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<mods:title id="186CA4227BAEACDE53DE8D77D50ABC52">A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India</mods:title>
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A.DC., 
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15 
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(1): 277 (de 
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– 
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As 
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<bibRefCitation id="EFAF65882F37FF8D0028FCD2E37E2D22" author="Chauhan A. S." box="[631,884,867,894]" pageId="90" pageNumber="91" pagination="174 - 176" refId="ref52920" refString="Chauhan A. S. 1996. Begoniaceae. In: Hajra P. K. (ed.) Flora of Namdapha, Arunachal-Pradesh: 174 - 176. Botanical Survey of India, Calcutta." type="book chapter" year="1996">Chauhan (1996: 176)</bibRefCitation>
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; as 
<taxonomicName id="4C3E63FA2F37FF8D03DAFC1BE40F2D98" authority=": Baruah &amp; Choudbury (2014: 41)" authorityName="Baruah &amp; Choudbury" authorityPageNumber="41" authorityYear="2014" box="[389,1029,938,964]" class="Magnoliopsida" family="Begoniaceae" genus="Casparya" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="90" pageNumber="91" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F37FF8D03DAFC1BE2782D98" box="[389,626,938,964]" italics="true" pageId="90" pageNumber="91">Casparya silletensis</emphasis>
: Baruah &amp; Choudbury (2014: 41)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F37FF8D02E2FC5AE2882B5B" pageId="90" pageNumber="91" type="materials_examined">
<paragraph id="8B8118792F37FF8D02E2FC5AE1732A59" blockId="90.[189,377,1003,1029]" box="[189,377,1003,1029]" pageId="90" pageNumber="91">
<heading id="D0C9AF152F37FF8D02E2FC5AE1732A59" bold="true" box="[189,377,1003,1029]" fontSize="11" level="3" pageId="90" pageNumber="91" reason="3">
<emphasis id="B94AC46B2F37FF8D02E2FC5AE1732A59" bold="true" box="[189,377,1003,1029]" pageId="90" pageNumber="91">Other material</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F37FF8D02E2FBABE2882B5B" blockId="90.[189,1399,1050,1287]" pageId="90" pageNumber="91">
<materialsCitation id="3B5612242F37FF8D02E2FBABE3DB2A68" ID-GBIF-Occurrence="2610421338" box="[189,977,1050,1076]" pageId="90" pageNumber="91">
<collectingCountry id="F32958E92F37FF8D02E2FBABE11F2A68" box="[189,277,1050,1076]" name="India" pageId="90" pageNumber="91">INDIA</collectingCountry>
: 
<emphasis id="B94AC46B2F37FF8D037CFBABE2132A68" bold="true" box="[291,537,1050,1076]" pageId="90" pageNumber="91">
<collectingRegion id="49FAD69B2F37FF8D037CFBABE2132A68" box="[291,537,1050,1076]" country="India" name="Arunachal-Pradesh" pageId="90" pageNumber="91">Arunachal-Pradesh</collectingRegion>
</emphasis>
: Amjee, 
<emphasis id="B94AC46B2F37FF8D00D4FBABE3222A68" box="[651,808,1050,1076]" italics="true" pageId="90" pageNumber="91">Sastry 40556</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D0164FBABE39A2A68" box="[827,912,1050,1076]" country="India" httpUri="http://grbio.org/cool/1csn-k189" name="Botanical Survey of India, Arunachal Pradesh Regional Centre" pageId="90" pageNumber="91" type="Herbarium">ARUN</collectionCode>
n.v.)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D01BEFBABE2392A0B" ID-GBIF-Occurrence="2610421338" pageId="90" pageNumber="91">
Bapu Mountain, Rotung, 
<date id="FF803EB92F37FF8D0749FBABE0F32A0B" pageId="90" pageNumber="91" value="1911-11-24">24 Nov. 1911</date>
, 
<emphasis id="B94AC46B2F37FF8D0359FB8CE1A22A0B" box="[262,424,1085,1111]" italics="true" pageId="90" pageNumber="91">Burkill 37376</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D03E6FB8CE1FF2A0B" box="[441,501,1085,1111]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="90" pageNumber="91" type="Herbarium">CAL</collectionCode>
n.v.)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D001FFB8CE4652A0B" ID-GBIF-Occurrence="2610421338" box="[576,1135,1085,1111]" pageId="90" pageNumber="91">
ibid., 
<date id="FF803EB92F37FF8D00DAFB8CE2F72A0B" box="[645,765,1085,1111]" pageId="90" pageNumber="91" value="1911-11">Nov. 1911</date>
– 
<date id="FF803EB92F37FF8D014FFB8CE3862A0B" box="[784,908,1085,1111]" pageId="90" pageNumber="91" value="1912-03">Mar. 1912</date>
, 
<emphasis id="B94AC46B2F37FF8D01C6FB8CE4312A0B" box="[921,1083,1085,1111]" italics="true" pageId="90" pageNumber="91">Burkill 36700</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D0612FB8CE46C2A0B" box="[1101,1126,1085,1111]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="90" pageNumber="91" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D0623FB8CE1562A27" ID-GBIF-Occurrence="2610421338" pageId="90" pageNumber="91">
Buyang, 
<emphasis id="B94AC46B2F37FF8D06B8FB8CE57C2A0B" box="[1255,1398,1085,1111]" italics="true" pageId="90" pageNumber="91">Dash 32695</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D029AFBD1E1102A26" box="[197,282,1120,1146]" country="India" httpUri="http://grbio.org/cool/1csn-k189" name="Botanical Survey of India, Arunachal Pradesh Regional Centre" pageId="90" pageNumber="91" type="Herbarium">ARUN</collectionCode>
n.v.)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D0333FBD0E4432A27" ID-GBIF-Occurrence="2610421338" box="[364,1097,1120,1147]" pageId="90" pageNumber="91">
Kameng District, 
<date id="FF803EB92F37FF8D001AFBD1E2E62A27" box="[581,748,1120,1147]" pageId="90" pageNumber="91" value="1957-03-24">24 Mar. 1957</date>
, 
<emphasis id="B94AC46B2F37FF8D00A3FBD1E3B32A26" box="[764,953,1120,1146]" italics="true" pageId="90" pageNumber="91">Panigrahi 6029</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D0194FBD1E40D2A26" box="[971,1031,1120,1146]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="90" pageNumber="91" type="Herbarium">CAL</collectionCode>
n.v.)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D0606FBD1E2DB2AC1" ID-GBIF-Occurrence="2610421338" pageId="90" pageNumber="91">
Mishmi Hills, Kamlang Valley, Glo, 
<date id="FF803EB92F37FF8D0311FB32E1E62AC1" box="[334,492,1155,1182]" pageId="90" pageNumber="91" value="1949-03-22">22 Mar. 1949</date>
, 
<emphasis id="B94AC46B2F37FF8D03A7FB32E28F2AC2" box="[504,645,1155,1182]" italics="true" pageId="90" pageNumber="91">Ward 18439</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D00CAFB32E2C22AC1" box="[661,712,1155,1181]" country="United Kingdom" name="Bristol Museum" pageId="90" pageNumber="91">BM</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D0083FB32E51E2AC2" ID-GBIF-Occurrence="2610421338" box="[732,1300,1155,1182]" pageId="90" pageNumber="91">
Sangram to Koloring, 
<emphasis id="B94AC46B2F37FF8D01BEFB32E4652AC2" box="[993,1135,1155,1182]" italics="true" pageId="90" pageNumber="91">Dash 31223</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D0622FB32E4D82AC1" box="[1149,1234,1155,1181]" country="India" httpUri="http://grbio.org/cool/1csn-k189" name="Botanical Survey of India, Arunachal Pradesh Regional Centre" pageId="90" pageNumber="91" type="Herbarium">ARUN</collectionCode>
n.v.).
</materialsCitation>
<materialsCitation id="3B5612242F37FF8D0744FB32E27B2A9D" ID-GBIF-Occurrence="2610421338" pageId="90" pageNumber="91">
<emphasis id="B94AC46B2F37FF8D0744FB32E5642AC1" bold="true" box="[1307,1390,1155,1181]" pageId="90" pageNumber="91">
<collectingRegion id="49FAD69B2F37FF8D0744FB32E5642AC1" box="[1307,1390,1155,1181]" country="India" name="Assam" pageId="90" pageNumber="91">Assam</collectingRegion>
</emphasis>
: Cachar, 
<date id="FF803EB92F37FF8D037DFB16E1962A9D" box="[290,412,1191,1217]" pageId="90" pageNumber="91" value="1873-10">Oct. 1873</date>
, 
<emphasis id="B94AC46B2F37FF8D03F2FB16E20D2A9C" box="[429,519,1191,1216]" italics="true" pageId="90" pageNumber="91">Keenan</emphasis>
s.n. (
<collectionCode id="ED2F80BC2F37FF8D0010FB16E2622A9D" box="[591,616,1191,1217]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="90" pageNumber="91" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D00DDFB17E4F22A9D" ID-GBIF-Occurrence="2610421338" box="[642,1272,1190,1217]" pageId="90" pageNumber="91">
Cachar, Katakhal Forest, 
<date id="FF803EB92F37FF8D01E6FB16E4322A9D" box="[953,1080,1191,1217]" pageId="90" pageNumber="91" value="1888-03">Mar. 1888</date>
, 
<emphasis id="B94AC46B2F37FF8D0616FB16E4872A9C" box="[1097,1165,1191,1216]" italics="true" pageId="90" pageNumber="91">Mann</emphasis>
s.n. (
<collectionCode id="ED2F80BC2F37FF8D0689FB16E4E52A9D" box="[1238,1263,1191,1217]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="90" pageNumber="91" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D0756FB17E2462AB8" ID-GBIF-Occurrence="2610421338" pageId="90" pageNumber="91">
Charduar Forest, 
<date id="FF803EB92F37FF8D0349FB7BE19A2AB8" box="[278,400,1226,1252]" pageId="90" pageNumber="91" value="1887-04">Apr. 1887</date>
, 
<emphasis id="B94AC46B2F37FF8D03C0FB7AE1E92AB8" box="[415,483,1227,1252]" italics="true" pageId="90" pageNumber="91">Mann</emphasis>
s.n. (
<collectionCode id="ED2F80BC2F37FF8D0075FB7BE2492AB8" box="[554,579,1226,1252]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="90" pageNumber="91" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D0004FB7BE3DC2AB8" ID-GBIF-Occurrence="2610421338" box="[603,982,1226,1252]" pageId="90" pageNumber="91">
ibid., 
<date id="FF803EB92F37FF8D00FFFB7BE3102AB8" box="[672,794,1226,1252]" pageId="90" pageNumber="91" value="1887-04">Apr. 1887</date>
, 
<emphasis id="B94AC46B2F37FF8D0176FB7AE3672AB8" box="[809,877,1227,1252]" italics="true" pageId="90" pageNumber="91">Mann</emphasis>
s.n. (
<collectionCode id="ED2F80BC2F37FF8D01EBFB7BE3C72AB8" box="[948,973,1226,1252]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="90" pageNumber="91" type="Herbarium">K</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B5612242F37FF8D01BAFB7BE15A2B5B" ID-GBIF-Occurrence="2610421338" pageId="90" pageNumber="91">
Namchung, 
<date id="FF803EB92F37FF8D0627FB7BE5122AB9" box="[1144,1304,1226,1253]" pageId="90" pageNumber="91" value="1885-04-18">18 Apr. 1885</date>
, 
<emphasis id="B94AC46B2F37FF8D0778FB7BE1122B5B" italics="true" pageId="90" pageNumber="91">Clarke 37937A</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D0376FB5CE1482B5B" box="[297,322,1261,1287]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="90" pageNumber="91" type="Herbarium">K</collectionCode>
).
</materialsCitation>
<materialsCitation id="3B5612242F37FF8D0308FB5CE2882B5B" ID-GBIF-Occurrence="2610421338" box="[343,642,1261,1287]" pageId="90" pageNumber="91">
<emphasis id="B94AC46B2F37FF8D0308FB5CE1CD2B5B" bold="true" box="[343,455,1261,1287]" pageId="90" pageNumber="91">
<collectingRegion id="49FAD69B2F37FF8D0308FB5CE1CD2B5B" box="[343,455,1261,1287]" country="India" name="Manipur" pageId="90" pageNumber="91">Manipur</collectingRegion>
</emphasis>
: 
<emphasis id="B94AC46B2F37FF8D0388FB5FE2442B5B" box="[471,590,1261,1287]" italics="true" pageId="90" pageNumber="91">Watt 6875</emphasis>
(
<collectionCode id="ED2F80BC2F37FF8D0001FB5CE27E2B5B" box="[606,628,1261,1287]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="90" pageNumber="91" type="Herbarium">E</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F37FF8C02E2FA9FE1202F14" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91" type="description">
<paragraph id="8B8118792F37FF8D02E2FA9FE1442B14" blockId="90.[189,334,1326,1352]" box="[189,334,1326,1352]" pageId="90" pageNumber="91">
<heading id="D0C9AF152F37FF8D02E2FA9FE1442B14" bold="true" box="[189,334,1326,1352]" fontSize="11" level="3" pageId="90" pageNumber="91" reason="3">
<emphasis id="B94AC46B2F37FF8D02E2FA9FE1442B14" bold="true" box="[189,334,1326,1352]" pageId="90" pageNumber="91">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F37FF8C02E2FAECE1202F14" blockId="90.[189,1399,1373,1998]" lastBlockId="91.[189,1399,267,328]" lastPageId="91" lastPageNumber="92" pageId="90" pageNumber="91">
Rhizomatous, stemless, monoecious herb, 
<quantity id="4CC6B59C2F37FF8D008EFAECE35E2B2B" box="[721,852,1373,1399]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="90" pageNumber="91" unit="cm" value="50.0" valueMax="60.0" valueMin="40.0">40–60 cm</quantity>
high. Rhizome: 
<quantity id="4CC6B59C2F37FF8D066FFAECE4A12B2B" box="[1072,1195,1373,1399]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="90" pageNumber="91" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
wide, glabrous. Stipules: lanceolate, 10–20 × 
<quantity id="4CC6B59C2F37FF8D0067FA31E28F2BC7" box="[568,645,1408,1435]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="90" pageNumber="91" unit="mm" value="4.0">4 mm</quantity>
, puberulent hairs on reverse, persistent. Leaves: petiole 
<quantity id="4CC6B59C2F37FF8D0718FA31E1022BE2" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="6.0" metricValueMin="1.0" pageId="90" pageNumber="91" unit="cm" value="35.0" valueMax="60.0" valueMin="10.0">10 – 60 cm</quantity>
long, short red tomentose or pubescent; lamina ovate to broadly ovate, basifixed, base cordate with lobes not overlapping, 10–33 × 
<quantity id="4CC6B59C2F37FF8D00C9FA77E32A2BBD" box="[662,800,1478,1505]" metricMagnitude="-1" metricUnit="m" metricValue="1.475" metricValueMax="2.1" metricValueMin="0.85" pageId="90" pageNumber="91" unit="cm" value="14.75" valueMax="21.0" valueMin="8.5">8.5–21 cm</quantity>
, asymmetric, upper surface green, very sparsely puberulous all over or on veins mostly near petiole attachment or glabrous, underside green, red puberulous all over, denser on veins, venation palmate, midrib 
<quantity id="4CC6B59C2F37FF8D01E6F9BCE42F287B" box="[953,1061,1549,1575]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="2.3" metricValueMin="0.7" pageId="90" pageNumber="91" unit="cm" value="15.0" valueMax="23.0" valueMin="7.0">7–23 cm</quantity>
long; margin entire or with small teeth at ends of main veins, with sparse short hairs; apex short acuminate or acute. Inflorescence: cymose, terminal, few; peduncle with puberulous hairs, branching once; male primary 
<quantity id="4CC6B59C2F37FF8D06A9F9E2E57D2832" box="[1270,1399,1619,1646]" metricMagnitude="-1" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="90" pageNumber="91" unit="cm" value="18.5" valueMax="25.0" valueMin="12.0">12 –25 cm</quantity>
long, secondary 
<quantity id="4CC6B59C2F37FF8D03D9F9C6E1E528CD" box="[390,495,1655,1681]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="90" pageNumber="91" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, 6–8 flowers; female primary 
<quantity id="4CC6B59C2F37FF8D01F4F9C7E41228CD" box="[939,1048,1654,1681]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="90" pageNumber="91" unit="cm" value="10.0" valueMax="15.0" valueMin="5.0">5–15 cm</quantity>
long, secondary 0.5(–4) cm long, 1–4 flowers; bracts ovate, 11–35 × 
<quantity id="4CC6B59C2F37FF8D00E7F928E33E28E8" box="[696,820,1689,1716]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="3.5" metricValueMin="0.2" pageId="90" pageNumber="91" unit="mm" value="18.5" valueMax="35.0" valueMin="2.0">2–35 mm</quantity>
, margin entire, deciduous. Male flower: pedicel 
<quantity id="4CC6B59C2F37FF8D02E2F90CE110288B" box="[189,282,1725,1751]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="90" pageNumber="91" unit="cm" value="2.5" valueMax="4.0" valueMin="1.0">1–4 cm</quantity>
long, red puberulous; tepals 4; outer tepals obovate, 4–20 × 
<quantity id="4CC6B59C2F37FF8D0656F90CE49A288B" box="[1033,1168,1725,1752]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="90" pageNumber="91" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
, white to pink, red puberulous on reverse denser near base, margin entire; inner tepals spathulate, 13–17 × 
<quantity id="4CC6B59C2F37FF8D06E6F951E52128A7" box="[1209,1323,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="12.0" metricValueMin="7.0" pageId="90" pageNumber="91" unit="mm" value="9.5" valueMax="12.0" valueMin="7.0">7–12 mm</quantity>
, white to pink, glabrous; androecium with 70–100 stamens, symmetric; filaments 
<quantity id="4CC6B59C2F37FF8D0608F8B2E4B52942" box="[1111,1215,1795,1822]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="90" pageNumber="91" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
long, unequal, slightly fused at base or free; anther oblong elliptic, 
<quantity id="4CC6B59C2F37FF8D014AF896E372291D" box="[789,888,1831,1857]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="90" pageNumber="91" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
long, dehiscing through slits running nearly the entire length of the anther, not hooded, connective extended. Female flower: pedicel 
<quantity id="4CC6B59C2F37FF8D068AF8F8E5392938" box="[1237,1331,1865,1892]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="90" pageNumber="91" unit="cm" value="3.5" valueMax="5.0" valueMin="2.0">2–5 cm</quantity>
long, red puberulous hairs; bracteoles absent; tepals 4, equal obovate-orbicular, ca 16 × 
<quantity id="4CC6B59C2F37FF8D0628F8DCE4C129DB" box="[1143,1227,1901,1928]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="90" pageNumber="91" unit="mm" value="15.0">15 mm</quantity>
, white to pink, puberulous on reverse denser near base, inner tepals similar yet smaller; ovary 3–4-locular, placentae bifid; capsule globose, densely red tomentose, without wings; styles 3–4, convoluted with twisted ends, caduceus. Fruit: upright, globose; capsule spherical, 8–23 × 
<quantity id="4CC6B59C2F36FF8C01F8FEBAE43A2F7A" box="[935,1072,267,294]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="91" pageNumber="92" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 mm</quantity>
, sparsely red tomentose to glabrous.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F36FF8C02E2FEC1E3682F80" pageId="91" pageNumber="92" type="distribution">
<paragraph id="8B8118792F36FF8C02E2FEC1E21F2FD6" blockId="91.[189,533,368,394]" box="[189,533,368,394]" pageId="91" pageNumber="92">
<heading id="D0C9AF152F36FF8C02E2FEC1E21F2FD6" bold="true" box="[189,533,368,394]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="B94AC46B2F36FF8C02E2FEC1E21F2FD6" bold="true" box="[189,533,368,394]" pageId="91" pageNumber="92">Distribution and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F36FF8C02E2FE2FE3682F80" blockId="91.[189,1399,414,476]" pageId="91" pageNumber="92">
Arunachal-Pradesh, Assam and Manipur; also in 
<collectingCountry id="F32958E92F36FF8C017DFE2FE3612FE4" box="[802,875,414,440]" name="China" pageId="91" pageNumber="92">China</collectingCountry>
, 
<collectingCountry id="F32958E92F36FF8C0122FE2FE40C2FE4" box="[893,1030,414,440]" name="Bangladesh" pageId="91" pageNumber="92">Bangladesh</collectingCountry>
and 
<collectingCountry id="F32958E92F36FF8C0614FE2EE4CB2FE5" box="[1099,1217,415,441]" name="Myanmar" pageId="91" pageNumber="92">Myanmar</collectingCountry>
; 
<quantity id="4CC6B59C2F36FF8C068DFE2FE5782FE5" box="[1234,1394,414,441]" metricMagnitude="2" metricUnit="m" metricValue="7.25" metricValueMax="10.0" metricValueMin="4.5" pageId="91" pageNumber="92" unit="m" value="725.0" valueMax="1000.0" valueMin="450.0">450–1000 m</quantity>
. Flowering: April to September; fruiting: April to October.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F36FF8C02E2FDB2E3DE2CEA" pageId="91" pageNumber="92" type="conservation">
<paragraph id="8B8118792F36FF8C02E2FDB2E1BF2C41" blockId="91.[189,437,515,541]" box="[189,437,515,541]" pageId="91" pageNumber="92">
<heading id="D0C9AF152F36FF8C02E2FDB2E1BF2C41" bold="true" box="[189,437,515,541]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="B94AC46B2F36FF8C02E2FDB2E1BF2C41" bold="true" box="[189,437,515,541]" pageId="91" pageNumber="92">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F36FF8C02E2FD83E3DE2CEA" blockId="91.[189,1399,560,694]" pageId="91" pageNumber="92">
Least Concern. 
<taxonomicName id="4C3E63FA2F36FF8C0326FD82E2522C10" authorityName="C. B. Clarke" authorityYear="1879" baseAuthorityName="A. DC." box="[377,600,562,588]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="91" pageNumber="92" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F36FF8C0326FD82E2522C10" box="[377,600,562,588]" italics="true" pageId="91" pageNumber="92">Begonia silletensis</emphasis>
</taxonomicName>
has an AOO of 
<quantity id="4CC6B59C2F36FF8C0140FD83E3602C10" box="[799,874,562,588]" metricMagnitude="4" metricUnit="m" metricValue="4.4" pageId="91" pageNumber="92" unit="km" value="44.0">44 km</quantity>
<superScript id="7C4BB5312F36FF8C0135FD81E3792C62" attach="left" box="[874,883,560,574]" fontSize="6" pageId="91" pageNumber="92">2</superScript>
and an EOO of 
<quantity id="4CC6B59C2F36FF8C0664FD83E4B02C10" box="[1083,1210,562,588]" metricMagnitude="7" metricUnit="m" metricValue="8.77" pageId="91" pageNumber="92" unit="km" value="87700.0">87,700 km</quantity>
<superScript id="7C4BB5312F36FF8C06E5FD81E4C92C62" attach="left" box="[1210,1219,560,574]" fontSize="6" pageId="91" pageNumber="92">2</superScript>
suggesting the species could be Vulnerable. There is, however, suitable habitat available in Northeast 
<collectingCountry id="F32958E92F36FF8C06F9FDE4E4EF2C33" box="[1190,1253,597,623]" name="India" pageId="91" pageNumber="92">India</collectingCountry>
, particularly the mountains of Arunachal-Pradesh and down into the Arakan mountain range. The species has been recorded near the Namphada Reserve and the Pakke Tiger Reserve.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F36FF8B02E2FD6CE1602CBA" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92" type="discussion">
<paragraph id="8B8118792F36FF8C02E2FD6CE1252CAB" blockId="91.[189,303,733,759]" box="[189,303,733,759]" pageId="91" pageNumber="92">
<heading id="D0C9AF152F36FF8C02E2FD6CE1252CAB" bold="true" box="[189,303,733,759]" fontSize="11" level="3" pageId="91" pageNumber="92" reason="3">
<emphasis id="B94AC46B2F36FF8C02E2FD6CE1252CAB" bold="true" box="[189,303,733,759]" pageId="91" pageNumber="92">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F36FF8B02E2FCBAE42C2FEE" blockId="91.[189,1399,779,1158]" lastBlockId="92.[189,1399,267,435]" lastPageId="92" lastPageNumber="93" pageId="91" pageNumber="92">
There has been much confusion over the delimitation of this species and how it can be separated from its nearest relative, 
<taxonomicName id="4C3E63FA2F36FF8C03F2FC9EE2482D14" authorityName="Dunn (Dunn" authorityYear="1920" box="[429,578,814,840]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="91" pageNumber="92" phylum="Tracheophyta" rank="species" species="aborensis">
<emphasis id="B94AC46B2F36FF8C03F2FC9EE2482D14" box="[429,578,814,840]" italics="true" pageId="91" pageNumber="92">B. aborensis</emphasis>
</taxonomicName>
, by using hair characters. 
<taxonomicName id="4C3E63FA2F36FF8C01D4FC9EE4662D14" authorityName="C. B. Clarke" authorityYear="1879" baseAuthorityName="A. DC." box="[907,1132,814,840]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="91" pageNumber="92" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F36FF8C01D4FC9EE4662D14" box="[907,1132,814,840]" italics="true" pageId="91" pageNumber="92">Begonia silletensis</emphasis>
</taxonomicName>
is described as being glabrous (
<bibRefCitation id="EFAF65882F36FF8C036EFCE3E2252D30" author="Tebbitt M. C. &amp; Guan K. Y." box="[305,559,850,876]" pageId="91" pageNumber="92" pagination="133 - 136" refId="ref57656" refString="Tebbitt M. C. &amp; Guan K. Y. 2002. Emended circumscription of Begonia silletensis (Begoniaceae) and description of a new subspecies from Yunnan, China. Novon 12: 133 - 136. https: // doi. org / 10.2307 / 3393252" type="journal article" year="2002">Tebbitt &amp; Guan 2002</bibRefCitation>
; 
<bibRefCitation id="EFAF65882F36FF8C0064FCE3E31C2D30" author="Dash S. S. &amp; Mao A. A." box="[571,790,850,876]" pageId="91" pageNumber="92" pagination="2095 - 2099" refId="ref53346" refString="Dash S. S. &amp; Mao A. A. 2011. Distribution of six little known plant species from Arunachal-Pradesh, India. Journal of Threatened Taxa 3 (9): 2095 - 2099. https: // doi. org / 10.11609 / jott. o 2688.2095 - 9" type="journal article" year="2011">Dash &amp; Mao 2011</bibRefCitation>
; 
<bibRefCitation id="EFAF65882F36FF8C017DFCE3E3C92D30" author="Morris R." box="[802,963,850,876]" pageId="91" pageNumber="92" pagination="89 - 93" refId="ref56705" refString="Morris R. 2011 c. Begonia aborensis Dunn. The Begonian 78: 89 - 93." type="journal article" year="2011">Morris 2011c</bibRefCitation>
, 
<bibRefCitation id="EFAF65882F36FF8C0191FCE3E4162D31" author="Morris R." box="[974,1052,850,877]" pageId="91" pageNumber="92" pagination="146 - 149" refId="ref56807" refString="Morris R. 2012 b. Begonia silletensis (A. DC.) C. B. Clarke. The Begonian 79: 146 - 149." type="journal article" year="2012">2012b</bibRefCitation>
), however, close examination of specimens cited in 
<bibRefCitation id="EFAF65882F36FF8C03E4FCC4E2CD2DD3" author="Tebbitt M. C. &amp; Guan K. Y." box="[443,711,885,911]" pageId="91" pageNumber="92" pagination="133 - 136" refId="ref57656" refString="Tebbitt M. C. &amp; Guan K. Y. 2002. Emended circumscription of Begonia silletensis (Begoniaceae) and description of a new subspecies from Yunnan, China. Novon 12: 133 - 136. https: // doi. org / 10.2307 / 3393252" type="journal article" year="2002">Tebbitt &amp; Guan (2002)</bibRefCitation>
reveal some with short sparse pubescence, and one (
<emphasis id="B94AC46B2F36FF8C076DFCC7E57C2DD3" box="[1330,1398,886,911]" italics="true" pageId="91" pageNumber="92">Mann</emphasis>
s.n.) with 
<quantity id="4CC6B59C2F36FF8C0370FC29E17B2DEF" box="[303,369,920,947]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="91" pageNumber="92" unit="mm" value="5.0">5 mm</quantity>
long hairs we consider to belong to 
<taxonomicName id="4C3E63FA2F36FF8C0153FC28E3962DEE" authorityName="Dunn (Dunn" authorityYear="1920" box="[780,924,920,946]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="91" pageNumber="92" phylum="Tracheophyta" rank="species" species="aborensis">
<emphasis id="B94AC46B2F36FF8C0153FC28E3962DEE" box="[780,924,920,946]" italics="true" pageId="91" pageNumber="92">B. aborensis</emphasis>
</taxonomicName>
. After consulting all available specimens including two sheets of type material and the Kew specimen (K000634631) designated as the 
<typeStatus id="5485A6DB2F36FF8C075BFC0AE5792D89" box="[1284,1395,955,981]" pageId="91" pageNumber="92" type="lectotype">lectotype</typeStatus>
, it is clear that 
<taxonomicName id="4C3E63FA2F36FF8C0338FC6EE1F62DA4" authorityName="C. B. Clarke" authorityYear="1879" baseAuthorityName="A. DC." box="[359,508,990,1016]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="91" pageNumber="92" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F36FF8C0338FC6EE1F62DA4" box="[359,508,990,1016]" italics="true" pageId="91" pageNumber="92">B. silletensis</emphasis>
</taxonomicName>
has soft short tomentose hairs along the veins on the lamina underside and petiole. The upper surface of the leaf looks glabrous at a glance, but with a hand lens or microscope small, sparsely spread hairs are often visible especially near the petiole attachment and along the veins. The same short tomentose pubescence is also found on the young ovaries but lost as the fruit ripens. The most reliable character for separating the two species is the longer peduncle in 
<taxonomicName id="4C3E63FA2F36FF8C0628FBDDE5042AD9" authorityName="C. B. Clarke" authorityYear="1879" baseAuthorityName="A. DC." box="[1143,1294,1131,1157]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="91" pageNumber="92" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F36FF8C0628FBDDE5042AD9" box="[1143,1294,1131,1157]" italics="true" pageId="91" pageNumber="92">B. silletensis</emphasis>
</taxonomicName>
, and the longer, red indumentum on 
<taxonomicName id="4C3E63FA2F31FF8B0058FEBDE2902F79" authorityName="Dunn (Dunn" authorityYear="1920" box="[519,666,267,293]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="species" species="aborensis">
<emphasis id="B94AC46B2F31FF8B0058FEBDE2902F79" box="[519,666,267,293]" italics="true" pageId="92" pageNumber="93">B. aborensis</emphasis>
</taxonomicName>
. In this account it has been decided to leave them as separate but very closely related species. Further studies on good flowering material and possibly a genetic analysis would be useful to give insight into whether they could be considered as a single, variable taxon. The plant photographed in 
<bibRefCitation id="EFAF65882F31FF8B0019FEC4E2FC2FD3" author="Morris R." box="[582,758,373,399]" pageId="92" pageNumber="93" pagination="89 - 93" refId="ref56705" refString="Morris R. 2011 c. Begonia aborensis Dunn. The Begonian 78: 89 - 93." type="journal article" year="2011">Morris (2011c)</bibRefCitation>
identified as 
<taxonomicName id="4C3E63FA2F31FF8B01CEFEC7E4282FD3" authorityName="Dunn (Dunn" authorityYear="1920" box="[913,1058,373,399]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="species" species="aborensis">
<emphasis id="B94AC46B2F31FF8B01CEFEC7E4282FD3" box="[913,1058,373,399]" italics="true" pageId="92" pageNumber="93">B. aborensis</emphasis>
</taxonomicName>
is referable to our concept of 
<taxonomicName id="4C3E63FA2F31FF8B02E2FE28E1582FEE" authorityName="C. B. Clarke" authorityYear="1879" baseAuthorityName="A. DC." box="[189,338,408,434]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F31FF8B02E2FE28E1582FEE" box="[189,338,408,434]" italics="true" pageId="92" pageNumber="93">B. silletensis</emphasis>
</taxonomicName>
considering its very short indumentum and distinct peduncle.
</paragraph>
<caption id="DF4148F12F36FF8C02E2F87BE48329B9" ID-DOI="http://doi.org/10.5281/zenodo.3787131" ID-Zenodo-Dep="3787131" box="[189,1161,1994,2021]" httpUri="https://zenodo.org/record/3787131/files/figure.png" pageId="91" pageNumber="92" startId="91.[189,232,1994,2020]" targetBox="[424,1163,1222,1959]" targetPageId="91">
<paragraph id="8B8118792F36FF8C02E2F87BE48329B9" blockId="91.[189,1161,1994,2021]" box="[189,1161,1994,2021]" pageId="91" pageNumber="92">
<emphasis id="B94AC46B2F36FF8C02E2F87BE11029B8" bold="true" box="[189,282,1994,2020]" pageId="91" pageNumber="92">Fig. 58.</emphasis>
Map showing the location of 
<taxonomicName id="4C3E63FA2F36FF8C0026F87AE30429B9" authorityName="C. B. Clarke" authorityYear="1879" baseAuthorityName="A. DC." box="[633,782,1995,2021]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="91" pageNumber="92" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F36FF8C0026F87AE30429B9" box="[633,782,1995,2021]" italics="true" pageId="91" pageNumber="92">B. silletensis</emphasis>
</taxonomicName>
(A.DC.) C.B.Clarke specimens.
</paragraph>
</caption>
<paragraph id="8B8118792F31FF8B02E2FE68E4D62C01" blockId="92.[189,1399,473,606]" pageId="92" pageNumber="93">
<bibRefCitation id="EFAF65882F31FF8B02E2FE68E17B2FA8" author="Morris R." box="[189,369,473,500]" pageId="92" pageNumber="93" pagination="146 - 149" refId="ref56807" refString="Morris R. 2012 b. Begonia silletensis (A. DC.) C. B. Clarke. The Begonian 79: 146 - 149." type="journal article" year="2012">Morris (2012b)</bibRefCitation>
shows photographs of a very large glabrous 
<taxonomicName id="4C3E63FA2F31FF8B01DEFE6BE3E92FAF" box="[897,995,474,499]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="genus">
<emphasis id="B94AC46B2F31FF8B01DEFE6BE3E92FAF" box="[897,995,474,499]" italics="true" pageId="92" pageNumber="93">Begonia</emphasis>
</taxonomicName>
from Arunachal-Pradesh, referred to as 
<taxonomicName id="4C3E63FA2F31FF8B035FFE4FE19D2C4B" authorityName="C. B. Clarke" authorityYear="1879" baseAuthorityName="A. DC." box="[256,407,509,535]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F31FF8B035FFE4FE19D2C4B" box="[256,407,509,535]" italics="true" pageId="92" pageNumber="93">B. silletensis</emphasis>
</taxonomicName>
. The plant has 5-locular fruit and flowers with 5–6 styles, and very large solitary female flowers over 
<quantity id="4CC6B59C2F31FF8B03F5FD91E1F82C67" box="[426,498,544,571]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="92" pageNumber="93" unit="cm" value="10.0">10 cm</quantity>
diameter with up to 10 fringed tepals. This differs considerably from the 
<typeStatus id="5485A6DB2F31FF8B071BFD90E57D2C67" box="[1348,1399,545,571]" pageId="92" pageNumber="93">type</typeStatus>
of 
<taxonomicName id="4C3E63FA2F31FF8B0282FDF5E17B2C01" authorityName="C. B. Clarke" authorityYear="1879" baseAuthorityName="A. DC." box="[221,369,579,605]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="species" species="silletensis">
<emphasis id="B94AC46B2F31FF8B0282FDF5E17B2C01" box="[221,369,579,605]" italics="true" pageId="92" pageNumber="93">B. silletensis</emphasis>
</taxonomicName>
and raises the possibility that the plant in the photograph is a new species.
</paragraph>
<paragraph id="8B8118792F31FF8B02E2FD34E1602CBA" blockId="92.[189,1399,645,742]" pageId="92" pageNumber="93">
There is variety of this species, 
<taxonomicName id="4C3E63FA2F31FF8B006BFD34E10E2C9E" authority="Tebbitt &amp; K. Y. Guan (Tebbitt &amp; Guan 2002)" authorityName="Tebbitt &amp; K. Y. Guan (Tebbitt &amp; Guan" authorityYear="2002" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="variety" species="silletensis" variety="mengyangensis">
<emphasis id="B94AC46B2F31FF8B006BFD34E2C32CC3" box="[564,713,645,671]" italics="true" pageId="92" pageNumber="93">B. silletensis</emphasis>
var. 
<emphasis id="B94AC46B2F31FF8B015BFD34E3BD2CC2" box="[772,951,645,670]" italics="true" pageId="92" pageNumber="93">mengyangensis</emphasis>
Tebbitt &amp; K.Y.Guan (
<bibRefCitation id="EFAF65882F31FF8B06E3FD34E0F62C9E" author="Tebbitt M. C. &amp; Guan K. Y." pageId="92" pageNumber="93" pagination="133 - 136" refId="ref57656" refString="Tebbitt M. C. &amp; Guan K. Y. 2002. Emended circumscription of Begonia silletensis (Begoniaceae) and description of a new subspecies from Yunnan, China. Novon 12: 133 - 136. https: // doi. org / 10.2307 / 3393252" type="journal article" year="2002">Tebbitt &amp; Guan 2002</bibRefCitation>
)
</taxonomicName>
, found in 
<collectingRegion id="49FAD69B2F31FF8B032CFD19E1DA2C9E" box="[371,464,680,706]" country="China" name="Yunnan" pageId="92" pageNumber="93">Yunnan</collectingRegion>
, 
<collectingCountry id="F32958E92F31FF8B0386FD19E2292C9E" box="[473,547,680,706]" name="China" pageId="92" pageNumber="93">China</collectingCountry>
.All specimens viewed for this account are of the 
<typeStatus id="5485A6DB2F31FF8B0609FD19E4832C9E" box="[1110,1161,680,706]" pageId="92" pageNumber="93">type</typeStatus>
variety 
<taxonomicName id="4C3E63FA2F31FF8B06BBFD18E1692CB9" authority="var. silletensis" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="92" pageNumber="93" phylum="Tracheophyta" rank="variety" species="silletensis" variety="silletensis">
<emphasis id="B94AC46B2F31FF8B06BBFD18E57D2C9E" box="[1252,1399,680,706]" italics="true" pageId="92" pageNumber="93">B. silletensis</emphasis>
var. 
<emphasis id="B94AC46B2F31FF8B02B0FD7AE1692CB9" box="[239,355,715,741]" italics="true" pageId="92" pageNumber="93">silletensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>