487 lines
74 KiB
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487 lines
74 KiB
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<document id="EE05E12CE7A5EA02D274F3C580A501C8" ID-CLB-Dataset="31448" ID-DOI="10.11646/zootaxa.4085.1.1" ID-GBIF-Dataset="429cf011-ec4c-4a61-b894-2bd32b2ffc9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1052381" ID-ZooBank="7F819439-FD77-4EF2-8E26-7B46F9B5FAAB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510872834225" checkinUser="plazi" docAuthor="Golovatch, Sergei, Evsyukov, Aleksandr & Reip, Hans" docDate="2016" docId="03D23F44F63C9D14FF075234FA38C1EA" docLanguage="en" docName="zootaxa.4085.1.1.pdf" docOrigin="Zootaxa 4085 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polydesmidae Leach 1815" docType="key" docVersion="7" lastPageNumber="48" masterDocId="FFEB473CF6129D3BFF905518FFA3C500" masterDocTitle="The millipede family Polydesmidae in the Caucasus (Diplopoda: Polydesmida)" masterLastPageNumber="51" masterPageNumber="1" pageNumber="47" updateTime="1698498788515" updateUser="ExternalLinkService">
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<mods:titleInfo id="ABC5B41E6AD21342BDB1A23015A6FC85">
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<mods:title id="F363185BF39EAACFEBD7B3E5654DA527">The millipede family Polydesmidae in the Caucasus (Diplopoda: Polydesmida)</mods:title>
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<mods:name id="0327B4AD450470EA8E249F387AFDA1FE" type="personal">
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<mods:namePart id="E3B65CC91A92C080E42EF118493B008F">Golovatch, Sergei</mods:namePart>
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<mods:namePart id="4350DE44209886ACE837CB803DBE7326">Evsyukov, Aleksandr</mods:namePart>
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</mods:name>
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<mods:name id="825BA4FAA40F99DF08B46853B9B84952" type="personal">
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<mods:namePart id="DEDB9DAEE217C3AFBC090FB4E22FA1D6">Reip, Hans</mods:namePart>
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<mods:typeOfResource id="FF4EC66D1305288A9544A0CDE5D36B2F">text</mods:typeOfResource>
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<mods:title id="FBA04A0A7C659954B934E39753A43E25">Zootaxa</mods:title>
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<mods:part id="F77D41E52DBC5492D34AB9354EE88EB5">
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<mods:date id="B03384CE66EAEAA661263ED9E380981F">2016</mods:date>
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<mods:number id="D9F403A7EF46E1C8CDCB59972C06004D">2016-03-01</mods:number>
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<mods:number id="8A2E2D4FF7C2369924B205D98B5FD4F1">4085</mods:number>
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<mods:number id="B4A158370363ED89C86CCCF3162E124A">1</mods:number>
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<mods:identifier id="536BBEB555676DBCA0D828B17A358032" type="DOI">10.11646/zootaxa.4085.1.1</mods:identifier>
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<treatment id="03D23F44F63C9D14FF075234FA38C1EA" ID-DOI="http://doi.org/10.5281/zenodo.6079510" ID-GBIF-Taxon="137308709" ID-Zenodo-Dep="6079510" LSID="urn:lsid:plazi:treatment:03D23F44F63C9D14FF075234FA38C1EA" httpUri="http://treatment.plazi.org/id/03D23F44F63C9D14FF075234FA38C1EA" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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<subSubSection id="C361DDD9F63C9D15FF075234FBA9C246" box="[151,1034,1836,1862]" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="8BC48E52F63C9D15FF075234FBA9C246" blockId="46.[151,1034,1836,1862]" box="[151,1034,1836,1862]" pageId="46" pageNumber="47">
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<heading id="D08C393EF63C9D15FF075234FBA9C246" bold="true" box="[151,1034,1836,1862]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
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<emphasis id="B90F5240F63C9D15FF075234FBA9C246" bold="true" box="[151,1034,1836,1862]" pageId="46" pageNumber="47">
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Key to
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<taxonomicName id="4C7BF5D1F63C9D15FF605234FE3EC246" ID-CoL="7NKL4" authorityName="Leach" authorityYear="1815" box="[240,413,1836,1862]" class="Diplopoda" family="Polydesmidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="family">Polydesmidae</taxonomicName>
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species presently known to occur in the Caucasus
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C361DDD9F63C9D14FF07526BFA38C1EA" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="key">
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<keyStep id="308F96F7F63C9D15FF07526BFA38C2E0" pageId="46" pageNumber="47">
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<paragraph id="8BC48E52F63C9D15FF07526BFA38C2A5" blockId="46.[151,1436,1907,2016]" pageId="46" pageNumber="47">
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<keyLead id="308A2D67F63C9D15FF07526BFA38C2A5" pageId="46" pageNumber="47">
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1. Adults with 20 segments, including telson. Collum often as broad as or broader than head. Gonopod telopodite often elongate and subgeniculate distally....................................................................
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<emphasis id="B90F5240F63C9D15FA815297FA20C2A5" box="[1297,1411,1935,1957]" italics="true" pageId="46" pageNumber="47">Polydesmus</emphasis>
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, 2
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</keyLead>
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</paragraph>
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<paragraph id="8BC48E52F63C9D15FF0752B5FA38C2E0" blockId="46.[151,1436,1907,2016]" pageId="46" pageNumber="47">
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<keyLead id="308A2D67F63C9D15FF0752B5FA38C2E0" pageId="46" pageNumber="47">
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- Adults with 19 segments, including telson. Head broader than collum. Gonopod telopodite stout and never subgeniculate distally...................................................................................
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<taxonomicName id="4C7BF5D1F63C9D15FB6852D2FA27C2E0" ID-CoL="7NTRJ" authorityName="Golovatch & Evsyukov & Reip" authorityYear="2016" box="[1272,1412,1994,2016]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="B90F5240F63C9D15FB6852D2FA27C2E0" box="[1272,1412,1994,2016]" italics="true" pageId="46" pageNumber="47">Brachydesmus</emphasis>
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</taxonomicName>
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, 5
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308F96F7F63D9D14FF07558FFA36C422" pageId="47" pageNumber="48">
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<paragraph id="8BC48E52F63D9D14FF07558FFA38C5E7" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
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<keyLead id="308A2D67F63D9D14FF07558FFA38C5E7" pageId="47" pageNumber="48">
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2. Live coloration usually light brown to brown (
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<figureCitation id="134092D7F63D9D14FD33558FFD57C5AC" box="[675,756,151,172]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H – J), P. lignaui Lohmander, 1936 (D – G) and P. abchasius Attems, 1898 (B, C). A, H – J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D – F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B – J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="47" pageNumber="48">Figs 3A</figureCitation>
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,
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<figureCitation id="134092D7F63D9D14FD6F558FFCBCC5AC" box="[767,799,151,172]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B – C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="47" pageNumber="48">5A</figureCitation>
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). Paraterga not upturned, subhorizontal to slightly declivous, dorsum slightly, but clearly convex in the middle (
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<figureCitation id="134092D7F63D9D14FD3355ADFC9EC5CA" box="[675,829,180,202]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H – J), P. lignaui Lohmander, 1936 (D – G) and P. abchasius Attems, 1898 (B, C). A, H – J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D – F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B – J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="47" pageNumber="48">
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Figs 3A, 3H
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<emphasis id="B90F5240F63D9D14FC8A55ACFC85C5CA" bold="true" box="[794,806,180,202]" pageId="47" pageNumber="48">–</emphasis>
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3J
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</figureCitation>
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,
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<figureCitation id="134092D7F63D9D14FCD855ADFCCBC5CA" box="[840,872,181,202]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B – C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="47" pageNumber="48">5A</figureCitation>
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). Collum clearly narrower than head (
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<figureCitation id="134092D7F63D9D14FB4455ADFAEEC5CA" box="[1236,1357,181,202]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H – J), P. lignaui Lohmander, 1936 (D – G) and P. abchasius Attems, 1898 (B, C). A, H – J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D – F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B – J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="47" pageNumber="48">Figs 3A, 3H</figureCitation>
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,
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<figureCitation id="134092D7F63D9D14FAC755ADFAD4C5CA" box="[1367,1399,181,202]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B – C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="47" pageNumber="48">5A</figureCitation>
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)...................................................................................................... 3
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</keyLead>
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</paragraph>
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<paragraph id="8BC48E52F63D9D14FF0755F7FA36C422" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
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<keyLead id="308A2D67F63D9D14FF0755F7FA36C422" pageId="47" pageNumber="48">
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- Coloration largely red to pinkish. Paraterga very broad, held high, about level to a flat mid-dorsal region, a few first postcollum paraterga slightly, but clearly upturned (
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<figureCitation id="134092D7F63D9D14FDF05415FD0FC422" box="[608,684,269,290]" captionStart="FIGURE 2" captionStartId="6.[151,249,1912,1934]" captionTargetBox="[242,1337,202,1870]" captionTargetId="figure@6.[241,1345,193,1881]" captionTargetPageId="6" captionText="FIGURE 2. Habitus of Polydesmus lignaui Lohmander, 1936. A: male from Kherkhvashi, Svanetia, Georgia, dorsal view. B – E: anterior, middle and caudal parts of body, and entire body, respectively, of male from Sukhumi University Campus, Abkhazia, dorsal (B – D) and lateral views (E). F: male ca 12 mm long from an unspecified locality in Abkhazia, dorsal view. Pictures A – E by K. Makarov, taken not to scale; watercolour F after Lignau (1915)." httpUri="https://zenodo.org/record/1052385/files/figure.png" pageId="47" pageNumber="48">Figs 2E</figureCitation>
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,
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<figureCitation id="134092D7F63D9D14FD275415FD70C422" box="[695,723,269,290]" captionStart="FIGURE 3" captionStartId="7.[151,249,1708,1730]" captionTargetBox="[151,1436,199,1687]" captionTargetId="figure@7.[151,1436,193,1687]" captionTargetPageId="7" captionText="FIGURE 3. Habitus of Polydesmus muralewiczi Lohmander, 1936 (A, H – J), P. lignaui Lohmander, 1936 (D – G) and P. abchasius Attems, 1898 (B, C). A, H – J: male from near Kursavka, Stavropol Prov., Russia, entire body, anterior body part, left half of metatergite 10 and caudal body part, respectively, dorsal views. B, C: male from Pasture Abago, Caucasian Nature Reserve, Adygea, Russia, anterior part of body, lateral view, and left half of metatergite 10, dorsal view, respectively. D – F: male from Dombai, Karachaevo-Cherkessia, Russia, anterior body part, left half of metatergite 10 and caudal part of body, respectively, dorsal views. G: female from Dombai, anterior part of body, lateral view. Picture A by K. Makarov, taken not to scale. Scale bar: 1.0 mm (B – J)." httpUri="https://zenodo.org/record/1052387/files/figure.png" pageId="47" pageNumber="48">3B</figureCitation>
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)........................................................ 4
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308F96F7F63D9D14FF075432FA38C47A" pageId="47" pageNumber="48">
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<paragraph id="8BC48E52F63D9D14FF075432FA38C43F" blockId="47.[151,1437,151,1258]" box="[151,1435,297,319]" pageId="47" pageNumber="48">
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<keyLead id="308A2D67F63D9D14FF075432FA38C43F" box="[151,1435,297,319]" pageId="47" pageNumber="48">
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3. Male prefemora strongly bulging laterad (
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<figureCitation id="134092D7F63D9D14FDE25432FD1DC43F" box="[626,702,298,319]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B – C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="47" pageNumber="48">Fig. 5B</figureCitation>
|
|||
|
|
). Gonopod telopodite stout (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FC405432FBBAC43F" box="[976,1049,298,319]" captionStart="FIGURE 5" captionStartId="11.[151,249,1531,1553]" captionTargetBox="[176,1382,214,1507]" captionTargetId="figure@11.[175,1399,193,1508]" captionTargetPageId="11" captionText="FIGURE 5. Polydesmus mediterraneus Daday, 1889: A: male from Gurzuf, Crimea, habitus, dorsal view. B – C: male from Yashtukha, Abkhazia, leg 7, lateral view, and left gonopod, mesal view, respectively. Designations: en, endomere, ex, exomere. Picture A by K. Makarov. Scale bars: 0.2 mm (B), 0.1 mm (C), not to scale (A)." httpUri="https://zenodo.org/record/1052391/files/figure.png" pageId="47" pageNumber="48">Fig. 5C</figureCitation>
|
|||
|
|
).........
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB095431FA38C43F" box="[1177,1435,297,319]" italics="true" pageId="47" pageNumber="48">
|
|||
|
|
Polydesmus
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FA815431FA38C43F" authorityName="Daday" authorityYear="1889" box="[1297,1435,297,319]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="mediterraneus">mediterraneus</taxonomicName>
|
|||
|
|
</emphasis>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF07545FFA38C47A" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF07545FFA38C47A" pageId="47" pageNumber="48">
|
|||
|
|
- Male prefemora only slightly bulging laterad. Gonopod telopodite strongly elongate, subgeniculate distally (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FB7F545FFAC0C45C" box="[1263,1379,327,348]" captionStart="FIGURE 4" captionStartId="8.[151,249,1796,1818]" captionTargetBox="[198,1381,227,1769]" captionTargetId="figure@8.[198,1388,193,1774]" captionTargetPageId="8" captionText="FIGURE 4. Left gonopods of Polydesmus abchasius Attems, 1898, P. lignaui Lohmander, 1936 and P. muralewiczi Lohmander, 1936 (A – C), and right gonopod of P. muralewiczi Lohmander, 1936 (D). A: male from near Lake Ritsa, Abkhazia, mesal view. B: male from Sukhumi Botanical Garden, Abkhazia, mesal view. C: male from near Kursavka, Stavropol Prov., Russia, mesal view. D: male lectotype from near Psebai, Krasnodar Prov., Russia (after Lohmander 1936). Designations: en, endomere; ex, exomere; d, exomere tooth; pu, hairy pulvillus; r, endomere rod. Scale bar: 0.2 mm (A – C), not to scale (D)." httpUri="https://zenodo.org/record/1052389/files/figure.png" pageId="47" pageNumber="48">Fig. 4C, 4D</figureCitation>
|
|||
|
|
)............................................................................................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FA9F547DFA38C47A" authorityName="Lohmander" authorityYear="1936" box="[1295,1435,356,378]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="muralewiczi">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FA9F547DFA38C47A" box="[1295,1435,356,378]" italics="true" pageId="47" pageNumber="48">P. muralewiczi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="308F96F7F63D9D14FF07549AFA3FC4EF" pageId="47" pageNumber="48">
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF07549AFA38C4B5" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF07549AFA38C4B5" pageId="47" pageNumber="48">
|
|||
|
|
4. Length ca
|
|||
|
|
<quantity id="4C8323B7F63D9D14FEDA549AFE11C497" box="[330,434,386,408]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="47" pageNumber="48" unit="mm" value="11.5" valueMax="13.0" valueMin="10.0">
|
|||
|
|
10
|
|||
|
|
<emphasis id="B90F5240F63D9D14FEF2549AFECDC498" bold="true" box="[354,366,386,408]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
13 mm
|
|||
|
|
</quantity>
|
|||
|
|
(male, female), width of midbody metazonae 1.25
|
|||
|
|
<emphasis id="B90F5240F63D9D14FC37549AFC10C498" bold="true" box="[935,947,386,408]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
1.7 (male) or
|
|||
|
|
<quantity id="4C8323B7F63D9D14FBAB549AFB13C497" box="[1083,1200,386,408]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="47" pageNumber="48" unit="mm" value="1.75" valueMax="2.1" valueMin="1.4">
|
|||
|
|
1.4
|
|||
|
|
<emphasis id="B90F5240F63D9D14FBCA549AFBC5C498" bold="true" box="[1114,1126,386,408]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
2.1 mm
|
|||
|
|
</quantity>
|
|||
|
|
(female), metazonite to prozonite width ratio <2.0 (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FE7E5487FD8AC4B4" box="[494,553,415,436]" captionStart="FIGURE 2" captionStartId="6.[151,249,1912,1934]" captionTargetBox="[242,1337,202,1870]" captionTargetId="figure@6.[241,1345,193,1881]" captionTargetPageId="6" captionText="FIGURE 2. Habitus of Polydesmus lignaui Lohmander, 1936. A: male from Kherkhvashi, Svanetia, Georgia, dorsal view. B – E: anterior, middle and caudal parts of body, and entire body, respectively, of male from Sukhumi University Campus, Abkhazia, dorsal (B – D) and lateral views (E). F: male ca 12 mm long from an unspecified locality in Abkhazia, dorsal view. Pictures A – E by K. Makarov, taken not to scale; watercolour F after Lignau (1915)." httpUri="https://zenodo.org/record/1052385/files/figure.png" pageId="47" pageNumber="48">Fig. 2</figureCitation>
|
|||
|
|
). Collum about as broad as head (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FCFF5487FBA4C4B4" box="[879,1031,415,436]" captionStart="FIGURE 2" captionStartId="6.[151,249,1912,1934]" captionTargetBox="[242,1337,202,1870]" captionTargetId="figure@6.[241,1345,193,1881]" captionTargetPageId="6" captionText="FIGURE 2. Habitus of Polydesmus lignaui Lohmander, 1936. A: male from Kherkhvashi, Svanetia, Georgia, dorsal view. B – E: anterior, middle and caudal parts of body, and entire body, respectively, of male from Sukhumi University Campus, Abkhazia, dorsal (B – D) and lateral views (E). F: male ca 12 mm long from an unspecified locality in Abkhazia, dorsal view. Pictures A – E by K. Makarov, taken not to scale; watercolour F after Lignau (1915)." httpUri="https://zenodo.org/record/1052385/files/figure.png" pageId="47" pageNumber="48">Fig. 2A, 2B, 2F</figureCitation>
|
|||
|
|
).........................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FAD054B8FA38C4B5" authorityName="Lohmander" authorityYear="1936" box="[1344,1435,415,437]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="lignaui">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FAD054B8FA38C4B5" box="[1344,1435,415,437]" italics="true" pageId="47" pageNumber="48">P. lignaui</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF0754A5FA3FC4EF" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF0754A5FA3FC4EF" pageId="47" pageNumber="48">
|
|||
|
|
- Length ca
|
|||
|
|
<quantity id="4C8323B7F63D9D14FED654A5FE0EC4D2" box="[326,429,444,466]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.9" metricValueMin="1.3" pageId="47" pageNumber="48" unit="mm" value="16.0" valueMax="19.0" valueMin="13.0">
|
|||
|
|
13
|
|||
|
|
<emphasis id="B90F5240F63D9D14FECE54A4FEC9C4D2" bold="true" box="[350,362,444,466]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
19 mm
|
|||
|
|
</quantity>
|
|||
|
|
(male, female), width of midbody metazonae 1.9
|
|||
|
|
<emphasis id="B90F5240F63D9D14FC1754A4FC30C4D2" bold="true" box="[903,915,444,466]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
3.3 (male) or
|
|||
|
|
<quantity id="4C8323B7F63D9D14FB8654A5FB2AC4D2" box="[1046,1161,444,466]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.5" metricValueMin="2.4" pageId="47" pageNumber="48" unit="mm" value="2.95" valueMax="3.5" valueMin="2.4">
|
|||
|
|
2.4
|
|||
|
|
<emphasis id="B90F5240F63D9D14FBA454A4FBE3C4D2" bold="true" box="[1076,1088,444,466]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
3.5 mm
|
|||
|
|
</quantity>
|
|||
|
|
(female), metazonite to prozonite width ratio> 2.0 (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FE5E54C2FDA5C4EF" box="[462,518,474,495]" captionStart="FIGURE 1" captionStartId="4.[151,249,1436,1458]" captionTargetBox="[161,1428,217,1393]" captionTargetId="figure@4.[151,1436,195,1411]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Polydesmus abchasius Attems, 1898. A: male from Gelgeluk, Abkhazia, lateral view. B: female from near Krasnaya Polyana, Krasnodar Prov., dorsal view. C: female ca 18 mm long from an unspecified locality in Abkhazia, dorsal view. Pictures A and B by K. Makarov, taken not to scale; watercolour C after Lignau (1915)." httpUri="https://zenodo.org/record/1052383/files/figure.png" pageId="47" pageNumber="48">Fig. 1</figureCitation>
|
|||
|
|
). Collum clearly broader than head (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FCF954C2FC03C4EF" box="[873,928,474,495]" captionStart="FIGURE 1" captionStartId="4.[151,249,1436,1458]" captionTargetBox="[161,1428,217,1393]" captionTargetId="figure@4.[151,1436,195,1411]" captionTargetPageId="4" captionText="FIGURE 1. Habitus of Polydesmus abchasius Attems, 1898. A: male from Gelgeluk, Abkhazia, lateral view. B: female from near Krasnaya Polyana, Krasnodar Prov., dorsal view. C: female ca 18 mm long from an unspecified locality in Abkhazia, dorsal view. Pictures A and B by K. Makarov, taken not to scale; watercolour C after Lignau (1915)." httpUri="https://zenodo.org/record/1052383/files/figure.png" pageId="47" pageNumber="48">Fig. 1</figureCitation>
|
|||
|
|
)..............................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FAB454C2FA3FC4EF" authorityName="Attems" authorityYear="1898" box="[1316,1436,473,495]" class="Pinopsida" family="Pinaceae" genus="Picea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="species" species="abchasius">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FAB454C2FA3FC4EF" box="[1316,1436,473,495]" italics="true" pageId="47" pageNumber="48">P. abchasius</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="308F96F7F63D9D14FF0754EFFA38C79F" pageId="47" pageNumber="48">
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF0754EFFA36C747" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF0754EFFA36C747" pageId="47" pageNumber="48">
|
|||
|
|
5. Paraterga poorly-developed, their fore margins and anterolateral corners largely broadly and regularly rounded, not angular (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FF78570DFEE3C72A" box="[232,320,533,554]" captionStart="FIGURE 10" captionStartId="19.[151,249,1583,1605]" captionTargetBox="[151,1417,193,1562]" captionTargetId="figure@19.[151,1436,193,1562]" captionTargetPageId="19" captionText="FIGURE 10. Brachydesmus furcatus Lohmander, 1936. A: habitus, entire body, dorsal view, male from Mount Kogot, Caucasian Nature Reserve, Krasnodar Prov., Russia. B – D: male from near Goryachiy Klyuch, Krasnodar Prov., Russia, B: anterior part of body, dorsal view; C: right half of metatergite 10, dorsal view; D: posterior part of body, dorsal view. E – F: male holotype of B. furcatus, right and left gonopods, lateral and mesal views, respectively (after Lohmander 1936); G: male holotype of B. furcatus exiguus Strasser, 1970, left gonopod, lateral view (after Strasser 1970). Designations: en, endomere; ex, exomere; t, tooth at base of femorite. Picture A by K. Makarov, taken not to scale. Line drawings B – D by S. Dashdamirov, scale bar: 1.0 mm (B – D), not to scale (E – G)." httpUri="https://zenodo.org/record/1052401/files/figure.png" pageId="47" pageNumber="48">Fig. 10C</figureCitation>
|
|||
|
|
). Metatergal sculpture strongly obliterate (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FD46570DFCC7C72A" box="[726,868,532,554]" captionStart="FIGURE 10" captionStartId="19.[151,249,1583,1605]" captionTargetBox="[151,1417,193,1562]" captionTargetId="figure@19.[151,1436,193,1562]" captionTargetPageId="19" captionText="FIGURE 10. Brachydesmus furcatus Lohmander, 1936. A: habitus, entire body, dorsal view, male from Mount Kogot, Caucasian Nature Reserve, Krasnodar Prov., Russia. B – D: male from near Goryachiy Klyuch, Krasnodar Prov., Russia, B: anterior part of body, dorsal view; C: right half of metatergite 10, dorsal view; D: posterior part of body, dorsal view. E – F: male holotype of B. furcatus, right and left gonopods, lateral and mesal views, respectively (after Lohmander 1936); G: male holotype of B. furcatus exiguus Strasser, 1970, left gonopod, lateral view (after Strasser 1970). Designations: en, endomere; ex, exomere; t, tooth at base of femorite. Picture A by K. Makarov, taken not to scale. Line drawings B – D by S. Dashdamirov, scale bar: 1.0 mm (B – D), not to scale (E – G)." httpUri="https://zenodo.org/record/1052401/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
Fig. 10A
|
|||
|
|
<emphasis id="B90F5240F63D9D14FCBB570CFC94C72A" bold="true" box="[811,823,532,554]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
10D
|
|||
|
|
</figureCitation>
|
|||
|
|
). Gonopod exomere (
|
|||
|
|
<emphasis id="B90F5240F63D9D14FBA3570CFBE9C72A" bold="true" box="[1075,1098,532,554]" pageId="47" pageNumber="48">ex</emphasis>
|
|||
|
|
) with a conspicuous densely pilose strip (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FE88572AFE06C747" box="[280,421,562,584]" captionStart="FIGURE 10" captionStartId="19.[151,249,1583,1605]" captionTargetBox="[151,1417,193,1562]" captionTargetId="figure@19.[151,1436,193,1562]" captionTargetPageId="19" captionText="FIGURE 10. Brachydesmus furcatus Lohmander, 1936. A: habitus, entire body, dorsal view, male from Mount Kogot, Caucasian Nature Reserve, Krasnodar Prov., Russia. B – D: male from near Goryachiy Klyuch, Krasnodar Prov., Russia, B: anterior part of body, dorsal view; C: right half of metatergite 10, dorsal view; D: posterior part of body, dorsal view. E – F: male holotype of B. furcatus, right and left gonopods, lateral and mesal views, respectively (after Lohmander 1936); G: male holotype of B. furcatus exiguus Strasser, 1970, left gonopod, lateral view (after Strasser 1970). Designations: en, endomere; ex, exomere; t, tooth at base of femorite. Picture A by K. Makarov, taken not to scale. Line drawings B – D by S. Dashdamirov, scale bar: 1.0 mm (B – D), not to scale (E – G)." httpUri="https://zenodo.org/record/1052401/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
Fig. 10E
|
|||
|
|
<emphasis id="B90F5240F63D9D14FEFC572AFEDBC748" bold="true" box="[364,376,562,584]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
10G
|
|||
|
|
</figureCitation>
|
|||
|
|
)................................................................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FB265729FA36C747" ID-CoL="MVX9" authorityName="Lohmander" authorityYear="1936" box="[1206,1429,561,583]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="furcatus">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB265729FA36C747" box="[1206,1429,561,583]" italics="true" pageId="47" pageNumber="48">Brachydesmus furcatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF075757FA38C79F" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF075757FA38C79F" pageId="47" pageNumber="48">
|
|||
|
|
- Paraterga variously developed, but their fore margins and anterolateral corners mostly clearly angulate (e.g.
|
|||
|
|
<figureCitation id="134092D7F63D9D14FA975757FAE5C764" box="[1287,1350,591,612]" captionStart="FIGURE 6" captionStartId="13.[151,249,1914,1936]" captionTargetBox="[267,1310,424,1889]" captionTargetId="figure@13.[256,1330,421,1889]" captionTargetPageId="13" captionText="FIGURE 6. Habitus of Brachydesmus assimilis Lohmander, 1936. A: male from Belakany, Azerbaijan. B – H: male (B), female (C – E) and male (F – H) from Upper Gunib, Dagestan, Russia. C, F: anterior part of body. D, G: right half of segment 10. E, H: caudal part of body. Pictures A and B by K. Makarov, taken not to scale. Line drawings C – H by S. Dashdamirov, scale bar: 1.0 mm." httpUri="https://zenodo.org/record/1052393/files/figure.png" pageId="47" pageNumber="48">Figs 6</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FAC25757FAD2C764" box="[1362,1393,591,612]" captionStart="FIGURE 9" captionStartId="17.[151,249,1949,1971]" captionTargetBox="[151,1406,333,1938]" captionTargetId="figure@17.[151,1436,333,1938]" captionTargetPageId="17" captionText="FIGURE 9. Brachydesmus superus Latzel, 1884. A: habitus, entire body, dorsal view, male from Stavropol, Russia. B: left gonopod, male from Zheleznovodsk, Stavropol Prov., Russia. Designation: pu, hairy pulvillus. Picture A taken by R. Zuev. Scale bars: 1.0 mm (A), 0.1 mm (B)." httpUri="https://zenodo.org/record/1052399/files/figure.png" pageId="47" pageNumber="48">9A</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FAED5757FA34C764" box="[1405,1431,591,612]" captionStart="FIGURE 12" captionStartId="22.[151,249,1819,1841]" captionTargetBox="[151,1423,209,1797]" captionTargetId="figure@22.[151,1436,193,1797]" captionTargetPageId="22" captionText="FIGURE 12. Morph A of Brachydesmus kalischewskyi Lignau, 1915. A: male (?) from an unspecified locality in Abkhazia, length 15 mm. B: male from Khosta, Krasnodar Prov., Russia. C: male from Pass Mamisoni, W Georgia. Watercolour A after Lignau (1915); pictures B and C by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/1052405/files/figure.png" pageId="47" pageNumber="48">12</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FF715775FE9DC782" box="[225,318,621,642]" captionStart="FIGURE 13" captionStartId="23.[151,249,1428,1450]" captionTargetBox="[159,1427,183,1397]" captionTargetId="figure@23.[159,1428,173,1410]" captionTargetPageId="23" captionText="FIGURE 13. Morph A of Brachydesmus kalischewskyi Lignau, 1915: A – C: female from Pass Ertso, South Ossetia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. D – F: male from Pass Ertso, South Ossetia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. G – H: gonopods of a male from near Batumi, Ajaria, Georgia, ventral and mesal views, respectively. Designations: en, endomere; ex, exomere; i, mesal apical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Line drawings A – F by S. Dashdamirov; G – H after Lohmander (1936). Scale bar: 1.0 mm (A – F), not to scale (G – H)." httpUri="https://zenodo.org/record/1052407/files/figure.png" pageId="47" pageNumber="48">13B, 13E</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FED85775FED1C782" box="[328,370,621,642]" captionStart="FIGURE 15" captionStartId="25.[151,249,1800,1822]" captionTargetBox="[178,1426,213,1734]" captionTargetId="figure@25.[153,1438,211,1747]" captionTargetPageId="25" captionText="FIGURE 15. Morph C of Brachydesmus kalischewskyi Lignau, 1915. A: male from Lower Roki, South Ossetia, habitus, lateral view. B: male from near Krasnaya Polyana, Krasnodar Prov., Russia, habitus, dorsal view. Pictures by K. Makarov, taken not to scale." httpUri="https://zenodo.org/record/1052411/files/figure.png" pageId="47" pageNumber="48">15B</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FEED5775FE78C782" box="[381,475,621,642]" captionStart="FIGURE 16" captionStartId="26.[151,249,1484,1506]" captionTargetBox="[161,1425,226,1439]" captionTargetId="figure@26.[150,1436,211,1445]" captionTargetPageId="26" captionText="FIGURE 16. Morph C of Brachydesmus kalischewskyi Lignau, 1915. A – C: male syntype of B. ferrugineus Lohmander, 1936 from near Borzhomi, Georgia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. D – F: male syntype of B. ferrugineus Lohmander, 1936 from Novyi Afon, Abkhazia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. G – H: gonopods of syntype of B. ferrugineus from Novyi Afon, Abkhazia, ventral and mesal views, respectively. Designations: en, endomere; ex, exomere; i, mesal apical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Line drawings A – F by S. Dashdamirov; G – H after Lohmander (1936). Scale bar: 1.0 mm (A – F), not to scale (G – H)." httpUri="https://zenodo.org/record/1052413/files/figure.png" pageId="47" pageNumber="48">16B, 16E</figureCitation>
|
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|
|
). Metatergal sculpture often more strongly developed. Gonopod exomere without a pilose strip, but endomere often with densely pilose patches................................................................ 6
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="308F96F7F63D9D14FF0757BFFDB9C614" pageId="47" pageNumber="48">
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF0757BFFA38C7DA" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF0757BFFA38C7DA" pageId="47" pageNumber="48">
|
|||
|
|
6. Tergal setae sharp (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FE0957BFFE45C7BC" box="[409,486,679,700]" captionStart="FIGURE 9" captionStartId="17.[151,249,1949,1971]" captionTargetBox="[151,1406,333,1938]" captionTargetId="figure@17.[151,1436,333,1938]" captionTargetPageId="17" captionText="FIGURE 9. Brachydesmus superus Latzel, 1884. A: habitus, entire body, dorsal view, male from Stavropol, Russia. B: left gonopod, male from Zheleznovodsk, Stavropol Prov., Russia. Designation: pu, hairy pulvillus. Picture A taken by R. Zuev. Scale bars: 1.0 mm (A), 0.1 mm (B)." httpUri="https://zenodo.org/record/1052399/files/figure.png" pageId="47" pageNumber="48">Fig. 9A</figureCitation>
|
|||
|
|
). Gonopod (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FDCC57BFFD05C7BC" box="[604,678,679,700]" captionStart="FIGURE 9" captionStartId="17.[151,249,1949,1971]" captionTargetBox="[151,1406,333,1938]" captionTargetId="figure@17.[151,1436,333,1938]" captionTargetPageId="17" captionText="FIGURE 9. Brachydesmus superus Latzel, 1884. A: habitus, entire body, dorsal view, male from Stavropol, Russia. B: left gonopod, male from Zheleznovodsk, Stavropol Prov., Russia. Designation: pu, hairy pulvillus. Picture A taken by R. Zuev. Scale bars: 1.0 mm (A), 0.1 mm (B)." httpUri="https://zenodo.org/record/1052399/files/figure.png" pageId="47" pageNumber="48">Fig. 9B</figureCitation>
|
|||
|
|
) unipartite, slender, only slightly falcate, ventrally with several teeth or spines.............................................................................................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FAA657DDFA38C7DA" ID-CoL="84Z92" authorityName="Latzel" authorityYear="1884" box="[1334,1435,709,730]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="superus">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FAA657DDFA38C7DA" box="[1334,1435,709,730]" italics="true" pageId="47" pageNumber="48">B. superus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF0757FAFDB9C614" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF0757FAFDB9C614" pageId="47" pageNumber="48">- Tergal setae mostly bacilliform or subclavate, only rarely sharpened. Gonopod telopodite bipartite, with a distinct exomere and a smaller, but evident endomere.</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="308F96F7F63D9D14FF075605FA38C66C" pageId="47" pageNumber="48">
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF075605FA38C64F" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF075605FA38C64F" pageId="47" pageNumber="48">
|
|||
|
|
7. Adults <
|
|||
|
|
<quantity id="4C8323B7F63D9D14FED15605FED8C632" box="[321,379,797,818]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="47" pageNumber="48" unit="mm" value="8.0">8 mm</quantity>
|
|||
|
|
long and ≤ 1.0 mm wide. Gonopod exomere with only a single apical tooth (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FB005605FA80C632" box="[1168,1315,796,818]" captionStart="FIGURE 28" captionStartId="45.[151,249,1860,1882]" captionTargetBox="[151,1436,325,1796]" captionTargetId="figure@45.[151,1436,325,1815]" captionTargetPageId="45" captionText="FIGURE 28. Brachydesmus kvavadzei sp. n. A: male paratype, habitus, dorsal view. B: leg 7, lateral view. C – D: right gonopod, mesal and lateral views, respectively. Designations: en, endomere; ex, exomere; e, apical tooth of exomere. Picture A by K. Makarov, taken not to scale. Scale bars: 0.25 mm (B), 0.1 mm (C – D)." httpUri="https://zenodo.org/record/1052437/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
Figs 28C
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB7C5604FB5BC632" bold="true" box="[1260,1272,796,818]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
28D
|
|||
|
|
</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FAA15605FA31C632" box="[1329,1426,796,818]" captionStart="FIGURE 29" captionStartId="46.[151,249,1652,1674]" captionTargetBox="[151,1427,193,1631]" captionTargetId="figure@46.[151,1436,193,1631]" captionTargetPageId="46" captionText="FIGURE 29. Brachydesmus simplex sp. n., male paratype from Sochi, Krasnodar Prov., Russia. A: habitus, dorsal view. B: leg 7, lateral view. C – D: left gonopod, ventral and dorsal views, respectively. E: right gonopod telopodite of male paratype from near Lake Ritsa, Abkhazia, lateral view. F – G: left gonopod of male paratype from Myussera Nature Reserve, Abkhazia, dorsal and ventral views, respectively. Designation: e, lateral apical tooth of exomere. Scale bar: 0.1 mm (B – G), not to scale (A)." httpUri="https://zenodo.org/record/1052439/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
29C
|
|||
|
|
<emphasis id="B90F5240F63D9D14FAC95604FAC6C632" bold="true" box="[1369,1381,796,818]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
29G
|
|||
|
|
</figureCitation>
|
|||
|
|
).
|
|||
|
|
<collectingCountry id="F36CCEC2F63D9D14FF715622FE8EC64F" box="[225,301,826,847]" name="Georgia" pageId="47" pageNumber="48">Colchis</collectingCountry>
|
|||
|
|
, Black Sea coast area............................................................................ 8
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF07564FFA38C66C" blockId="47.[151,1437,151,1258]" box="[151,1435,855,876]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF07564FFA38C66C" box="[151,1435,855,876]" pageId="47" pageNumber="48">
|
|||
|
|
- Adults ≥
|
|||
|
|
<quantity id="4C8323B7F63D9D14FEAA564FFED2C66C" box="[314,369,855,876]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="47" pageNumber="48" unit="mm" value="9.0">9 mm</quantity>
|
|||
|
|
long and ≥ 1.0 mm wide.................................................................... 9
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="308F96F7F63D9D14FF07566DFA3FC100" pageId="47" pageNumber="48">
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF07566DFA3FC6A8" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<materialsCitation id="3B13840FF63D9D14FF07566DFA3FC6A8" ID-GBIF-Occurrence="1701200773" country="Georgia" location="Ajaria" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Ajaria">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF07566DFA3FC6A8" pageId="47" pageNumber="48">
|
|||
|
|
8. Length
|
|||
|
|
<quantity id="4C8323B7F63D9D14FEBB566DFE2CC68A" box="[299,399,884,906]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" pageId="47" pageNumber="48" unit="mm" value="6.75" valueMax="7.5" valueMin="6.0">
|
|||
|
|
6
|
|||
|
|
<emphasis id="B90F5240F63D9D14FEA7566CFEE0C68A" bold="true" box="[311,323,884,906]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
7.5 mm
|
|||
|
|
</quantity>
|
|||
|
|
, width of midbody metazonae 0.8
|
|||
|
|
<emphasis id="B90F5240F63D9D14FD46566CFD41C68A" bold="true" box="[726,738,884,906]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
1.0 mm, metazonite to prozonite width ratio ca 1.5
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB59566CFB76C68A" bold="true" box="[1225,1237,884,906]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
1.6 (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FA90566DFAF9C68A" box="[1280,1370,885,906]" captionStart="FIGURE 29" captionStartId="46.[151,249,1652,1674]" captionTargetBox="[151,1427,193,1631]" captionTargetId="figure@46.[151,1436,193,1631]" captionTargetPageId="46" captionText="FIGURE 29. Brachydesmus simplex sp. n., male paratype from Sochi, Krasnodar Prov., Russia. A: habitus, dorsal view. B: leg 7, lateral view. C – D: left gonopod, ventral and dorsal views, respectively. E: right gonopod telopodite of male paratype from near Lake Ritsa, Abkhazia, lateral view. F – G: left gonopod of male paratype from Myussera Nature Reserve, Abkhazia, dorsal and ventral views, respectively. Designation: e, lateral apical tooth of exomere. Scale bar: 0.1 mm (B – G), not to scale (A)." httpUri="https://zenodo.org/record/1052439/files/figure.png" pageId="47" pageNumber="48">Fig. 29A</figureCitation>
|
|||
|
|
). Male legs slightly, but clearly incrassate (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FDA7568AFD33C6A7" box="[567,656,914,935]" captionStart="FIGURE 29" captionStartId="46.[151,249,1652,1674]" captionTargetBox="[151,1427,193,1631]" captionTargetId="figure@46.[151,1436,193,1631]" captionTargetPageId="46" captionText="FIGURE 29. Brachydesmus simplex sp. n., male paratype from Sochi, Krasnodar Prov., Russia. A: habitus, dorsal view. B: leg 7, lateral view. C – D: left gonopod, ventral and dorsal views, respectively. E: right gonopod telopodite of male paratype from near Lake Ritsa, Abkhazia, lateral view. F – G: left gonopod of male paratype from Myussera Nature Reserve, Abkhazia, dorsal and ventral views, respectively. Designation: e, lateral apical tooth of exomere. Scale bar: 0.1 mm (B – G), not to scale (A)." httpUri="https://zenodo.org/record/1052439/files/figure.png" pageId="47" pageNumber="48">Fig. 29B</figureCitation>
|
|||
|
|
). Gonopod exomere longer (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FC31568AFB90C6A7" box="[929,1075,914,936]" captionStart="FIGURE 29" captionStartId="46.[151,249,1652,1674]" captionTargetBox="[151,1427,193,1631]" captionTargetId="figure@46.[151,1436,193,1631]" captionTargetPageId="46" captionText="FIGURE 29. Brachydesmus simplex sp. n., male paratype from Sochi, Krasnodar Prov., Russia. A: habitus, dorsal view. B: leg 7, lateral view. C – D: left gonopod, ventral and dorsal views, respectively. E: right gonopod telopodite of male paratype from near Lake Ritsa, Abkhazia, lateral view. F – G: left gonopod of male paratype from Myussera Nature Reserve, Abkhazia, dorsal and ventral views, respectively. Designation: e, lateral apical tooth of exomere. Scale bar: 0.1 mm (B – G), not to scale (A)." httpUri="https://zenodo.org/record/1052439/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
Figs 29C
|
|||
|
|
<emphasis id="B90F5240F63D9D14FC6A568AFBA5C6A8" bold="true" box="[1018,1030,914,936]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
29G
|
|||
|
|
</figureCitation>
|
|||
|
|
).
|
|||
|
|
<collectingRegion id="49BF40B0F63D9D14FBD3568AFB22C6A7" box="[1091,1153,914,935]" country="Georgia" name="Ajaria" pageId="47" pageNumber="48">Ajaria</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="F36CCEC2F63D9D14FB1B568AFB78C6A7" box="[1163,1243,914,935]" name="Georgia" pageId="47" pageNumber="48">Georgia</collectingCountry>
|
|||
|
|
..
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FB6C568AFAFCC6A7" ID-CoL="8H7RW" authority="Golovatch & Evsyukov & Reip, 2016" authorityName="Golovatch & Evsyukov & Reip" authorityYear="2016" box="[1276,1375,913,935]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="simplex" status="sp. nov.">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB6C568AFAFCC6A7" box="[1276,1375,913,935]" italics="true" pageId="47" pageNumber="48">B. simplex</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A23CEF3BF63D9D14FAF6568AFA3FC6A8" box="[1382,1436,914,936]" pageId="47" pageNumber="48" rank="species">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FAF6568AFA3FC6A8" bold="true" box="[1382,1436,914,936]" pageId="47" pageNumber="48">sp. n.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
</keyLead>
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF0756B7FA3FC100" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF0756B7FA3FC100" pageId="47" pageNumber="48">
|
|||
|
|
-
|
|||
|
|
<materialsCitation id="3B13840FF63D9D14FF7156B7FBABC6E2" ID-GBIF-Occurrence="1701200771" location="Abkhazia" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Abkhazia">
|
|||
|
|
Length
|
|||
|
|
<quantity id="4C8323B7F63D9D14FEBB56B7FEDEC6C4" box="[299,381,943,965]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="47" pageNumber="48" unit="mm" value="5.5" valueMax="6.0" valueMin="5.0">
|
|||
|
|
5
|
|||
|
|
<emphasis id="B90F5240F63D9D14FEA756B7FEE0C6C5" bold="true" box="[311,323,943,965]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
6 mm
|
|||
|
|
</quantity>
|
|||
|
|
, width of midbody metazonae
|
|||
|
|
<quantity id="4C8323B7F63D9D14FD3756B7FC8AC6C4" box="[679,809,943,965]" metricMagnitude="-4" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="47" pageNumber="48" unit="mm" value="0.675" valueMax="0.7" valueMin="0.65">
|
|||
|
|
0.65
|
|||
|
|
<emphasis id="B90F5240F63D9D14FD4156B7FD7EC6C5" bold="true" box="[721,733,943,965]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
0.7 mm
|
|||
|
|
</quantity>
|
|||
|
|
, metazonite to prozonite width ratio ca 1.3 (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FB4256B7FA8FC6C4" box="[1234,1324,943,964]" captionStart="FIGURE 28" captionStartId="45.[151,249,1860,1882]" captionTargetBox="[151,1436,325,1796]" captionTargetId="figure@45.[151,1436,325,1815]" captionTargetPageId="45" captionText="FIGURE 28. Brachydesmus kvavadzei sp. n. A: male paratype, habitus, dorsal view. B: leg 7, lateral view. C – D: right gonopod, mesal and lateral views, respectively. Designations: en, endomere; ex, exomere; e, apical tooth of exomere. Picture A by K. Makarov, taken not to scale. Scale bars: 0.25 mm (B), 0.1 mm (C – D)." httpUri="https://zenodo.org/record/1052437/files/figure.png" pageId="47" pageNumber="48">Fig. 28A</figureCitation>
|
|||
|
|
). Male legs slender (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FEA356D5FE28C6E2" box="[307,395,973,994]" captionStart="FIGURE 28" captionStartId="45.[151,249,1860,1882]" captionTargetBox="[151,1436,325,1796]" captionTargetId="figure@45.[151,1436,325,1815]" captionTargetPageId="45" captionText="FIGURE 28. Brachydesmus kvavadzei sp. n. A: male paratype, habitus, dorsal view. B: leg 7, lateral view. C – D: right gonopod, mesal and lateral views, respectively. Designations: en, endomere; ex, exomere; e, apical tooth of exomere. Picture A by K. Makarov, taken not to scale. Scale bars: 0.25 mm (B), 0.1 mm (C – D)." httpUri="https://zenodo.org/record/1052437/files/figure.png" pageId="47" pageNumber="48">Fig. 28B</figureCitation>
|
|||
|
|
). Gonopod exomere shorter (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FD3356D5FC97C6E2" box="[675,820,972,994]" captionStart="FIGURE 28" captionStartId="45.[151,249,1860,1882]" captionTargetBox="[151,1436,325,1796]" captionTargetId="figure@45.[151,1436,325,1815]" captionTargetPageId="45" captionText="FIGURE 28. Brachydesmus kvavadzei sp. n. A: male paratype, habitus, dorsal view. B: leg 7, lateral view. C – D: right gonopod, mesal and lateral views, respectively. Designations: en, endomere; ex, exomere; e, apical tooth of exomere. Picture A by K. Makarov, taken not to scale. Scale bars: 0.25 mm (B), 0.1 mm (C – D)." httpUri="https://zenodo.org/record/1052437/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
Figs 28C
|
|||
|
|
<emphasis id="B90F5240F63D9D14FD6B56D4FCA4C6E2" bold="true" box="[763,775,972,994]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
28D
|
|||
|
|
</figureCitation>
|
|||
|
|
).
|
|||
|
|
<collectingRegion id="49BF40B0F63D9D14FCD456D5FC03C6E2" box="[836,928,973,994]" country="Georgia" name="Abkhazia" pageId="47" pageNumber="48">Abkhazia</collectingRegion>
|
|||
|
|
and Sochi
|
|||
|
|
</materialsCitation>
|
|||
|
|
,
|
|||
|
|
<materialsCitation id="3B13840FF63D9D14FB8256D5FA3FC100" ID-GBIF-Occurrence="1701200770" country="Russia" location="Krasnodar" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Krasnodar">
|
|||
|
|
<collectingRegion id="49BF40B0F63D9D14FB8256D5FBD5C6E2" box="[1042,1142,973,994]" country="Russia" name="Krasnodar" pageId="47" pageNumber="48">Krasnodar</collectingRegion>
|
|||
|
|
Prov.,
|
|||
|
|
<collectingCountry id="F36CCEC2F63D9D14FB2956D5FB5AC6E2" box="[1209,1273,973,994]" name="Russia" pageId="47" pageNumber="48">Russia</collectingCountry>
|
|||
|
|
...................................................................................................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FB7656F2FAFCC6FF" ID-CoL="8H7S4" authority="Golovatch & Evsyukov & Reip, 2016" authorityName="Golovatch & Evsyukov & Reip" authorityYear="2016" box="[1254,1375,1001,1023]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kvavadzei" status="sp. nov.">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB7656F2FAFCC6FF" box="[1254,1375,1001,1023]" italics="true" pageId="47" pageNumber="48">B. kvavadzei</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A23CEF3BF63D9D14FAF656F2FA3FC100" box="[1382,1436,1002,1024]" pageId="47" pageNumber="48" rank="species">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FAF656F2FA3FC100" bold="true" box="[1382,1436,1002,1024]" pageId="47" pageNumber="48">sp. n.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
</materialsCitation>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="308F96F7F63D9D14FF07511FFA38C192" pageId="47" pageNumber="48">
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF07511FFA3FC175" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF07511FFA3FC175" pageId="47" pageNumber="48">
|
|||
|
|
9. Gonopod endomere (
|
|||
|
|
<emphasis id="B90F5240F63D9D14FE3E511FFE65C11D" bold="true" box="[430,454,1031,1053]" pageId="47" pageNumber="48">en</emphasis>
|
|||
|
|
) partially densely pilose and clearly bifid; exomere (
|
|||
|
|
<emphasis id="B90F5240F63D9D14FC58511FFC7CC11D" bold="true" box="[968,991,1031,1053]" pageId="47" pageNumber="48">ex</emphasis>
|
|||
|
|
) with 1
|
|||
|
|
<emphasis id="B90F5240F63D9D14FBBB511FFB94C11D" bold="true" box="[1067,1079,1031,1053]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
2 apical teeth
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB2F511FFB69C11D" bold="true" box="[1215,1226,1031,1053]" pageId="47" pageNumber="48">e</emphasis>
|
|||
|
|
and
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB69511FFAA3C11D" bold="true" box="[1273,1280,1031,1053]" pageId="47" pageNumber="48">i</emphasis>
|
|||
|
|
(the latter often subapical), often also with another subapical lobule or denticle
|
|||
|
|
<emphasis id="B90F5240F63D9D14FCDB513CFCF1C13A" bold="true" box="[843,850,1060,1082]" pageId="47" pageNumber="48">l</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="134092D7F63D9D14FCF0513DFC51C13A" box="[864,1010,1060,1082]" captionStart="FIGURE 13" captionStartId="23.[151,249,1428,1450]" captionTargetBox="[159,1427,183,1397]" captionTargetId="figure@23.[159,1428,173,1410]" captionTargetPageId="23" captionText="FIGURE 13. Morph A of Brachydesmus kalischewskyi Lignau, 1915: A – C: female from Pass Ertso, South Ossetia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. D – F: male from Pass Ertso, South Ossetia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. G – H: gonopods of a male from near Batumi, Ajaria, Georgia, ventral and mesal views, respectively. Designations: en, endomere; ex, exomere; i, mesal apical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Line drawings A – F by S. Dashdamirov; G – H after Lohmander (1936). Scale bar: 1.0 mm (A – F), not to scale (G – H)." httpUri="https://zenodo.org/record/1052407/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
Figs 13G
|
|||
|
|
<emphasis id="B90F5240F63D9D14FC2B513CFC64C13A" bold="true" box="[955,967,1060,1082]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
13H
|
|||
|
|
</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FC6D513DFBFAC13A" box="[1021,1113,1060,1082]" captionStart="FIGURE 14" captionStartId="24.[151,249,1831,1853]" captionTargetBox="[173,1411,475,1816]" captionTargetId="figure@24.[166,1426,457,1816]" captionTargetPageId="24" captionText="FIGURE 14. Morph B of Brachydesmus kalischewskyi Lignau, 1915: A: male from Yarymdja, E Caucasus Major, Azerbaijan, habitus, lateral view. B: male from Khosrov Nature Reserve, Caucasus Minor, S Armenia, habitus, dorsal view. C: left gonopod of male holotype of B. karawajewi Lohmander, 1928, from Teberda, Karachaevo-Cherkessia, Russia, mesal view. D – E: male from Verkhniy Agdan (now Gandzakar), Idjevan Distr., Armenia (not Georgia, as mistakenly stated by Lohmander 1936), ventral and mesal views, respectively. Designations: en, endomere; ex, exomere; i, mesal subapical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Pictures A and B by K. Makarov; C – E after Lohmander (1928, 1936). Not to scale." httpUri="https://zenodo.org/record/1052409/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
14C
|
|||
|
|
<emphasis id="B90F5240F63D9D14FBB5513CFB92C13A" bold="true" box="[1061,1073,1060,1082]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
14E
|
|||
|
|
</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FBF5513DFB66C13A" box="[1125,1221,1060,1082]" captionStart="FIGURE 16" captionStartId="26.[151,249,1484,1506]" captionTargetBox="[161,1425,226,1439]" captionTargetId="figure@26.[150,1436,211,1445]" captionTargetPageId="26" captionText="FIGURE 16. Morph C of Brachydesmus kalischewskyi Lignau, 1915. A – C: male syntype of B. ferrugineus Lohmander, 1936 from near Borzhomi, Georgia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. D – F: male syntype of B. ferrugineus Lohmander, 1936 from Novyi Afon, Abkhazia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. G – H: gonopods of syntype of B. ferrugineus from Novyi Afon, Abkhazia, ventral and mesal views, respectively. Designations: en, endomere; ex, exomere; i, mesal apical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Line drawings A – F by S. Dashdamirov; G – H after Lohmander (1936). Scale bar: 1.0 mm (A – F), not to scale (G – H)." httpUri="https://zenodo.org/record/1052413/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
16G
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB1E513CFB39C13A" bold="true" box="[1166,1178,1060,1082]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
16H
|
|||
|
|
</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FB41513DFA8CC13A" box="[1233,1327,1060,1082]" captionStart="FIGURE 18" captionStartId="30.[151,249,1169,1191]" captionTargetBox="[158,1423,184,1156]" captionTargetId="figure@30.[151,1436,171,1168]" captionTargetPageId="30" captionText="FIGURE 18. Morph D of Brachydesmus kalischewskyi Lignau, 1915, male holotype of B. talyschanus Lohmander, 1936: A – C: anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. D – E: left gonopod, mesal and ventral views, respectively. Designations: i, mesal subapical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Line drawings A – C by S. Dashdamirov; D – E after Lohmander (1936). Scale bar: 1.0 mm (A – C), not to scale (D – E)." httpUri="https://zenodo.org/record/1052417/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
18D
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB6A513CFAA5C13A" bold="true" box="[1274,1286,1060,1082]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
18E
|
|||
|
|
</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FAAA513DFA3BC13A" box="[1338,1432,1060,1082]" captionStart="FIGURE 19" captionStartId="34.[151,249,1871,1893]" captionTargetBox="[113,1450,206,1838]" captionTargetId="figure@34.[113,1474,193,1850]" captionTargetPageId="34" captionText="FIGURE 19. Morph E of Brachydesmus kalischewskyi Lignau, 1915, males from near Mestia, Svanetia, Georgia. A – B: habitus, dorsal views. C: leg 7, lateral view. D – E: right gonopod, lateral and mesal views, respectively. Designations: i, mesal subapical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Pictures A and B by K. Makarov. Scale bars: 0.2 mm (C – E), not to scale (A – B)." httpUri="https://zenodo.org/record/1052419/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
19D
|
|||
|
|
<emphasis id="B90F5240F63D9D14FAF3513CFACCC13A" bold="true" box="[1379,1391,1060,1082]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
19E
|
|||
|
|
</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FF71515AFE98C157" box="[225,315,1090,1112]" captionStart="FIGURE 21" captionStartId="36.[151,249,1884,1906]" captionTargetBox="[171,1421,605,1858]" captionTargetId="figure@36.[155,1440,591,1871]" captionTargetPageId="36" captionText="FIGURE 21. Morph F of Brachydesmus kalischewskyi Lignau, 1915. A – C: male from Agveran, Armenia, anterior part of body, right half of metatergite 10 and caudal part of body, respectively, dorsal views. D – E: male from near Shvanidzor, Megri Mt. Range, Armenia, right gonopod, ventral and mesal views, respectively. F: male from near Badara, Mountainous Karabakh, Azerbaijan, right gonopod, mesal view. Line drawings A – C by S. Dashdamirov. Scale bars: 1.0 mm (A – C), 0.2 mm (D – F)." httpUri="https://zenodo.org/record/1052423/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
21D
|
|||
|
|
<emphasis id="B90F5240F63D9D14FE9A515AFEB5C158" bold="true" box="[266,278,1090,1112]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
21F
|
|||
|
|
</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FED6515AFE27C157" box="[326,388,1090,1112]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStart-2="FIGURE 24" captionStart-3="FIGURE 25" captionStartId-0="37.[151,249,1867,1889]" captionStartId-1="38.[151,249,1838,1860]" captionStartId-2="39.[151,249,1335,1357]" captionStartId-3="40.[151,249,1899,1921]" captionTargetBox-0="[185,1478,223,1836]" captionTargetBox-1="[165,1412,213,1803]" captionTargetBox-2="[182,1388,199,1308]" captionTargetBox-3="[230,1376,301,1868]" captionTargetId-0="figure@37.[171,1502,193,1853]" captionTargetId-1="figure@38.[154,1433,193,1817]" captionTargetId-2="figure@39.[166,1420,193,1319]" captionTargetId-3="figure@40.[198,1388,279,1878]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionTargetPageId-3="40" captionText-0="FIGURE 22. Examples of gonopods of Brachydesmus kalischewskyi Lignau, 1915. A: male from Pasture Abago, Caucasian Nature Reserve, Adygea, morph A. B: male from Khosrov Nature Reserve, S Armenia, morph B. C: male from Pass Ertso, South Ossetia, morph A. D: male from Nadirkhanly, Mountainous Karabakh, Azerbaijan, morph F. E: male from Pass Voratan, Zangezur Mt. Range, Armenia, morph B. F: male from near Pass Krestovyi, Georgia, morph B. Designations: i, mesal subapical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Scale bar: 0.2 mm." captionText-1="FIGURE 23. Examples of gonopods of Brachydesmus kalischewskyi Lignau, 1915. A – C: male from Psebe, Krasnodar Prov., Russia, morph A?, lateral, mesal and ventral views, respectively. D – F: male from Zeraboseli, Ajaria, Georgia, morph A?, sublateral, dorsal and ventral views, respectively. Designations: i, mesal subapical tooth of exomere; e, lateral apical tooth of exomere. Scale bars: 0.1 mm." captionText-2="FIGURE 24. Examples of gonopods of Brachydesmus kalischewskyi Lignau, 1915. A – C: male from Farm Azerbaijan, Talysh Mts, Azerbaijan, morph D, dorsal, ventral and mesal views, respectively. D – F: male from Shishkert, Shikahoh Nature Reserve, Armenia, morph B?, mesal, lateral and ventral views, respectively. Designations: i, mesal, subapical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Scale bar: 0.1 mm." captionText-3="FIGURE 25. Examples of gonopods of Brachydesmus kalischewskyi Lignau, 1915. A – C: male from Farm Azerbaijan, Talysh Mts, Azerbaijan, morph D, lateral, mesal and ventral views, respectively. D: male from Arasbaran Wildlife Refuge, East Azerbaijan Prov., Iran, morph D, mesal view. Designations: i, mesal subapical tooth of exomere; e, lateral apical tooth of exomere; l, subapical lobe of exomere. Scale bars: 0.2 mm." httpUri-0="https://zenodo.org/record/1052425/files/figure.png" httpUri-1="https://zenodo.org/record/1052427/files/figure.png" httpUri-2="https://zenodo.org/record/1052429/files/figure.png" httpUri-3="https://zenodo.org/record/1052431/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
22
|
|||
|
|
<emphasis id="B90F5240F63D9D14FECE515AFEC9C158" bold="true" box="[350,362,1090,1112]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
25
|
|||
|
|
</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FE1E515AFE53C157" box="[398,496,1090,1111]" captionStart="FIGURE 27" captionStartId="42.[151,249,1799,1821]" captionTargetBox="[271,1308,206,1762]" captionTargetId="figure@42.[264,1322,193,1778]" captionTargetPageId="42" captionText="FIGURE 27. Brachydesmus kalischewskyi Lignau, 1915, morph B, males from Pasture Abago and Arkhyz, Adygea, Russia, respectively. A: leg 7, lateral view. B: left gonopod, mesal view. C: leg 7, lateral view. D: left gonopod, mesal view. Scale bars: 1.15 mm (A), 0.2 mm (B and D), 0.1 mm (C)." httpUri="https://zenodo.org/record/1052435/files/figure.png" pageId="47" pageNumber="48">27B, 27D</figureCitation>
|
|||
|
|
). Extremely widespread (Map 2) and variable in all other characters, morphism outlined in Table 1......................................................................................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FB6D5178FA3FC175" ID-CoL="MVY5" authorityName="Lignau" authorityYear="1915" box="[1277,1436,1119,1141]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="kalischewskyi">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB6D5178FA3FC175" box="[1277,1436,1119,1141]" italics="true" pageId="47" pageNumber="48">B. kalischewskyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF075165FA38C192" blockId="47.[151,1437,151,1258]" box="[151,1435,1148,1170]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF075165FA38C192" box="[151,1435,1148,1170]" pageId="47" pageNumber="48">
|
|||
|
|
- Gonopods subfalcate, endomere (
|
|||
|
|
<emphasis id="B90F5240F63D9D14FDB35164FD98C192" bold="true" box="[547,571,1148,1170]" pageId="47" pageNumber="48">en</emphasis>
|
|||
|
|
) simple, more or less finger-shaped (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FC305165FC7DC192" box="[928,990,1149,1170]" captionStart="FIGURE 7" captionStartId="14.[151,249,1837,1859]" captionTargetBox="[279,1301,218,1816]" captionTargetId="figure@14.[252,1335,193,1816]" captionTargetPageId="14" captionText="FIGURE 7. Gonopods of Brachydesmus assimilis Lohmander, 1936. A – B: male from Mariamdjvari Nature Reserve, Georgia, lateral and mesal views, respectively. C: male from near Pass Krestovyi, Georgia. D: male syntype from Tbilisi, Georgia (after Lohmander 1936). Designations: en, endomere; ex, exomere; s, spine on exomere; t, tooth near base of exomere; k, spine at base of endomere; b, tooth at base of femorite. Scale bar: 0.1 mm (A – C), not to scale (D)." httpUri="https://zenodo.org/record/1052395/files/figure.png" pageId="47" pageNumber="48">Figs 7</figureCitation>
|
|||
|
|
,
|
|||
|
|
<figureCitation id="134092D7F63D9D14FC785165FB8AC192" box="[1000,1065,1148,1170]" captionStart="FIGURE 8" captionStartId="16.[151,249,1708,1730]" captionTargetBox="[157,1425,306,1672]" captionTargetId="figure@16.[151,1436,303,1685]" captionTargetPageId="16" captionText="FIGURE 8. Brachydesmus pigmentatus Attems, 1951, males from Hyrcan Nature Reserve, Azerbaijan. A: habitus, entire body, dorsal view. B: anterior part of body, dorsal view. C: right half of metatergite 10, dorsal view. D: posterior part of body, dorsal view. E: gonopods in situ, ventral view. F: right gonopod, mesal view. Designations: en, endomere; ex, exomere; k, spine at base of endomere; t, tooth near base of femorite. Pictures A and E by K. Makarov, taken not to scale. Line drawings B – D by S. Dashdamirov, scale bar: 1.0 mm (B – D), 0.3 mm (F)." httpUri="https://zenodo.org/record/1052397/files/figure.png" pageId="47" pageNumber="48">
|
|||
|
|
8E
|
|||
|
|
<emphasis id="B90F5240F63D9D14FB935164FBACC192" bold="true" box="[1027,1039,1148,1170]" pageId="47" pageNumber="48">–</emphasis>
|
|||
|
|
8F
|
|||
|
|
</figureCitation>
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)........................... 10
|
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="308F96F7F63D9D14FF075182FA38C1EA" pageId="47" pageNumber="48">
|
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<paragraph id="8BC48E52F63D9D14FF075182FA36C1CD" blockId="47.[151,1437,151,1258]" pageId="47" pageNumber="48">
|
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<keyLead id="308A2D67F63D9D14FF075182FA36C1CD" pageId="47" pageNumber="48">
|
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10. Tergal setae mostly subclavate (
|
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<figureCitation id="134092D7F63D9D14FD875182FD2EC1AF" box="[535,653,1178,1200]" captionStart="FIGURE 6" captionStartId="13.[151,249,1914,1936]" captionTargetBox="[267,1310,424,1889]" captionTargetId="figure@13.[256,1330,421,1889]" captionTargetPageId="13" captionText="FIGURE 6. Habitus of Brachydesmus assimilis Lohmander, 1936. A: male from Belakany, Azerbaijan. B – H: male (B), female (C – E) and male (F – H) from Upper Gunib, Dagestan, Russia. C, F: anterior part of body. D, G: right half of segment 10. E, H: caudal part of body. Pictures A and B by K. Makarov, taken not to scale. Line drawings C – H by S. Dashdamirov, scale bar: 1.0 mm." httpUri="https://zenodo.org/record/1052393/files/figure.png" pageId="47" pageNumber="48">
|
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|
|
Fig. 6C
|
|||
|
|
<emphasis id="B90F5240F63D9D14FDF15182FDCEC1B0" bold="true" box="[609,621,1178,1200]" pageId="47" pageNumber="48">–</emphasis>
|
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6H
|
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</figureCitation>
|
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|
|
). Gonopod exomere (
|
|||
|
|
<emphasis id="B90F5240F63D9D14FCF15182FCDBC1B0" bold="true" box="[865,888,1178,1200]" pageId="47" pageNumber="48">ex</emphasis>
|
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|
|
) more elaborate (
|
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|
|
<figureCitation id="134092D7F63D9D14FBB55182FBC2C1AF" box="[1061,1121,1178,1199]" captionStart="FIGURE 7" captionStartId="14.[151,249,1837,1859]" captionTargetBox="[279,1301,218,1816]" captionTargetId="figure@14.[252,1335,193,1816]" captionTargetPageId="14" captionText="FIGURE 7. Gonopods of Brachydesmus assimilis Lohmander, 1936. A – B: male from Mariamdjvari Nature Reserve, Georgia, lateral and mesal views, respectively. C: male from near Pass Krestovyi, Georgia. D: male syntype from Tbilisi, Georgia (after Lohmander 1936). Designations: en, endomere; ex, exomere; s, spine on exomere; t, tooth near base of exomere; k, spine at base of endomere; b, tooth at base of femorite. Scale bar: 0.1 mm (A – C), not to scale (D)." httpUri="https://zenodo.org/record/1052395/files/figure.png" pageId="47" pageNumber="48">Fig. 7</figureCitation>
|
|||
|
|
). Much of both Caucasus Major and Caucasus Minor, but not Hyrcania (Map 3).....................................................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FAB651A0FA36C1CD" ID-CoL="MVW3" authorityName="Lohmander" authorityYear="1936" box="[1318,1429,1207,1229]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="assimilis">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FAB651A0FA36C1CD" box="[1318,1429,1207,1229]" italics="true" pageId="47" pageNumber="48">B. assimilis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BC48E52F63D9D14FF0751CDFA38C1EA" blockId="47.[151,1437,151,1258]" box="[151,1435,1236,1258]" pageId="47" pageNumber="48">
|
|||
|
|
<keyLead id="308A2D67F63D9D14FF0751CDFA38C1EA" box="[151,1435,1236,1258]" pageId="47" pageNumber="48">
|
|||
|
|
- Tergal setae sharp (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FE0A51CDFE44C1EA" box="[410,487,1237,1258]" captionStart="FIGURE 9" captionStartId="17.[151,249,1949,1971]" captionTargetBox="[151,1406,333,1938]" captionTargetId="figure@17.[151,1436,333,1938]" captionTargetPageId="17" captionText="FIGURE 9. Brachydesmus superus Latzel, 1884. A: habitus, entire body, dorsal view, male from Stavropol, Russia. B: left gonopod, male from Zheleznovodsk, Stavropol Prov., Russia. Designation: pu, hairy pulvillus. Picture A taken by R. Zuev. Scale bars: 1.0 mm (A), 0.1 mm (B)." httpUri="https://zenodo.org/record/1052399/files/figure.png" pageId="47" pageNumber="48">Fig. 9A</figureCitation>
|
|||
|
|
). Gonopod exomere (
|
|||
|
|
<emphasis id="B90F5240F63D9D14FD2851CCFD6DC1EA" bold="true" box="[696,718,1236,1258]" pageId="47" pageNumber="48">ex</emphasis>
|
|||
|
|
) simple (
|
|||
|
|
<figureCitation id="134092D7F63D9D14FCBA51CDFCD1C1EA" box="[810,882,1237,1258]" captionStart="FIGURE 8" captionStartId="16.[151,249,1708,1730]" captionTargetBox="[157,1425,306,1672]" captionTargetId="figure@16.[151,1436,303,1685]" captionTargetPageId="16" captionText="FIGURE 8. Brachydesmus pigmentatus Attems, 1951, males from Hyrcan Nature Reserve, Azerbaijan. A: habitus, entire body, dorsal view. B: anterior part of body, dorsal view. C: right half of metatergite 10, dorsal view. D: posterior part of body, dorsal view. E: gonopods in situ, ventral view. F: right gonopod, mesal view. Designations: en, endomere; ex, exomere; k, spine at base of endomere; t, tooth near base of femorite. Pictures A and E by K. Makarov, taken not to scale. Line drawings B – D by S. Dashdamirov, scale bar: 1.0 mm (B – D), 0.3 mm (F)." httpUri="https://zenodo.org/record/1052397/files/figure.png" pageId="47" pageNumber="48">Fig. 8F</figureCitation>
|
|||
|
|
). Hyrcania (Map 3)...................
|
|||
|
|
<taxonomicName id="4C7BF5D1F63D9D14FA9C51CDFA38C1EA" ID-CoL="MVYX" authorityName="Attems" authorityYear="1951" box="[1292,1435,1237,1258]" class="Diplopoda" family="Polydesmidae" genus="Brachydesmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Polydesmida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pigmentatus">
|
|||
|
|
<emphasis id="B90F5240F63D9D14FA9C51CDFA38C1EA" box="[1292,1435,1237,1258]" italics="true" pageId="47" pageNumber="48">B. pigmentatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|