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157 lines
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<document id="5221169DC41BC2645BD2622B55B12BAB" ID-CLB-Dataset="300009" ID-DOI="10.32225/bkpmnh.2021.50_55" ID-GBIF-Dataset="70224806-ca38-47ca-bd8a-63da3ac532f9" ID-ISSN="2189-6720" ID-Zenodo-Dep="12774639" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1721260925551" checkinUser="felipe" docAuthor="Watanabe, Kyohei" docDate="2021" docId="03F987931B66A730FC0260E4FE8677D0" docLanguage="en" docName="BullKanagawaPrefectMus.50.55-136.pdf" docOrigin="Bulletin of the Kanagawa Prefectural Museum (Natural Science) 50" docSource="https://doi.org/10.32225/bkpmnh.2021.50_55" docStyle="DocumentStyle:020B98DC1285EBBD02620724ECB44EDE.1:BullKanagawaPrefectMus.2013-.journal_article.english" docStyleId="020B98DC1285EBBD02620724ECB44EDE" docStyleName="BullKanagawaPrefectMus.2013-.journal_article.english" docStyleVersion="1" docTitle="Ethelurgus Forster 1869" docType="treatment" docVersion="5" lastPageNumber="91" masterDocId="FFC0FFEB1B45A714FFCA6633FFD77447" masterDocTitle="Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species" masterLastPageNumber="136" masterPageNumber="55" pageNumber="90" updateTime="1721329445643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="21E9899CD3D9E8B79AD86662E73BFDAD">Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species</mods:title>
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<mods:namePart id="435D96F32415D14900011E5FB7C24C5C">Watanabe, Kyohei</mods:namePart>
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<mods:affiliation id="EB3183F5A6EFD86B17DFEA09F48E7EF2">Kanagawa Prefectural Museum of Natural History, 499 Iryuda, Odawara, Kanagawa 250 – 0031, Japan</mods:affiliation>
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<mods:nameIdentifier id="6BF500E099A52775BDFB5EA818B0C947" type="email">watanabe-k@nh‛kanagawa-museum‛jp</mods:nameIdentifier>
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<mods:title id="21A7D965ADF5CE1A5D59477C9D015F97">Bulletin of the Kanagawa Prefectural Museum (Natural Science)</mods:title>
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<mods:date id="2688D11A49B3BBC50AB386F5E355C5BF">2021</mods:date>
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<mods:number id="AD375ECA93BABE5B9F484660ABE36C8E">2021-03-30</mods:number>
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<treatment id="03F987931B66A730FC0260E4FE8677D0" ID-DOI="http://doi.org/10.5281/zenodo.12774791" ID-GBIF-Taxon="235610840" ID-Zenodo-Dep="12774791" LSID="urn:lsid:plazi:treatment:03F987931B66A730FC0260E4FE8677D0" httpUri="http://treatment.plazi.org/id/03F987931B66A730FC0260E4FE8677D0" lastPageId="36" lastPageNumber="91" pageId="35" pageNumber="90">
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<subSubSection id="C34A650E1B66A737FC0260E4FB0C72B7" box="[968,1243,1751,1776]" pageId="35" pageNumber="90" type="nomenclature">
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<paragraph id="8BEF36851B66A737FC0260E4FB0C72B7" blockId="35.[872,1338,1751,1815]" box="[968,1243,1751,1776]" pageId="35" pageNumber="90">
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<heading id="D0A781E91B66A737FC0260E4FB0C72B7" box="[968,1243,1751,1776]" centered="true" fontSize="10" level="2" pageId="35" pageNumber="90" reason="4">
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<emphasis id="B924EA971B66A737FC0260E4FB0C72B7" bold="true" box="[968,1243,1751,1776]" pageId="35" pageNumber="90">
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<taxonomicName id="4C504D061B66A737FC0260E4FB9272A8" ID-CoL="62RFV" ID-ENA="2886593" authority="Forster, 1869" authorityName="Forster" authorityYear="1869" box="[968,1093,1751,1775]" class="Insecta" family="Ichneumonidae" genus="Ethelurgus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="90" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA971B66A737FC0260E4FB9272A8" bold="true" box="[968,1093,1751,1775]" italics="true" pageId="35" pageNumber="90">Ethelurgus</emphasis>
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</taxonomicName>
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genus group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A650E1B66A737FCA260CDFAED7350" box="[872,1338,1790,1815]" pageId="35" pageNumber="90" type="reference_group">
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<paragraph id="8BEF36851B66A737FCA260CDFAED7350" blockId="35.[872,1338,1751,1815]" box="[872,1338,1790,1815]" pageId="35" pageNumber="90">
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(subtribe
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<taxonomicName id="4C504D061B66A737FC0460CDFAED7350" authority="sensu Townes (1970))" authorityName="Townes" authorityYear="1970" box="[974,1338,1790,1815]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="90" phylum="Arthropoda" rank="subTribe" subTribe="Ethelurgina">
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Ethelurgina sensu
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<bibRefCitation id="EFC14B741B66A737FB5860CCFAED7350" author="Townes, H." box="[1170,1338,1791,1815]" pageId="35" pageNumber="90" pagination="1 - 537" refId="ref58785" refString="Townes, H., 1970. The genera of Ichneumonidae, Part 2. Memoirs of the American Entomological Institute, 12: 1 - 537." type="journal article" year="1970">Townes (1970))</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A650E1B66A737FC8F617EFB5D739E" pageId="35" pageNumber="90" type="description">
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<paragraph id="8BEF36851B66A737FC8F617EFB5D739E" blockId="35.[813,1398,1868,2009]" pageId="35" pageNumber="90">
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Two genera,
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<taxonomicName id="4C504D061B66A737FC00617FFB127323" authority="Forster, 1869" authorityName="Forster" authorityYear="1869" box="[970,1221,1868,1892]" class="Insecta" family="Ichneumonidae" genus="Ethelurgus" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="90" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA971B66A737FC00617FFBE07323" box="[970,1079,1868,1892]" italics="true" pageId="35" pageNumber="90">Ethelurgus</emphasis>
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<bibRefCitation id="EFC14B741B66A737FBF5617EFB127323" author="Forster, A." box="[1087,1221,1868,1892]" pageId="35" pageNumber="90" pagination="135 - 221" refId="ref56071" refString="Forster, A., 1869. Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 25: 135 - 221." type="journal article" year="1869">Förster, 1869</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C504D061B66A737FB33617FFC6873CC" authority="Forster, 1869" authorityName="Forster" authorityYear="1869" class="Insecta" family="Ichneumonidae" genus="Rhembobius" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="90" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA971B66A737FB33617FFAA17323" box="[1273,1398,1868,1892]" italics="true" pageId="35" pageNumber="90">Rhembobius</emphasis>
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<bibRefCitation id="EFC14B741B66A737FCE76147FC6873CC" author="Forster, A." box="[813,959,1907,1931]" pageId="35" pageNumber="90" pagination="135 - 221" refId="ref56071" refString="Forster, A., 1869. Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 25: 135 - 221." type="journal article" year="1869">Förster, 1869</bibRefCitation>
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</taxonomicName>
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, have been recorded from
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<collectingCountry id="F34776151B66A737FB2F6147FAF273CC" box="[1253,1317,1908,1931]" name="Japan" pageId="35" pageNumber="90">Japan</collectingCountry>
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. In this study, I review both genera. The Japanese genera can be distinguished by the following key.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A650E1B61A730FF0966A8FCD674F4" box="[195,769,155,179]" pageId="36" pageNumber="91" type="multiple">
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<paragraph id="8BEF36851B61A730FF0966A8FCD674F4" blockId="36.[195,769,155,179]" box="[195,769,155,179]" pageId="36" pageNumber="91">
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<heading id="D0A781E91B61A730FF0966A8FCD674F4" box="[195,769,155,179]" centered="true" fontSize="36" level="2" pageId="36" pageNumber="91" reason="5">
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<emphasis id="B924EA971B61A730FF0966A8FCD674F4" bold="true" box="[195,769,155,179]" pageId="36" pageNumber="91">
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Key to Japanese genera of
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<taxonomicName id="4C504D061B61A730FE3966A8FDB974F4" ID-CoL="62RFV" ID-ENA="2886593" authorityName="Forster" authorityYear="1869" box="[499,622,155,179]" class="Insecta" family="Ichneumonidae" genus="Ethelurgus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="91" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA971B61A730FE3966A8FDB974F4" bold="true" box="[499,622,155,179]" italics="true" pageId="36" pageNumber="91">Ethelurgus</emphasis>
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</taxonomicName>
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genus group
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34A650E1B61A730FF7766DAFCD0767F" pageId="36" pageNumber="91" type="key">
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<key id="FFC161B41B61A730FF7766DAFCD0767F" pageId="36" pageNumber="91">
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<keyStep id="30A42E201B61A730FF7766DAFCD0767F" pageId="36" pageNumber="91">
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<paragraph id="8BEF36851B61A730FF7766DAFCD175DA" blockId="36.[189,775,233,568]" pageId="36" pageNumber="91">
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<keyLead id="30A195B01B61A730FF7766DAFCD175DA" pageId="36" pageNumber="91">
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1. Vein 2m-cu vertical (
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<figureCitation id="136B2A001B61A730FE0466DAFDE07546" box="[462,567,233,257]" captionStart="Fig" captionStartId="39.[151,184,2008,2030]" captionTargetBox="[151,1398,1321,1987]" captionTargetId="figure-409@39.[151,1398,1321,1987]" captionTargetPageId="39" captionText="Fig. 32. Rhembobius perscrutator (Thunberg,1822), KPM-NK 81957 (A, C, D) and 81961 (C), females from Japan ―A, lateral habitus; B, head, frontal view; C, fore wing; D, propodeum, T I, and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774707" httpUri="https://zenodo.org/record/12774707/files/figure.png" pageId="36" pageNumber="91">Fig. 32C</figureCitation>
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). Propodeum with strong, projecting apophyses (
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<figureCitation id="136B2A001B61A730FDFA6723FD14756F" box="[560,707,272,296]" captionStart="Fig" captionStartId="39.[151,184,2008,2030]" captionTargetBox="[151,1398,1321,1987]" captionTargetId="figure-409@39.[151,1398,1321,1987]" captionTargetPageId="39" captionText="Fig. 32. Rhembobius perscrutator (Thunberg,1822), KPM-NK 81957 (A, C, D) and 81961 (C), females from Japan ―A, lateral habitus; B, head, frontal view; C, fore wing; D, propodeum, T I, and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774707" httpUri="https://zenodo.org/record/12774707/files/figure.png" pageId="36" pageNumber="91">Figs 32A, D</figureCitation>
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). T II largely smooth except for longitudinally striate basally in Japanese species (
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<figureCitation id="136B2A001B61A730FE36676DFD897531" box="[508,606,350,374]" captionStart="Fig" captionStartId="39.[151,184,2008,2030]" captionTargetBox="[151,1398,1321,1987]" captionTargetId="figure-409@39.[151,1398,1321,1987]" captionTargetPageId="39" captionText="Fig. 32. Rhembobius perscrutator (Thunberg,1822), KPM-NK 81957 (A, C, D) and 81961 (C), females from Japan ―A, lateral habitus; B, head, frontal view; C, fore wing; D, propodeum, T I, and T II, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774707" httpUri="https://zenodo.org/record/12774707/files/figure.png" pageId="36" pageNumber="91">Fig. 32D</figureCitation>
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). ................................................
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<taxonomicName id="4C504D061B61A730FE3F67B6FD2B75DA" ID-CoL="78H7" authority="Forster, 1869" authorityName="Forster" authorityYear="1869" box="[501,764,389,413]" class="Insecta" family="Ichneumonidae" genus="Rhembobius" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="91" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA971B61A730FE3F67B6FDA575DA" box="[501,626,389,413]" italics="true" pageId="36" pageNumber="91">Rhembobius</emphasis>
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<bibRefCitation id="EFC14B741B61A730FDBD67B6FD2B75DA" author="Forster, A." box="[631,764,389,413]" pageId="36" pageNumber="91" pagination="135 - 221" refId="ref56071" refString="Forster, A., 1869. Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 25: 135 - 221." type="journal article" year="1869">Förster, 1869</bibRefCitation>
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</taxonomicName>
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*
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</keyLead>
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</paragraph>
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<paragraph id="8BEF36851B61A730FF77679FFCD0767F" blockId="36.[189,775,233,568]" pageId="36" pageNumber="91">
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<keyLead id="30A195B01B61A730FF77679FFCD0767F" pageId="36" pageNumber="91">
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-. Vein 2m-cu inclivous. Propodeum without distinct apophyses (
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<figureCitation id="136B2A001B61A730FE9967E0FE6D75AC" box="[339,442,467,491]" captionStart="Fig" captionStartId="37.[151,184,2014,2036]" captionTargetBox="[151,1401,1328,1993]" captionTargetId="figure-464@37.[151,1401,1328,1993]" captionTargetPageId="37" captionText="Fig.30. Ethelurgus episyrphicola Kusigemati, 1983, KPM-NK 81913 (B, C) and 81924 (A, D), females from Japan ―A, lateral habitus;B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesosoma, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774703" httpUri="https://zenodo.org/record/12774703/files/figure.png" pageId="36" pageNumber="91">Figs 30A</figureCitation>
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,
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<figureCitation id="136B2A001B61A730FE0D67E0FE2E75AC" box="[455,505,467,491]" captionStart="Fig" captionStartId="38.[190,223,2014,2036]" captionTargetBox="[189,1436,1388,1992]" captionTargetId="figure-440@38.[189,1436,1388,1992]" captionTargetPageId="38" captionText="Fig. 31. Ethelurgus sodalis kumatai Kusigemati, 1983, KPM-NK 81947 (C) and 81950 (A, B, D), females from Japan ―A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesosoma, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774705" httpUri="https://zenodo.org/record/12774705/files/figure.png" pageId="36" pageNumber="91">31A</figureCitation>
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). T II largely punctate in Japanese species. .................................................
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<taxonomicName id="4C504D061B61A730FE336413FD38767F" ID-CoL="62RFV" authority="Forster, 1869" authorityName="Forster" authorityYear="1869" box="[505,751,544,568]" class="Insecta" family="Ichneumonidae" genus="Ethelurgus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="91" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA971B61A730FE336413FDB1767F" box="[505,614,544,568]" italics="true" pageId="36" pageNumber="91">Ethelurgus</emphasis>
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<bibRefCitation id="EFC14B741B61A730FDA16412FD38767F" author="Forster, A." box="[619,751,544,568]" pageId="36" pageNumber="91" pagination="135 - 221" refId="ref56071" refString="Forster, A., 1869. Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 25: 135 - 221." type="journal article" year="1869">Förster, 1869</bibRefCitation>
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</taxonomicName>
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**
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</keyLead>
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</paragraph>
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</keyStep>
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</key>
|
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</subSubSection>
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<subSubSection id="C34A650E1B61A730FF1F645CFE8677D0" pageId="36" pageNumber="91" type="description">
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<paragraph id="8BEF36851B61A730FF1F645CFDB876BC" blockId="36.[189,775,622,919]" pageId="36" pageNumber="91">
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* This genus is sometimes misidentified with
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<taxonomicName id="4C504D061B61A730FD5C645DFE8C76EA" ID-CoL="6SGJ" authority="Townes, 1962" authorityName="Townes" authorityYear="1962" class="Insecta" family="Ichneumonidae" genus="Pleolophus" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="91" phylum="Arthropoda" rank="genus">
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<emphasis id="B924EA971B61A730FD5C645DFCD176C1" box="[662,774,622,646]" italics="true" pageId="36" pageNumber="91">Pleolophus</emphasis>
|
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Townes, 1962
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</taxonomicName>
|
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(Aptesini Smith & Shenfelt, 1955 of
|
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<taxonomicName id="4C504D061B61A730FF77648FFEFF7693" box="[189,296,700,724]" class="Insecta" family="Ichneumonidae" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="55" phylum="Arthropoda" rank="subFamily" subFamily="Cryptinae">Cryptinae</taxonomicName>
|
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) because vein 2m-cu vertical. Users should carefully check the character states of them.
|
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</paragraph>
|
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<paragraph id="8BEF36851B61A730FF1F6539FE8677D0" blockId="36.[189,775,622,919]" pageId="36" pageNumber="91">
|
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** This genus is sometimes misidentified with
|
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|
<taxonomicName id="4C504D061B61A730FD676539FE9F770E" ID-CoL="645MG" authority="Forster, 1869" authorityName="Forster" authorityYear="1869" class="Insecta" family="Ichneumonidae" genus="Stibeutes" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="91" phylum="Arthropoda" rank="genus">
|
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|
<emphasis id="B924EA971B61A730FD676539FCD07765" box="[685,775,778,802]" italics="true" pageId="36" pageNumber="91">Stibeutes</emphasis>
|
|||
|
|
<bibRefCitation id="EFC14B741B61A730FF776501FE9F770E" author="Forster, A." box="[189,328,817,841]" pageId="36" pageNumber="91" pagination="135 - 221" refId="ref56071" refString="Forster, A., 1869. Synopsis der Familien und Gattungen der Ichneumonen. Verhandlungen des Naturhistorischen Vereins der Preussischen Rheinlande und Westfalens, 25: 135 - 221." type="journal article" year="1869">Förster, 1869</bibRefCitation>
|
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|
|
</taxonomicName>
|
|||
|
|
(
|
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|
|
<taxonomicName id="4C504D061B61A730FE936502FE0C770E" ID-CoL="6PGC" authorityName="Gravenhorst" authorityYear="1829" box="[345,475,817,841]" class="Insecta" family="Ichneumonidae" genus="Phygadeuon" kingdom="Animalia" order="Hymenoptera" pageId="36" pageNumber="91" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B924EA971B61A730FE936502FE0C770E" box="[345,475,817,841]" italics="true" pageId="36" pageNumber="91">Phygadeuon</emphasis>
|
|||
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</taxonomicName>
|
|||
|
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genus group). Users should carefully check the character states of clypeus, propodeum, and ovipositor.
|
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|
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</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
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</document>
|