captionStartId="4.[151,190,1507,1526]" captionTargetBox="[168,1121,220,1471]" captionTargetId="figure-10@4.[159,1121,210,1474]" captionTargetPageId="4" captionText="Figs. 1–11. 1–2 – Philonthus aminius Tottenham, 1955; 3–7 – P. aonyx sp. nov.; 8–11 – P. biguttulus Fauvel, 1907 (8–9: Angola; 10–11: Ethiopia). 1, 3, 8, 10 – aedeagus, ventral view; 2 – male sternite VIII, ventral view; 4 – aedeagus, lateral view; 5 – apex of paramere with sensory peg setae, ventral view; 6 – apical portion of male sternite VIII, ventral view; 7 – elytra; 9, 11 – paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="3" pageNumber="164">Fig. 2</figureCitation>
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), aedeagus (
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<figureCitation id="04B32A66FFC5FF8CF486FC87FE80FC93" box="[226,289,827,852]" captionStart="Figs" captionStartId="4.[151,190,1507,1526]" captionTargetBox="[168,1121,220,1471]" captionTargetId="figure-10@4.[159,1121,210,1474]" captionTargetPageId="4" captionText="Figs. 1–11. 1–2 – Philonthus aminius Tottenham, 1955; 3–7 – P. aonyx sp. nov.; 8–11 – P. biguttulus Fauvel, 1907 (8–9: Angola; 10–11: Ethiopia). 1, 3, 8, 10 – aedeagus, ventral view; 2 – male sternite VIII, ventral view; 4 – aedeagus, lateral view; 5 – apex of paramere with sensory peg setae, ventral view; 6 – apical portion of male sternite VIII, ventral view; 7 – elytra; 9, 11 – paramere with sensory peg setae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5404278" httpUri="https://zenodo.org/record/5404278/files/figure.png" pageId="3" pageNumber="164">Fig. 1</figureCitation>
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).
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</paragraph>
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<paragraph id="9C3736E3FFC5FF8CF4F4FCE7FEFDFCB4" blockId="3.[113,1089,219,1044]" box="[144,348,859,883]" pageId="3" pageNumber="164">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="D4926568FFC5FF8CF415FCC7FEC7FC34" pageId="3" pageNumber="164" type="diagnosis">
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<paragraph id="9C3736E3FFC5FF8CF415FCC7FC95FC14" blockId="3.[113,1089,219,1044]" pageId="3" pageNumber="164">
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<emphasis id="AEFCEAF1FFC5FF8CF415FCC7FECBFC54" bold="true" box="[113,362,891,915]" pageId="3" pageNumber="164">Differential diagnosis.</emphasis>
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This species is very similar to
|
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<taxonomicName id="5B884D60FFC5FF8CF6C8FCC1FC9DFC53" authorityName="Hromadka" authorityYear="2006" box="[684,828,893,916]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="leptophilus">
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<emphasis id="AEFCEAF1FFC5FF8CF6C8FCC1FC9DFC53" box="[684,828,893,916]" italics="true" pageId="3" pageNumber="164">P. leptophilus</emphasis>
|
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</taxonomicName>
|
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, from which it differs by longer eyes and denser punctation of the scutellum and abdomen. The species differs from all species of
|
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<taxonomicName id="5B884D60FFC5FF8CF567FC01FEFAFC13" authorityName="Boheman" authorityYear="1848" box="[259,347,957,980]" class="Insecta" family="Staphylinidae" genus="Philonthus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="164" phylum="Arthropoda" rank="species" species="caffer">
|
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<emphasis id="AEFCEAF1FFC5FF8CF567FC01FEFAFC13" box="[259,347,957,980]" italics="true" pageId="3" pageNumber="164">P. caffer</emphasis>
|
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</taxonomicName>
|
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species group by the shape of the aedeagus.
|
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</paragraph>
|
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<paragraph id="9C3736E3FFC5FF8CF415FC67FEC7FC34" blockId="3.[113,1089,219,1044]" box="[113,358,987,1011]" pageId="3" pageNumber="164">
|
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|
<emphasis id="AEFCEAF1FFC5FF8CF415FC67FF4EFC34" bold="true" box="[113,239,987,1011]" pageId="3" pageNumber="164">Bionomics.</emphasis>
|
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|
Unknown.
|
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</paragraph>
|
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</subSubSection>
|
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|
<subSubSection id="D4926568FFC5FF8CF415FC47FEF6FBD3" box="[113,343,1019,1044]" pageId="3" pageNumber="164" type="distribution">
|
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|
<paragraph id="9C3736E3FFC5FF8CF415FC47FEF6FBD3" blockId="3.[113,1089,219,1044]" box="[113,343,1019,1044]" pageId="3" pageNumber="164">
|
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|
|
<emphasis id="AEFCEAF1FFC5FF8CF415FC47FEA5FBD4" bold="true" box="[113,260,1019,1043]" pageId="3" pageNumber="164">Distribution.</emphasis>
|
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|
|
<collectingCountry id="E49F7673FFC5FF8CF56EFC40FEF2FBD3" box="[266,339,1020,1044]" name="Kenya" pageId="3" pageNumber="164">Kenya</collectingCountry>
|
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.
|
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</paragraph>
|
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</subSubSection>
|
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</treatment>
|
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</document>
|