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<document id="05D158935A75DB0D62F9BC0F804B5157" ID-DOI="10.11646/zootaxa.4803.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3909047" ID-ZooBank="0C0F5754-9530-4072-AF45-BDFE4A8A2B57" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593158336852" checkinUser="plazi" docAuthor="Meyer, Marc De, Goergen, Georg &amp; Jordaens, Kurt" docDate="2020" docId="03F94D2CFFA7FFE60EAD3785FF76FA4C" docLanguage="en" docName="zootaxa.4803.2.1.pdf" docOrigin="Zootaxa 4803 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Phytomia austeni Meyer &amp; Goergen &amp; Jordaens 2020, sp. nov." docType="treatment" docUuid="E7389C49-209A-453B-8913-12A1DDA0ABA6" docUuidSource="ZooBank" docVersion="3" lastPageNumber="218" masterDocId="FFC03554FFB7FFF70E3A3578FFACFFEA" masterDocTitle="Taxonomic revision of the Afrotropical Phytomia Guérin-Méneville (Diptera: Syrphidae)" masterLastPageNumber="250" masterPageNumber="201" pageNumber="217" updateTime="1700277132632" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="6F159DD77E50E7A1615E72CF8A73779A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6376583A5EEB0E4F62CDB561A6FB4F3C">
<mods:title id="E6A9CF73528F70CEC7B801EDC7DEF5DF">Taxonomic revision of the Afrotropical Phytomia Guérin-Méneville (Diptera: Syrphidae)</mods:title>
</mods:titleInfo>
<mods:name id="62AE649107830E9152C80D23F8CF6CF8" type="personal">
<mods:role id="4ECAFD80D7A338BCD44011EB65945D6C">
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<mods:namePart id="E200AF4B3320F95F49FF57926A0C1A01">Meyer, Marc De</mods:namePart>
<mods:affiliation id="6D6B4FFCD73522C060209369F8697230">Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B 3080 Tervuren Belgium.</mods:affiliation>
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<mods:namePart id="313EDB5DE31A3F4EE1DAFD74A3E96971">Goergen, Georg</mods:namePart>
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<mods:name id="7852DC5F0D8A84F57EE088922246438B" type="personal">
<mods:role id="C9FE285ED3B7CB4514CBE796FF34AADE">
<mods:roleTerm id="0D34593D65CB60B4C5326BB43B7E3B6D">Author</mods:roleTerm>
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<mods:namePart id="8C8C0C9F26F72B00B10D629F7281B1B7">Jordaens, Kurt</mods:namePart>
<mods:affiliation id="4E79276CBF38486DFBE2EFD06614CC9E">Royal Museum for Central Africa, Invertebrates Section and JEMU, Leuvensesteenweg 13, B 3080 Tervuren Belgium. &amp; kurt. jordaens @ africamuseum. be; https: // orcid. org / 0000 - 0003 - 4321 - 5944</mods:affiliation>
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<mods:title id="296B32D50B86C6C0525EBB70DD377AB9">Zootaxa</mods:title>
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<mods:part id="F7E27166ED54E3F28395191E796E3509">
<mods:date id="3C0AE5DEA0D908F68DD67507EAD82845">2020</mods:date>
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<mods:number id="DA14DC8A1EE47CB73938AD1D3D0B8A78">2020-06-26</mods:number>
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<mods:identifier id="24DB7E2626FB20B2BA1128A09ACF5FA6" type="DOI">10.11646/zootaxa.4803.2.1</mods:identifier>
<mods:identifier id="90561057F3BB132A98002A7335C187A9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="006CF68CDA0BC124ECE643B439062FB7" type="Zenodo-Dep">3909047</mods:identifier>
<mods:identifier id="789B475E0DA8989E63C00D51ECB51B46" type="ZooBank">0C0F5754-9530-4072-AF45-BDFE4A8A2B57</mods:identifier>
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<treatment id="03F94D2CFFA7FFE60EAD3785FF76FA4C" LSID="urn:lsid:zoobank.org:act:E7389C49-209A-453B-8913-12A1DDA0ABA6" httpUri="http://treatment.plazi.org/id/03F94D2CFFA7FFE60EAD3785FF76FA4C" lastPageId="17" lastPageNumber="218" pageId="16" pageNumber="217">
<subSubSection id="C34AAFB1FFA7FFE70EAD3785FE63FCFD" box="[151,463,765,792]" pageId="16" pageNumber="217" type="nomenclature">
<paragraph id="8BEFFC3AFFA7FFE70EAD3785FE63FCFD" blockId="16.[151,977,765,863]" box="[151,463,765,792]" pageId="16" pageNumber="217">
<heading id="D0A74B56FFA7FFE70EAD3785FE63FCFD" bold="true" box="[151,463,765,792]" fontSize="11" level="1" pageId="16" pageNumber="217" reason="1">
<emphasis id="B9242028FFA7FFE70EAD3785FE63FCFD" bold="true" box="[151,463,765,792]" pageId="16" pageNumber="217">
<taxonomicName id="4C5087B9FFA7FFE70EAD3785FEC6FCFD" ID-CoL="9HXYT" authority="Meyer &amp; Goergen &amp; Jordaens, 2020" authorityName="Meyer &amp; Goergen &amp; Jordaens" authorityYear="2020" box="[151,362,765,792]" class="Insecta" family="Syrphidae" genus="Phytomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="austeni" status="sp. nov.">
<emphasis id="B9242028FFA7FFE70EAD3785FEC6FCFD" bold="true" box="[151,362,765,792]" italics="true" pageId="16" pageNumber="217">Phytomia austeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2179D53FFA7FFE70F483785FE63FCFD" box="[370,463,765,791]" pageId="16" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C34AAFB1FFA7FFE70EAD3659FC7DFCB5" pageId="16" pageNumber="217" type="description">
<paragraph id="8BEFFC3AFFA7FFE70EAD3659FE6DFCD1" blockId="16.[151,977,765,863]" box="[151,449,801,827]" pageId="16" pageNumber="217">
(
<figureCitation id="136BE0BFFFA7FFE70EA53659FF44FCD1" box="[159,232,801,827]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1931,1956]" captionTargetBox="[162,1426,181,1905]" captionTargetId="figure@4.[161,1426,181,1905]" captionTargetPageId="4" captionText="FIGURES 1–10. Phytomia spp., habitus, lateral view. 1. P. bullata (Loew) ♂ 2. P. aurigera Bezzi ♂ 3. P. kroeberi (Bezzi) ♂ 4. P. serena (Curran) ♀ 5. P. bulligera (Austen) ♂ 6. P. austeni sp. nov. ♂ 7. P. bezzii Curran ♂ 8. P. memnon sp. nov. ♂ 9. P. pubipennis Bezzi ♂ 10. P. natalensis (Macquart) ♂" figureDoi="http://doi.org/10.5281/zenodo.3909049" httpUri="https://zenodo.org/record/3909049/files/figure.png" pageId="16" pageNumber="217">Figs 6</figureCitation>
,
<figureCitation id="136BE0BFFFA7FFE70ECE3659FEBEFCD1" box="[244,274,801,827]" captionStart="FIGURES 29–36" captionStartId="13.[151,264,1945,1970]" captionTargetBox="[242,1360,181,1919]" captionTargetId="figure@13.[227,1360,181,1919]" captionTargetPageId="13" captionText="FIGURES 29–36. Figs 29–34. Phytomia spp., hind leg, anterior view. 29. P. bullata (Loew) ♂ 30. P. serena (Curran) ♀ 31. P. bulligera (Austen) ♂ 32. P. austeni sp. nov. ♂ 33. P. bezzii Curran ♂ 34. P. memnon sp. nov. ♂ Figs 35–36: Phytomia spp., scutellum, dorsal view. 35. P. melas (Bezzi) ♂ 36. P. poensis (Bezzi) ♀" figureDoi="http://doi.org/10.5281/zenodo.3909055" httpUri="https://zenodo.org/record/3909055/files/figure.png" pageId="16" pageNumber="217">32</figureCitation>
,
<figureCitation id="136BE0BFFFA7FFE70F243659FECAFCD1" box="[286,358,801,827]" captionStart="FIGURES 37–48" captionStartId="21.[151,264,1945,1970]" captionTargetBox="[157,1428,201,1919]" captionTargetId="figure@21.[157,1431,181,1919]" captionTargetPageId="21" captionText="FIGURES 37–48. Phytomia spp., right wing. 37. P. bullata (Loew) ♀ 38. P. aurigera Bezzi ♂ 39–40. P. kroeberi (Bezzi) ♀ 41. P. serena (Curran) ♀ 42. P. bulligera (Austen) ♀ 43. P. austeni sp. nov. ♂ 44. P. austeni sp. nov. ♀ 45. P. bezzii Curran ♂ 46. P. bezzii Curran ♀ 47. P. memnon sp. nov. ♀ 48. P. pubipennis Bezzi ♂" figureDoi="http://doi.org/10.5281/zenodo.3909057" httpUri="https://zenodo.org/record/3909057/files/figure.png" pageId="16" pageNumber="217">43–44</figureCitation>
,
<figureCitation id="136BE0BFFFA7FFE70F483659FE3CFCD1" box="[370,400,801,827]" captionStart="FIGURES 61–72" captionStartId="29.[151,264,1945,1970]" captionTargetBox="[182,1431,181,1918]" captionTargetId="figure@29.[156,1431,181,1919]" captionTargetPageId="29" captionText="FIGURES 61–72. Phytomia spp., abdomen, dorsal view. 61. P. bullata (Loew) ♂ 62. P. aurigera Bezzi ♂ 63. P. kroeberi (Bezzi) ♂ 64–65. P. kroeberi (Bezzi) ♀ 66. P. serena (Curran) ♀ 67. P. bulligera (Austen) ♂ 68. P. austeni sp. nov. ♂ 69. P. bezzii Curran ♀ 70. P. memnon sp. nov. ♀ 71. P. pubipennis Bezzi ♂ 72. P. natalensis (Macquart) (M)." figureDoi="http://doi.org/10.5281/zenodo.3909061" httpUri="https://zenodo.org/record/3909061/files/figure.png" pageId="16" pageNumber="217">68</figureCitation>
,
<figureCitation id="136BE0BFFFA7FFE70FA63659FE15FCD1" box="[412,441,801,827]" captionStart="FIGURES 85–100" captionStartId="37.[151,264,1727,1752]" captionTargetBox="[151,1436,381,1702]" captionTargetId="figure@37.[151,1436,381,1702]" captionTargetPageId="37" captionText="FIGURES 85–100. Phytomia spp., male terminalia, dorsal view. 85. P. bullata (Loew). 86. P. aurigera Bezzi. 87. P. kroeberi (Bezzi). 88. P. bulligera (Austen). 89. P. austeni sp. nov. 90. P. bezzii Curran. 91. P. memnon sp. nov. 92. P. pubipennis Bezzi. 93. P. natalensis (Macquart). 94. P. curta (Loew). 95. P. pallida sp. nov. 96. P. erratica (Bezzi). 97. P. fucoides Bezzi. 98. P. incisa (Wiedemann). 99. P. melas (Bezzi). 100. P. poensis (Bezzi)." figureDoi="http://doi.org/10.5281/zenodo.3909065" httpUri="https://zenodo.org/record/3909065/files/figure.png" pageId="16" pageNumber="217">89</figureCitation>
)
</paragraph>
<paragraph id="8BEFFC3AFFA7FFE70EAD363CFC7DFCB5" blockId="16.[151,977,765,863]" box="[151,977,836,863]" pageId="16" pageNumber="217">
<uri id="FFC1F038FFA7FFE70EAD363CFC7DFCB5" box="[151,977,836,863]" pageId="16" pageNumber="217">
urn:lsid:zoobank.org:act:
<uuid id="FFF6C6EFFFA7FFE70F93363CFC7DFCB5" box="[425,977,836,863]" pageId="16" pageNumber="217">E7389C49-209A-453B-8913-12A1DDA0ABA6</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C34AAFB1FFA7FFE70EAD36F4FBC6FA91" pageId="16" pageNumber="217" type="materials_examined">
<paragraph id="8BEFFC3AFFA7FFE70EAD36F4FB6BFABD" blockId="16.[151,1437,908,2015]" pageId="16" pageNumber="217">
<emphasis id="B9242028FFA7FFE70EAD36F4FED6FC4D" bold="true" box="[151,378,908,935]" pageId="16" pageNumber="217">Material examined</emphasis>
.
<materialsCitation id="3B38F667FFA7FFE70FB336F5FB27FC4D" box="[393,1163,909,935]" collectingDate="2002-03" collectionCode="KMMA" collectorName="G. Goergen" country="Togo" location="Kloto" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54EB4298FFA7FFE70FB336F5FD8DFC4D" box="[393,545,909,935]" pageId="16" pageNumber="217" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D5637B3FFA7FFE70C1636F5FDEFFC4D" box="[556,579,909,935]" pageId="16" pageNumber="217" type="male">♂</specimenCount>
,
<collectingCountry id="F347BCAAFFA7FFE70C7536F5FD33FC4D" box="[591,671,909,935]" name="Togo" pageId="16" pageNumber="217">TOGO</collectingCountry>
,
<location id="8E8FAAE1FFA7FFE70C9136F5FD42FC4D" LSID="urn:lsid:plazi:treatment:03F94D2CFFA7FFE60EAD3785FF76FA4C:8E8FAAE1FFA7FFE70C9136F5FD42FC4D" box="[683,750,909,935]" country="Togo" name="Kloto" pageId="16" pageNumber="217">Kloto</location>
,
<date id="FFEEDAFAFFA7FFE70CC336F5FC29FC4D" box="[761,901,909,935]" pageId="16" pageNumber="217" value="2002-03">
<collectingDate id="EFAA2312FFA7FFE70CC336F5FC29FC4D" box="[761,901,909,935]" pageId="16" pageNumber="217" value="2002-03">March 2002</collectingDate>
</date>
,
<collectorName id="26A599ECFFA7FFE70DAA36F5FBBFFC4D" box="[912,1043,909,935]" pageId="16" pageNumber="217">G. Goergen</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70A1836F5FB2FFC4D" box="[1058,1155,909,935]" country="Belgium" httpUri="http://grbio.org/cool/p87v-9846" name="Koninklijk Museum voor Midden Afrika" pageId="16" pageNumber="217" type="Museum">KMMA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B38F667FFA7FFE70AAC36F5FE76FC21" collectingDate="2006-02" country="Benin" location="Aplaoue" pageId="16" pageNumber="217" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="54EB4298FFA7FFE70AAC36F5FA99FC4D" box="[1174,1333,909,935]" pageId="16" pageNumber="217" type="paratype">PARATYPES</typeStatus>
:
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,
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,
<location id="8E8FAAE1FFA7FFE70EFC36C9FE84FC21" LSID="urn:lsid:plazi:treatment:03F94D2CFFA7FFE60EAD3785FF76FA4C:8E8FAAE1FFA7FFE70EFC36C9FE84FC21" box="[198,296,945,971]" country="Benin" name="Aplaoue" pageId="16" pageNumber="217">Aplaoué</location>
,
<date id="FFEEDAFAFFA7FFE70F0936C9FE76FC21" box="[307,474,945,971]" pageId="16" pageNumber="217" value="2006-02">
<collectingDate id="EFAA2312FFA7FFE70F0936C9FE76FC21" box="[307,474,945,971]" pageId="16" pageNumber="217" value="2006-02">February 2006</collectingDate>
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;
<specimenCount id="9D5637B3FFA7FFE70FDF36C9FDA4FC21" box="[485,520,945,971]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, Pénéssoulou,
<date id="FFEEDAFAFFA7FFE70C9536C9FCCBFC21" box="[687,871,945,971]" pageId="16" pageNumber="217" value="2003-11">November 2003</date>
,
<materialsCitation id="3B38F667FFA7FFE70D4B36C9FBC4FC21" box="[881,1128,945,971]" collectionCode="IITA" collectorName="Goergen" country="Benin" pageId="16" pageNumber="217" specimenCount="1" typeStatus="paratype">
all G.
<collectorName id="26A599ECFFA7FFE70D8C36C9FBBAFC21" box="[950,1046,945,971]" pageId="16" pageNumber="217">Goergen</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70A1E36C9FBCCFC21" box="[1060,1120,945,971]" country="Nigeria" httpUri="http://grbio.org/cool/8h49-s777" name="International Institute of Tropical Agriculture" pageId="16" pageNumber="217">IITA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B38F667FFA7FFE70A4836C9FEE2FC05" collectingDate="2004-12" country="Ghana" location="Cape Coast" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<collectingCountry id="F347BCAAFFA7FFE70A4836C9FB76FC21" box="[1138,1242,945,971]" name="Ghana" pageId="16" pageNumber="217">GHANA</collectingCountry>
,
<specimenCount id="9D5637B3FFA7FFE70ADF36C9FAA5FC21" box="[1253,1289,945,971]" pageId="16" pageNumber="217" type="male">1♂</specimenCount>
,
<location id="8E8FAAE1FFA7FFE70B2F36C9FA34FC21" LSID="urn:lsid:plazi:treatment:03F94D2CFFA7FFE60EAD3785FF76FA4C:8E8FAAE1FFA7FFE70B2F36C9FA34FC21" box="[1301,1432,945,971]" country="Ghana" name="Cape Coast" pageId="16" pageNumber="217">Cape Coast</location>
,
<date id="FFEEDAFAFFA7FFE70EAD36ADFEE2FC05" box="[151,334,981,1007]" pageId="16" pageNumber="217" value="2004-12">
<collectingDate id="EFAA2312FFA7FFE70EAD36ADFEE2FC05" box="[151,334,981,1007]" pageId="16" pageNumber="217" value="2004-12">December 2004</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B38F667FFA7FFE70F6136ADFD98FC05" box="[347,564,981,1007]" collectionCode="IITA" collectorName="Goergen" country="Ghana" pageId="16" pageNumber="217" specimenCount="1" typeStatus="paratype">
G.
<collectorName id="26A599ECFFA7FFE70F4536ADFE73FC05" box="[383,479,981,1007]" pageId="16" pageNumber="217">Goergen</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70FCA36ADFD80FC05" box="[496,556,981,1007]" country="Nigeria" httpUri="http://grbio.org/cool/8h49-s777" name="International Institute of Tropical Agriculture" pageId="16" pageNumber="217">IITA</collectionCode>
)
</materialsCitation>
;
<specimenCount id="9D5637B3FFA7FFE70C7B36ADFDC8FC05" box="[577,612,981,1007]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, Wati Waterfalls,
<date id="FFEEDAFAFFA7FFE70D0936ADFC70FC05" box="[819,988,981,1007]" pageId="16" pageNumber="217" value="2003-02">February 2003</date>
,
<materialsCitation id="3B38F667FFA7FFE70DD336ADFB6EFC05" box="[1001,1218,981,1007]" collectionCode="IITA" collectorName="Goergen" country="Ghana" pageId="16" pageNumber="217" specimenCount="1" typeStatus="paratype">
G.
<collectorName id="26A599ECFFA7FFE70A3736ADFBC1FC05" box="[1037,1133,981,1007]" pageId="16" pageNumber="217">Goergen</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70A4436ADFB16FC05" box="[1150,1210,981,1007]" country="Nigeria" httpUri="http://grbio.org/cool/8h49-s777" name="International Institute of Tropical Agriculture" pageId="16" pageNumber="217">IITA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B38F667FFA7FFE70AF436ADFE49FBF9" collectionCode="CNC" country="Sierra Leone" location="Foxcroft" pageId="16" pageNumber="217" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<collectingCountry id="F347BCAAFFA7FFE70AF436ADFA34FC05" box="[1230,1432,981,1007]" name="Sierra Leone" pageId="16" pageNumber="217">SIERRA LEONE</collectingCountry>
,
<specimenCount id="9D5637B3FFA7FFE70EAD3681FF16FBF9" box="[151,186,1017,1043]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, 1858, J.
<location id="8E8FAAE1FFA7FFE70F163681FE20FBF9" LSID="urn:lsid:plazi:treatment:03F94D2CFFA7FFE60EAD3785FF76FA4C:8E8FAAE1FFA7FFE70F163681FE20FBF9" box="[300,396,1017,1043]" country="Sierra Leone" name="Foxcroft" pageId="16" pageNumber="217">Foxcroft</location>
(
<collectionCode id="ED4164FFFFA7FFE70FA43681FE71FBF9" box="[414,477,1017,1043]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="16" pageNumber="217">CNC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B38F667FFA7FFE70FC83681FB29FBF9" box="[498,1157,1017,1043]" collectionCode="BMNH" collectorName="Clements" country="Sierra Leone" pageId="16" pageNumber="217" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="9D5637B3FFA7FFE70FC83681FDBBFBF9" box="[498,535,1017,1043]" pageId="16" pageNumber="217" type="male">1♂</specimenCount>
,
<collectorName id="26A599ECFFA7FFE70C1F3681FD23FBF9" box="[549,655,1017,1043]" pageId="16" pageNumber="217">Clements</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70CA53681FD5AFBF9" box="[671,758,1017,1043]" pageId="16" pageNumber="217">BMNH</collectionCode>
) (both as
<typeStatus id="54EB4298FFA7FFE70D513681FC61FBF9" box="[875,973,1017,1043]" pageId="16" pageNumber="217" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="4C5087B9FFA7FFE70DCC3682FBD7FBF9" authorityName=", Austen" authorityYear="1909" box="[1014,1147,1017,1043]" class="Insecta" family="Syrphidae" genus="Megaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="bulligera">
<emphasis id="B9242028FFA7FFE70DCC3682FBD7FBF9" box="[1014,1147,1017,1043]" italics="true" pageId="16" pageNumber="217">P. bulligera</emphasis>
</taxonomicName>
)
</materialsCitation>
.
<materialsCitation id="3B38F667FFA7FFE70AA93681FA9FFBF9" box="[1171,1331,1017,1043]" country="Togo" location="Kloto" pageId="16" pageNumber="217" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F347BCAAFFA7FFE70AA93681FB4FFBF9" box="[1171,1251,1017,1043]" name="Togo" pageId="16" pageNumber="217">TOGO</collectingCountry>
,
<location id="8E8FAAE1FFA7FFE70ACA3681FA9FFBF9" LSID="urn:lsid:plazi:treatment:03F94D2CFFA7FFE60EAD3785FF76FA4C:8E8FAAE1FFA7FFE70ACA3681FA9FFBF9" box="[1264,1331,1017,1043]" country="Togo" name="Kloto" pageId="16" pageNumber="217">Kloto</location>
</materialsCitation>
,
<specimenCount id="9D5637B3FFA7FFE70B053681FACEFBF9" box="[1343,1378,1017,1043]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, Au- gust 2003;
<specimenCount id="9D5637B3FFA7FFE70F2D3165FE97FBDD" box="[279,315,1053,1079]" pageId="16" pageNumber="217" type="male">1♂</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70F733165FE4BFBDD" box="[329,487,1053,1079]" pageId="16" pageNumber="217" value="2003-10">October 2003</date>
;
<specimenCount id="9D5637B3FFA7FFE70FCE3165FDBBFBDD" box="[500,535,1053,1079]" pageId="16" pageNumber="217" type="male">3♂</specimenCount>
<specimenCount id="9D5637B3FFA7FFE70C253165FDEEFBDD" box="[543,578,1053,1079]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70C683165FD41FBDD" box="[594,749,1053,1079]" pageId="16" pageNumber="217" value="2004-01">January 2004</date>
;
<specimenCount id="9D5637B3FFA7FFE70CC33165FCB0FBDD" box="[761,796,1053,1079]" pageId="16" pageNumber="217" type="male">2♂</specimenCount>
<specimenCount id="9D5637B3FFA7FFE70D1F3165FCE5FBDD" box="[805,841,1053,1079]" pageId="16" pageNumber="217" type="female">3♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70D6D3165FBADFBDD" box="[855,1025,1053,1079]" pageId="16" pageNumber="217" value="2004-02">February 2004</date>
;
<specimenCount id="9D5637B3FFA7FFE70A373165FB9CFBDD" box="[1037,1072,1053,1079]" pageId="16" pageNumber="217" type="male">2♂</specimenCount>
<specimenCount id="9D5637B3FFA7FFE70A033165FBF1FBDD" box="[1081,1117,1053,1079]" pageId="16" pageNumber="217" type="female">4♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70A513165FB55FBDD" box="[1131,1273,1053,1079]" pageId="16" pageNumber="217" value="2004-03">March 2004</date>
;
<specimenCount id="9D5637B3FFA7FFE70B3F3165FA84FBDD" box="[1285,1320,1053,1079]" pageId="16" pageNumber="217" type="male">7♂</specimenCount>
<specimenCount id="9D5637B3FFA7FFE70B0A3165FAF9FBDD" box="[1328,1365,1053,1079]" pageId="16" pageNumber="217" type="female">2♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70B5B3165FF7FFBB1" pageId="16" pageNumber="217" value="2004-04">April 2004</date>
;
<specimenCount id="9D5637B3FFA7FFE70EDB3139FEA9FBB1" box="[225,261,1089,1115]" pageId="16" pageNumber="217" type="male">1♂</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70F2F3139FE7DFBB1" box="[277,465,1089,1115]" pageId="16" pageNumber="217" value="2004-11">November 2004</date>
;
<specimenCount id="9D5637B3FFA7FFE70FE53139FDAEFBB1" box="[479,514,1089,1115]" pageId="16" pageNumber="217" type="male">2♂</specimenCount>
<specimenCount id="9D5637B3FFA7FFE70C363139FD83FBB1" box="[524,559,1089,1115]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70C7A3139FD55FBB1" box="[576,761,1089,1115]" pageId="16" pageNumber="217" value="2004-12">December 2004</date>
;
<specimenCount id="9D5637B3FFA7FFE70D3D3139FC80FBB1" box="[775,812,1089,1115]" pageId="16" pageNumber="217" type="male">2♂</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70D013139FC49FBB1" box="[827,997,1088,1115]" pageId="16" pageNumber="217" value="2005-02">February 2005</date>
;
<specimenCount id="9D5637B3FFA7FFE70DCE3139FBBBFBB1" box="[1012,1047,1089,1115]" pageId="16" pageNumber="217" type="male">1♂</specimenCount>
<specimenCount id="9D5637B3FFA7FFE70A1B3139FBE8FBB1" box="[1057,1092,1089,1115]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70A6F3139FB4FFBB1" box="[1109,1251,1088,1115]" pageId="16" pageNumber="217" value="2005-03">March 2005</date>
;
<specimenCount id="9D5637B3FFA7FFE70AC83139FAB9FBB1" box="[1266,1301,1089,1115]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70B1C3139FF7EFB95" pageId="16" pageNumber="217" value="2005-11">November 2005</date>
;
<specimenCount id="9D5637B3FFA7FFE70EE5311DFEA9FB95" box="[223,261,1125,1151]" pageId="16" pageNumber="217" type="male">2♂</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70F2B311DFE25FB95" box="[273,393,1125,1151]" pageId="16" pageNumber="217" value="2006-05">May 2006</date>
;
<specimenCount id="9D5637B3FFA7FFE70FAE311DFE1BFB95" box="[404,439,1125,1151]" pageId="16" pageNumber="217" type="male">2♂</specimenCount>
<specimenCount id="9D5637B3FFA7FFE70FFA311DFE48FB95" box="[448,484,1125,1151]" pageId="16" pageNumber="217" type="female">4♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70FC8311DFD23FB95" box="[498,655,1125,1151]" pageId="16" pageNumber="217" value="2007-10">October 2007</date>
;
<specimenCount id="9D5637B3FFA7FFE70CA6311DFD6CFB95" box="[668,704,1125,1151]" pageId="16" pageNumber="217" type="male">1♂</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70CF4311DFCC4FB95" box="[718,872,1125,1151]" pageId="16" pageNumber="217" value="2008-01">January 2008</date>
;
<specimenCount id="9D5637B3FFA7FFE70D4F311DFC35FB95" box="[885,921,1125,1151]" pageId="16" pageNumber="217" type="male">1♂</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70D9D311DFBB5FB95" box="[935,1049,1125,1151]" pageId="16" pageNumber="217" value="2008-07">July 2008</date>
;
<specimenCount id="9D5637B3FFA7FFE70A1C311DFBE6FB95" box="[1062,1098,1125,1151]" pageId="16" pageNumber="217" type="male">1♂</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70A6C311DFB46FB95" box="[1110,1258,1124,1151]" pageId="16" pageNumber="217" value="2015-08">August 2015</date>
;
<specimenCount id="9D5637B3FFA7FFE70ACD311DFAB6FB95" box="[1271,1306,1125,1151]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
,
<date id="FFEEDAFAFFA7FFE70B13311DFF7EFB49" pageId="16" pageNumber="217" value="2016-12">December 2016</date>
,
<materialsCitation id="3B38F667FFA7FFE70EDA31F1FE73FB49" box="[224,479,1161,1187]" collectionCode="IITA" collectorName="Goergen" country="Togo" pageId="16" pageNumber="217" specimenCount="1" typeStatus="paratype">
all G.
<collectorName id="26A599ECFFA7FFE70F1031F1FE26FB49" box="[298,394,1161,1187]" pageId="16" pageNumber="217">Goergen</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70FA131F1FE7BFB49" box="[411,471,1161,1187]" country="Nigeria" httpUri="http://grbio.org/cool/8h49-s777" name="International Institute of Tropical Agriculture" pageId="16" pageNumber="217">IITA</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B38F667FFA7FFE70FD631F1FB6EFABD" collectingDate="1910-02" collectingDateMax="1960-02-03" collectingDateMin="1910-02" collectionCode="KMMA, KBIN, AMNH" collectorName="J. Vrydagh &amp; F. J. Francois &amp; P. Hulstaert &amp; Lang &amp; Chapin &amp; Manyema, R" country="Democratic Republic of the Congo" location="Associated" municipality="Bambesa" pageId="16" pageNumber="217" specimenCount="7" specimenCount-female="7" typeStatus="holotype">
<typeStatus id="54EB4298FFA7FFE70FD631F1FDF8FB49" box="[492,596,1161,1187]" pageId="16" pageNumber="217" type="holotype">Holotype</typeStatus>
deposited in KMMA,
<typeStatus id="54EB4298FFA7FFE70D6631F1FC6BFB49" box="[860,967,1161,1187]" pageId="16" pageNumber="217" type="paratype">paratypes</typeStatus>
in collections of AMNH, BMNH, CNC, IITA, and KMMA.
<location id="8E8FAAE1FFA7FFE70F4C31D5FE5DFB2D" LSID="urn:lsid:plazi:treatment:03F94D2CFFA7FFE60EAD3785FF76FA4C:8E8FAAE1FFA7FFE70F4C31D5FE5DFB2D" box="[374,497,1197,1223]" country="Democratic Republic of the Congo" municipality="Bambesa" name="Associated" pageId="16" pageNumber="217">Associated</location>
material (not included in type series; females only)
<collectingCountry id="F347BCAAFFA7FFE70A7D31D5FB07FB2D" box="[1095,1195,1197,1223]" name="Democratic Republic of the Congo" pageId="16" pageNumber="217">CONGO</collectingCountry>
(DR),
<specimenCount id="9D5637B3FFA7FFE70AC631D5FAB3FB2D" box="[1276,1311,1197,1223]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
,
<collectingMunicipality id="6B8B6640FFA7FFE70B1531D5FA35FB2D" box="[1327,1433,1197,1223]" pageId="16" pageNumber="217">Bambesa</collectingMunicipality>
,
<date id="FFEEDAFAFFA7FFE70EAD31A9FEADFB01" box="[151,257,1233,1259]" pageId="16" pageNumber="217" value="1938-10-01">
<collectingDate id="EFAA2312FFA7FFE70EAD31A9FEADFB01" box="[151,257,1233,1259]" pageId="16" pageNumber="217" value="1938-10-01">1.X.1938</collectingDate>
</date>
,
<collectorName id="26A599ECFFA7FFE70F3631A9FE2FFB01" box="[268,387,1233,1259]" pageId="16" pageNumber="217">J. Vrydagh</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70FAB31A9FE5EFB01" box="[401,498,1233,1259]" country="Belgium" httpUri="http://grbio.org/cool/p87v-9846" name="Koninklijk Museum voor Midden Afrika" pageId="16" pageNumber="217" type="Museum">KMMA</collectionCode>
);
<specimenCount id="9D5637B3FFA7FFE70C3E31A9FD8BFB01" box="[516,551,1233,1259]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, Dekese, Itunda, Kasai,
<date id="FFEEDAFAFFA7FFE70D0E31A9FC31FB01" box="[820,925,1233,1259]" pageId="16" pageNumber="217" value="1960-02-03">
<collectingDate id="EFAA2312FFA7FFE70D0E31A9FC31FB01" box="[820,925,1233,1259]" pageId="16" pageNumber="217" value="1960-02-03">3.II.1960</collectingDate>
</date>
,
<collectorName id="26A599ECFFA7FFE70D9231A9FB98FB01" box="[936,1076,1233,1259]" pageId="16" pageNumber="217">F.J. François</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70A7831A9FB20FB01" box="[1090,1164,1233,1259]" pageId="16" pageNumber="217">KBIN</collectionCode>
);
<specimenCount id="9D5637B3FFA7FFE70AA531A9FB6EFB01" box="[1183,1218,1233,1259]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, Flandria, Tshuapa,
<date id="FFEEDAFAFFA7FFE70EAD318CFEBCFAE5" box="[151,272,1268,1295]" pageId="16" pageNumber="217" value="1945-10-18">
<collectingDate id="EFAA2312FFA7FFE70EAD318CFEBCFAE5" box="[151,272,1268,1295]" pageId="16" pageNumber="217" value="1945-10-18">18.X.1945</collectingDate>
</date>
,
<collectorName id="26A599ECFFA7FFE70F24318DFE0EFAE5" box="[286,418,1269,1295]" pageId="16" pageNumber="217">P. Hulstaert</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70F89318DFDB8FAE5" box="[435,532,1269,1295]" country="Belgium" httpUri="http://grbio.org/cool/p87v-9846" name="Koninklijk Museum voor Midden Afrika" pageId="16" pageNumber="217" type="Museum">KMMA</collectionCode>
);
<specimenCount id="9D5637B3FFA7FFE70C13318DFDE0FAE5" box="[553,588,1269,1295]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, Gamangui,
<date id="FFEEDAFAFFA7FFE70CD8318DFC27FAE5" box="[738,907,1269,1295]" pageId="16" pageNumber="217" value="1910-02">
<collectingDate id="EFAA2312FFA7FFE70CD8318DFC27FAE5" box="[738,907,1269,1295]" pageId="16" pageNumber="217" value="1910-02">February 1910</collectingDate>
</date>
,
<collectorName id="26A599ECFFA7FFE70DA3318DFC7FFAE5" box="[921,979,1269,1295]" pageId="16" pageNumber="217">Lang</collectorName>
&amp;
<collectorName id="26A599ECFFA7FFE70DC1318DFBE0FAE5" box="[1019,1100,1269,1295]" pageId="16" pageNumber="217">Chapin</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70A67318DFB15FAE5" box="[1117,1209,1269,1295]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="16" pageNumber="217">AMNH</collectionCode>
);
<specimenCount id="9D5637B3FFA7FFE70AF5318DFB5EFAE5" box="[1231,1266,1269,1295]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
,
<collectorName id="26A599ECFFA7FFE70B38318DFA35FAE5" box="[1282,1433,1269,1295]" pageId="16" pageNumber="217">Manyema, R</collectorName>
. Mayné (
<collectionCode id="ED4164FFFFA7FFE70ECF3061FEFAFAD9" box="[245,342,1305,1331]" country="Belgium" httpUri="http://grbio.org/cool/p87v-9846" name="Koninklijk Museum voor Midden Afrika" pageId="16" pageNumber="217" type="Museum">KMMA</collectionCode>
);
<specimenCount id="9D5637B3FFA7FFE70F503061FE21FAD9" box="[362,397,1305,1331]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, Medje, Ituri,
<date id="FFEEDAFAFFA7FFE70C083061FD41FAD9" box="[562,749,1305,1331]" pageId="16" pageNumber="217" value="1910-09">
<collectingDate id="EFAA2312FFA7FFE70C083061FD41FAD9" box="[562,749,1305,1331]" pageId="16" pageNumber="217" value="1910-09">September 1910</collectingDate>
</date>
,
<collectorName id="26A599ECFFA7FFE70CC33061FC9FFAD9" box="[761,819,1305,1331]" pageId="16" pageNumber="217">Lang</collectorName>
&amp;
<collectorName id="26A599ECFFA7FFE70D633061FC06FAD9" box="[857,938,1305,1331]" pageId="16" pageNumber="217">Chapin</collectorName>
(
<collectionCode id="ED4164FFFFA7FFE70D803061FBB7FAD9" box="[954,1051,1305,1331]" country="Belgium" httpUri="http://grbio.org/cool/p87v-9846" name="Koninklijk Museum voor Midden Afrika" pageId="16" pageNumber="217" type="Museum">KMMA</collectionCode>
);
<specimenCount id="9D5637B3FFA7FFE70A153061FBFEFAD9" box="[1071,1106,1305,1331]" pageId="16" pageNumber="217" type="female">1♀</specimenCount>
, Nyangwe,
<date id="FFEEDAFAFFA7FFE70AE63061FACBFAD9" box="[1244,1383,1305,1331]" pageId="16" pageNumber="217" value="1910-12-20">
<collectingDate id="EFAA2312FFA7FFE70AE63061FACBFAD9" box="[1244,1383,1305,1331]" pageId="16" pageNumber="217" value="1910-12-20">20.XII.1910</collectingDate>
</date>
, Be- quaert (
<collectionCode id="ED4164FFFFA7FFE70ED63045FEE6FABD" box="[236,330,1341,1367]" country="Belgium" httpUri="http://grbio.org/cool/p87v-9846" name="Koninklijk Museum voor Midden Afrika" pageId="16" pageNumber="217" type="Museum">KMMA</collectionCode>
) (labelled as
<typeStatus id="54EB4298FFA7FFE70FE53045FDEDFABD" box="[479,577,1341,1367]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5087B9FFA7FFE70C5D3046FD51FABD" box="[615,765,1341,1367]" class="Insecta" family="Syrphidae" genus="Phytomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="pubipennis">
<emphasis id="B9242028FFA7FFE70C5D3046FD51FABD" box="[615,765,1341,1367]" italics="true" pageId="16" pageNumber="217">P. pubipennis</emphasis>
</taxonomicName>
but see Comments under
<taxonomicName id="4C5087B9FFA7FFE70A1B3046FB1BFABD" box="[1057,1207,1341,1367]" class="Insecta" family="Syrphidae" genus="Phytomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="pubipennis">
<emphasis id="B9242028FFA7FFE70A1B3046FB1BFABD" box="[1057,1207,1341,1367]" italics="true" pageId="16" pageNumber="217">P. pubipennis</emphasis>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEFFC3AFFA7FFE70EFD3019FBC6FA91" blockId="16.[151,1437,908,2015]" box="[199,1130,1376,1403]" pageId="16" pageNumber="217">
<emphasis id="B9242028FFA7FFE70EFD3019FE94FA91" bold="true" box="[199,312,1376,1403]" pageId="16" pageNumber="217">Body size</emphasis>
. Body length (n=10): 22.4–
<quantity id="4CA851DFFFA7FFE70C483019FD74FA91" box="[626,728,1377,1403]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="16" pageNumber="217" unit="mm" value="13.2">13.2 mm</quantity>
. Wing length (n=10): 8.9–10.0 mm.
</paragraph>
</subSubSection>
<subSubSection id="C34AAFB1FFA7FFE60EFD30FCFB39FD29" lastPageId="17" lastPageNumber="218" pageId="16" pageNumber="217" type="description">
<paragraph id="8BEFFC3AFFA7FFE70EFD30FCFC82F971" blockId="16.[151,1437,908,2015]" pageId="16" pageNumber="217">
<emphasis id="B9242028FFA7FFE70EFD30FCFEFDFA75" bold="true" box="[199,337,1412,1439]" pageId="16" pageNumber="217">Description</emphasis>
.
<emphasis id="B9242028FFA7FFE70F6530FCFE1FFA75" bold="true" box="[351,435,1412,1439]" pageId="16" pageNumber="217">MALE</emphasis>
(
<figureCitation id="136BE0BFFFA7FFE70FFB30FDFDA8FA4A" box="[449,516,1413,1440]" captionStart="FIGURES 1–10" captionStartId="4.[151,264,1931,1956]" captionTargetBox="[162,1426,181,1905]" captionTargetId="figure@4.[161,1426,181,1905]" captionTargetPageId="4" captionText="FIGURES 1–10. Phytomia spp., habitus, lateral view. 1. P. bullata (Loew) ♂ 2. P. aurigera Bezzi ♂ 3. P. kroeberi (Bezzi) ♂ 4. P. serena (Curran) ♀ 5. P. bulligera (Austen) ♂ 6. P. austeni sp. nov. ♂ 7. P. bezzii Curran ♂ 8. P. memnon sp. nov. ♂ 9. P. pubipennis Bezzi ♂ 10. P. natalensis (Macquart) ♂" figureDoi="http://doi.org/10.5281/zenodo.3909049" httpUri="https://zenodo.org/record/3909049/files/figure.png" pageId="16" pageNumber="217">Fig. 6</figureCitation>
).
<emphasis id="B9242028FFA7FFE70C2330FCFDFBFA75" bold="true" box="[537,599,1412,1439]" italics="true" pageId="16" pageNumber="217">Head</emphasis>
. Eye bare; holoptic, touching for 2–3 times length of ocellar triangle, facets enlarged in dorsal half. Frons brown to black; with greyish brown pollinosity; dispersed pilosity of medium long pale pile; rugose area shining black, occupying one-third of entire width and two-fifths of entire height; with shorter dispersed pale pile. Face ground colour black; with dispersed greyish to greyish brown pollinosity, in parts weakly shining black; dispersed pilosity of short pale pile; facial tubercle elongated, strongly pronounced. Gena colour and pollinosity as face; with long pale pilosity. Occiput dorsal third black, with greyish pollinosity; ventrally more densely yellow-white, with dispersed pale pile. Antennal segments brown to black-brown; arista yellow-brown, in basal half with medium long pile equal to width of pedicel.
</paragraph>
<paragraph id="8BEFFC3AFFA7FFE70EFD33DDFE38F8C1" blockId="16.[151,1437,908,2015]" pageId="16" pageNumber="217">
<emphasis id="B9242028FFA7FFE70EFD33DDFE8CF955" bold="true" box="[199,288,1701,1727]" pageId="16" pageNumber="217">Thorax</emphasis>
. Scutum and scutellum black ground colour, with dense orange-brown pollinosity; covered with dense pile of long orange to orange-brown pilosity. Pleural sclerites ground colour black, with greyish brown pollinosity; covered with dispersed pile of long golden yellow to orange pile except meron, anterior part of katepisternum and anterior anepisternum.
</paragraph>
<paragraph id="8BEFFC3AFFA7FFE60EFD324DFB72FF13" blockId="16.[151,1437,908,2015]" lastBlockId="17.[151,1437,153,1446]" lastPageId="17" lastPageNumber="218" pageId="16" pageNumber="217">
<emphasis id="B9242028FFA7FFE70EFD324DFF53F8A5" bold="true" box="[199,255,1845,1871]" pageId="16" pageNumber="217">Legs</emphasis>
. Yellow-orange to black-brown. Fore leg, femur black brown, apical margin narrowly paler; with dense short black pilosity, anteriorly and posteriorly longer, in few specimens also partially paler at base; tibia pale in basal third, otherwise orange-brown; with short black pilosity, except in basal third where pale; tarsal segments yelloworange. Mid leg as in fore leg, except tibia pale for basal half, and pilosity pale over entire length; sometimes pilosity on femur more golden yellow basally, especially along posterior part. Hind leg (
<figureCitation id="136BE0BFFFA7FFE70A2032BDFBC2F835" box="[1050,1134,1989,2015]" captionStart="FIGURES 29–36" captionStartId="13.[151,264,1945,1970]" captionTargetBox="[242,1360,181,1919]" captionTargetId="figure@13.[227,1360,181,1919]" captionTargetPageId="13" captionText="FIGURES 29–36. Figs 29–34. Phytomia spp., hind leg, anterior view. 29. P. bullata (Loew) ♂ 30. P. serena (Curran) ♀ 31. P. bulligera (Austen) ♂ 32. P. austeni sp. nov. ♂ 33. P. bezzii Curran ♂ 34. P. memnon sp. nov. ♂ Figs 35–36: Phytomia spp., scutellum, dorsal view. 35. P. melas (Bezzi) ♂ 36. P. poensis (Bezzi) ♀" figureDoi="http://doi.org/10.5281/zenodo.3909055" httpUri="https://zenodo.org/record/3909055/files/figure.png" pageId="16" pageNumber="217">Fig. 32</figureCitation>
), femur slightly thickened; black-brown, at extremities narrowly paler; dorsally with long pale pilosity, ventrally with long dark pilosity, towards base paler; tibia orange-brown to black brown, pale at basal tip; dorsally with short pale to golden yellow pilosity, ventrally with short to medium-long dense black pilosity; tarsal segments yellow-orange.
</paragraph>
<paragraph id="8BEFFC3AFFA6FFE60EFD347AFE77FE6C" blockId="17.[151,1437,153,1446]" pageId="17" pageNumber="218">
<emphasis id="B9242028FFA6FFE60EFD347AFEA4FEF7" bold="true" box="[199,264,258,285]" pageId="17" pageNumber="218">Wing</emphasis>
(fig. 43). Mainly hyaline. Microtrichia along base; also patch in medial part of wing anteriorly from apex of vein
<emphasis id="B9242028FFA6FFE60ED6345EFEAAFEAA" box="[236,262,294,320]" italics="true" pageId="17" pageNumber="218">Sc</emphasis>
, posteriorly along medial part of cell
<emphasis id="B9242028FFA6FFE60C93345FFD10FEAF" box="[681,700,295,325]" italics="true" pageId="17" pageNumber="218">
r
<subScript id="17D4FE7FFFA6FFE60C8E344FFD10FEAF" attach="left" box="[692,700,311,325]" fontSize="6" pageId="17" pageNumber="218">1</subScript>
</emphasis>
, basal part of cell
<emphasis id="B9242028FFA6FFE60DB2345FFC02FEAF" box="[904,942,295,325]" italics="true" pageId="17" pageNumber="218">
r
<subScript id="17D4FE7FFFA6FFE60DA9344FFC02FEAF" attach="left" box="[915,942,311,325]" fontSize="6" pageId="17" pageNumber="218">2+3</subScript>
</emphasis>
, and into cell
<taxonomicName id="4C5087B9FFA6FFE60A71345FFB0DFEAA" box="[1099,1185,294,320]" pageId="17" pageNumber="218" rank="subSpecies" subSpecies="where">
<emphasis id="B9242028FFA6FFE60A71345FFBFAFEAA" box="[1099,1110,295,320]" italics="true" pageId="17" pageNumber="218">r</emphasis>
where
</taxonomicName>
it reaches the junction of vein
<emphasis id="B9242028FFA6FFE60ED03432FEADFE89" box="[234,257,330,355]" italics="true" pageId="17" pageNumber="218">M</emphasis>
with crossvein
<emphasis id="B9242028FFA6FFE60F953431FE59FE89" box="[431,501,329,355]" italics="true" pageId="17" pageNumber="218">bm-cu</emphasis>
; otherwise with faint streaks of microtrichia in cells
<emphasis id="B9242028FFA6FFE60A033432FBE9FE89" box="[1081,1093,330,355]" italics="true" pageId="17" pageNumber="218">c</emphasis>
,
<emphasis id="B9242028FFA6FFE60A693431FBD9FE89" box="[1107,1141,329,355]" italics="true" pageId="17" pageNumber="218">bm</emphasis>
,
<emphasis id="B9242028FFA6FFE60AB83432FB06FE89" box="[1154,1194,330,355]" italics="true" pageId="17" pageNumber="218">cup</emphasis>
,
<emphasis id="B9242028FFA6FFE60A8D3432FB66FE82" box="[1207,1226,330,360]" italics="true" pageId="17" pageNumber="218">
r
<subScript id="17D4FE7FFFA6FFE60AF83422FB66FE82" attach="left" box="[1218,1226,346,360]" fontSize="6" pageId="17" pageNumber="218">1</subScript>
</emphasis>
and
<emphasis id="B9242028FFA6FFE60AC43432FA88FE82" box="[1278,1316,330,360]" italics="true" pageId="17" pageNumber="218">
r
<subScript id="17D4FE7FFFA6FFE60B333422FA88FE82" attach="left" box="[1289,1316,346,360]" fontSize="6" pageId="17" pageNumber="218">2+3</subScript>
</emphasis>
. Calypters dark with fringe of dark pile.
</paragraph>
<paragraph id="8BEFFC3AFFA6FFE60EFD34F6FDDBFE1A" blockId="17.[151,1437,153,1446]" pageId="17" pageNumber="218">
<emphasis id="B9242028FFA6FFE60EFD34F6FE90FE43" bold="true" box="[199,316,398,425]" pageId="17" pageNumber="218">Abdomen</emphasis>
(
<figureCitation id="136BE0BFFFA6FFE60F7134F7FE0CFE40" box="[331,416,399,426]" captionStart="FIGURES 61–72" captionStartId="29.[151,264,1945,1970]" captionTargetBox="[182,1431,181,1918]" captionTargetId="figure@29.[156,1431,181,1919]" captionTargetPageId="29" captionText="FIGURES 61–72. Phytomia spp., abdomen, dorsal view. 61. P. bullata (Loew) ♂ 62. P. aurigera Bezzi ♂ 63. P. kroeberi (Bezzi) ♂ 64–65. P. kroeberi (Bezzi) ♀ 66. P. serena (Curran) ♀ 67. P. bulligera (Austen) ♂ 68. P. austeni sp. nov. ♂ 69. P. bezzii Curran ♀ 70. P. memnon sp. nov. ♀ 71. P. pubipennis Bezzi ♂ 72. P. natalensis (Macquart) (M)." figureDoi="http://doi.org/10.5281/zenodo.3909061" httpUri="https://zenodo.org/record/3909061/files/figure.png" pageId="17" pageNumber="218">Fig. 68</figureCitation>
). Mainly orange-brown to brown ground colour. Terga 2–4 with darker medial bulla; pilosity with dispersed short to medium long pale pile, longer along margins. Sterna black-brown, with widely dispersed long pale pile. Male genitalia as in
<figureCitation id="136BE0BFFFA6FFE60C1B34AEFDDFFE1A" box="[545,627,470,496]" captionStart="FIGURES 85–100" captionStartId="37.[151,264,1727,1752]" captionTargetBox="[151,1436,381,1702]" captionTargetId="figure@37.[151,1436,381,1702]" captionTargetPageId="37" captionText="FIGURES 85–100. Phytomia spp., male terminalia, dorsal view. 85. P. bullata (Loew). 86. P. aurigera Bezzi. 87. P. kroeberi (Bezzi). 88. P. bulligera (Austen). 89. P. austeni sp. nov. 90. P. bezzii Curran. 91. P. memnon sp. nov. 92. P. pubipennis Bezzi. 93. P. natalensis (Macquart). 94. P. curta (Loew). 95. P. pallida sp. nov. 96. P. erratica (Bezzi). 97. P. fucoides Bezzi. 98. P. incisa (Wiedemann). 99. P. melas (Bezzi). 100. P. poensis (Bezzi)." figureDoi="http://doi.org/10.5281/zenodo.3909065" httpUri="https://zenodo.org/record/3909065/files/figure.png" pageId="17" pageNumber="218">Fig. 89</figureCitation>
.
</paragraph>
<paragraph id="8BEFFC3AFFA6FFE60EFD3480FB39FD29" blockId="17.[151,1437,153,1446]" pageId="17" pageNumber="218">
<emphasis id="B9242028FFA6FFE60EFD3480FE93FDF9" bold="true" box="[199,319,504,531]" pageId="17" pageNumber="218">FEMALE</emphasis>
. As male except for the following character states. Eye, facets of equal size; dichoptic. Frons black ground colour; with greyish brown pollinosity, except in dorsal third where dark brown; covered with short dispersed pale to yellow-orange pilosity; rugose area shining black-brown; occupying one-third of entire width and equal in height to at most twice length of ocellar triangle. Face near antennal insertion more orange-brown. Wing (
<figureCitation id="136BE0BFFFA6FFE60E9A37FEFF5CFD4A" box="[160,240,646,672]" captionStart="FIGURES 37–48" captionStartId="21.[151,264,1945,1970]" captionTargetBox="[157,1428,201,1919]" captionTargetId="figure@21.[157,1431,181,1919]" captionTargetPageId="21" captionText="FIGURES 37–48. Phytomia spp., right wing. 37. P. bullata (Loew) ♀ 38. P. aurigera Bezzi ♂ 39–40. P. kroeberi (Bezzi) ♀ 41. P. serena (Curran) ♀ 42. P. bulligera (Austen) ♀ 43. P. austeni sp. nov. ♂ 44. P. austeni sp. nov. ♀ 45. P. bezzii Curran ♂ 46. P. bezzii Curran ♀ 47. P. memnon sp. nov. ♀ 48. P. pubipennis Bezzi ♂" figureDoi="http://doi.org/10.5281/zenodo.3909057" httpUri="https://zenodo.org/record/3909057/files/figure.png" pageId="17" pageNumber="218">Fig. 44</figureCitation>
) more extensively covered with microtrichia, covering most of wing surface except along posterior margin; giving wing a darker appearance, medial patch along cross-vein
<emphasis id="B9242028FFA6FFE60D5337D2FC3DFD29" box="[873,913,682,707]" italics="true" pageId="17" pageNumber="218">r-m</emphasis>
more densely covered.
</paragraph>
</subSubSection>
<subSubSection id="C34AAFB1FFA6FFE60EFD37B3FD8CFCE3" pageId="17" pageNumber="218" type="diagnosis">
<paragraph id="8BEFFC3AFFA6FFE60EFD37B3FD8CFCE3" blockId="17.[151,1437,153,1446]" pageId="17" pageNumber="218">
<emphasis id="B9242028FFA6FFE60EFD37B3FE96FD0C" bold="true" box="[199,314,715,742]" pageId="17" pageNumber="218">Diagnosis</emphasis>
. Recognizable by the black fringe on the ventral calypter in both sexes and the extensive mictrotrichose covering in the female wing.
</paragraph>
</subSubSection>
<subSubSection id="C34AAFB1FFA6FFE60EFD366AFC2BFCC7" box="[199,903,786,813]" pageId="17" pageNumber="218" type="distribution">
<paragraph id="8BEFFC3AFFA6FFE60EFD366AFC2BFCC7" blockId="17.[151,1437,153,1446]" box="[199,903,786,813]" pageId="17" pageNumber="218">
<emphasis id="B9242028FFA6FFE60EFD366AFEF5FCC7" bold="true" box="[199,345,786,813]" pageId="17" pageNumber="218">Distribution</emphasis>
.
<collectingCountry id="F347BCAAFFA6FFE60F5D366AFE01FCC6" box="[359,429,786,812]" name="Benin" pageId="17" pageNumber="218">Benin</collectingCountry>
,
<collectingCountry id="F347BCAAFFA6FFE60F82366BFDAFFCC7" box="[440,515,787,813]" name="Democratic Republic of the Congo" pageId="17" pageNumber="218">Congo</collectingCountry>
(DR),
<collectingCountry id="F347BCAAFFA6FFE60C68366AFD32FCC6" box="[594,670,786,812]" name="Ghana" pageId="17" pageNumber="218">Ghana</collectingCountry>
,
<collectingCountry id="F347BCAAFFA6FFE60C93366AFC97FCC7" box="[681,827,786,813]" name="Sierra Leone" pageId="17" pageNumber="218">Sierra Leone</collectingCountry>
,
<collectingCountry id="F347BCAAFFA6FFE60D7C366BFC2FFCC7" box="[838,899,787,813]" name="Togo" pageId="17" pageNumber="218">Togo</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C34AAFB1FFA6FFE60EFD364EFC43FC7D" pageId="17" pageNumber="218" type="etymology">
<paragraph id="8BEFFC3AFFA6FFE60EFD364EFC43FC7D" blockId="17.[151,1437,153,1446]" pageId="17" pageNumber="218">
<emphasis id="B9242028FFA6FFE60EFD364EFEE4FCBA" bold="true" box="[199,328,822,848]" pageId="17" pageNumber="218">Etymology</emphasis>
. Named after the dipterist Ernest E. Austen (
<date id="FFEEDAFAFFA6FFE60D0E364EFC19FCBB" box="[820,949,822,849]" pageId="17" pageNumber="218" value="1867" valueMax="1938">1867–1938</date>
) who, when describing
<taxonomicName id="4C5087B9FFA6FFE60A86364FFA91FCBA" authorityName=", Austen" authorityYear="1909" box="[1212,1341,822,848]" class="Insecta" family="Syrphidae" genus="Megaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="bulligera">
<emphasis id="B9242028FFA6FFE60A86364FFA91FCBA" box="[1212,1341,822,848]" italics="true" pageId="17" pageNumber="218">P. bulligera</emphasis>
</taxonomicName>
, already indicated that some specimens where deviant in morphological characters, which led to the description of this new species. The specific epithet should be treated as a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C34AAFB1FFA6FFE60EFD36E6FF76FA4C" pageId="17" pageNumber="218" type="discussion">
<paragraph id="8BEFFC3AFFA6FFE60EFD36E6FF76FA4C" blockId="17.[151,1437,153,1446]" pageId="17" pageNumber="218">
<emphasis id="B9242028FFA6FFE60EFD36E6FEE4FC53" bold="true" box="[199,328,926,953]" pageId="17" pageNumber="218">Comments</emphasis>
. When describing
<taxonomicName id="4C5087B9FFA6FFE60C1A36D8FCF9FC53" authority=", Austen (1909)" authorityName=", Austen" authorityYear="1909" box="[544,853,927,954]" class="Insecta" family="Syrphidae" genus="Megaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="bulligera">
<emphasis id="B9242028FFA6FFE60C1A36D8FD09FC53" box="[544,677,927,953]" italics="true" pageId="17" pageNumber="218">P. bulligera</emphasis>
,
<bibRefCitation id="EFC181CBFFA6FFE60C8836D8FCF9FC53" author="Austen, E. E." box="[690,853,927,954]" pageId="17" pageNumber="218" pagination="85 - 102" refId="ref44103" refString="Austen, E. E. (1909) Ruwenzori expedition reports. 10. Diptera. Transactions of the Zoological Society of London, 19, 85 - 102. https: // doi. org / 10.1111 / j. 1469 - 7998.1909. tb 08562. x" type="journal article" year="1909">Austen (1909)</bibRefCitation>
</taxonomicName>
already indicated that the male specimen collected in
<collectingCountry id="F347BCAAFFA6FFE60E8E36BAFEE9FC37" box="[180,325,962,989]" name="Sierra Leone" pageId="17" pageNumber="218">Sierra Leone</collectingCountry>
deviated in abdominal coloration from the male
<typeStatus id="54EB4298FFA6FFE60D5636BBFC30FC37" box="[876,924,963,989]" pageId="17" pageNumber="218">type</typeStatus>
specimen from
<collectingCountry id="F347BCAAFFA6FFE60A6836BAFB01FC36" box="[1106,1197,962,988]" name="Uganda" pageId="17" pageNumber="218">Uganda</collectingCountry>
. After morphological examination of long series, the former specimen turned out to belong to a different species described here and that differs from
<taxonomicName id="4C5087B9FFA6FFE60F133172FE02FBC9" authorityName=", Austen" authorityYear="1909" box="[297,430,1033,1059]" class="Insecta" family="Syrphidae" genus="Megaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="bulligera">
<emphasis id="B9242028FFA6FFE60F133172FE02FBC9" box="[297,430,1033,1059]" italics="true" pageId="17" pageNumber="218">P. bulligera</emphasis>
</taxonomicName>
by the following main characteristics: ventral calypter with fringe of black pile (fringe of pale pile in
<taxonomicName id="4C5087B9FFA6FFE60F7F3155FE67FBAC" authorityName=", Austen" authorityYear="1909" box="[325,459,1068,1094]" class="Insecta" family="Syrphidae" genus="Megaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="bulligera">
<emphasis id="B9242028FFA6FFE60F7F3155FE67FBAC" box="[325,459,1068,1094]" italics="true" pageId="17" pageNumber="218">P. bulligera</emphasis>
</taxonomicName>
), abdominal tergum 2 along anterior margin at most with dispersed short white pile anterolaterally (with conspicuous long white pile along whole anterior margin, distinctly widened anterolaterally in
<taxonomicName id="4C5087B9FFA6FFE60E8C310BFE97FB66" authorityName=", Austen" authorityYear="1909" box="[182,315,1138,1164]" class="Insecta" family="Syrphidae" genus="Megaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="bulligera">
<emphasis id="B9242028FFA6FFE60E8C310BFE97FB66" box="[182,315,1138,1164]" italics="true" pageId="17" pageNumber="218">P. bulligera</emphasis>
</taxonomicName>
), and female with microtrichosity very extensive covering most of the wing surface except along posterior margin (microtrichosity restricted to base, anterior margin and weakly developed medial patch in
<taxonomicName id="4C5087B9FFA6FFE60B6931EFFF76FB39" authorityName=", Austen" authorityYear="1909" class="Insecta" family="Syrphidae" genus="Megaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="bulligera">
<emphasis id="B9242028FFA6FFE60B6931EFFF76FB39" italics="true" pageId="17" pageNumber="218">P. bulligera</emphasis>
</taxonomicName>
). DNA barcoding showed that specimens of both species clustered into different groups. The mean p-distance between both species is 9.6 % and much higher than both mean intraspecific p-distances (0.2 % for
<taxonomicName id="4C5087B9FFA6FFE60AD031A5FAC7FB1C" authorityName=", Austen" authorityYear="1909" box="[1258,1387,1244,1270]" class="Insecta" family="Syrphidae" genus="Megaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="bulligera">
<emphasis id="B9242028FFA6FFE60AD031A5FAC7FB1C" box="[1258,1387,1244,1270]" italics="true" pageId="17" pageNumber="218">P. bulligera</emphasis>
</taxonomicName>
; 0.1 % for
<taxonomicName id="4C5087B9FFA6FFE60EE53078FEE1FAF3" authorityName="Meyer &amp; Goergen &amp; Jordaens" authorityYear="2020" box="[223,333,1280,1305]" class="Insecta" family="Syrphidae" genus="Phytomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="austeni">
<emphasis id="B9242028FFA6FFE60EE53078FEE1FAF3" box="[223,333,1280,1305]" italics="true" pageId="17" pageNumber="218">P. austeni</emphasis>
</taxonomicName>
) and well above the mean interspecific p-distance for
<taxonomicName id="4C5087B9FFA6FFE60D8C3187FB8DFAF3" box="[950,1057,1279,1305]" class="Insecta" family="Syrphidae" genus="Phytomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="genus">
<emphasis id="B9242028FFA6FFE60D8C3187FB8DFAF3" box="[950,1057,1279,1305]" italics="true" pageId="17" pageNumber="218">Phytomia</emphasis>
</taxonomicName>
(6.6 %), confirming the morphological differentiation. A number of females are not included in the
<typeStatus id="54EB4298FFA6FFE60DAC305BFC6AFAD7" box="[918,966,1315,1341]" pageId="17" pageNumber="218">type</typeStatus>
series. They originate from
<collectingCountry id="F347BCAAFFA6FFE60B33305BFAF8FAD7" box="[1289,1364,1315,1341]" name="Democratic Republic of the Congo" pageId="17" pageNumber="218">Congo</collectingCountry>
(DR), geographically distinct from the main series which originates from western Africa. As this material is relatively old and the majority in poor condition, no DNA could be extracted to verify whether they would group in the
<taxonomicName id="4C5087B9FFA6FFE60B0A3012FA30FA69" authorityName="Meyer &amp; Goergen &amp; Jordaens" authorityYear="2020" box="[1328,1436,1386,1411]" class="Insecta" family="Syrphidae" genus="Phytomia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="austeni">
<emphasis id="B9242028FFA6FFE60B0A3012FA30FA69" box="[1328,1436,1386,1411]" italics="true" pageId="17" pageNumber="218">P. austeni</emphasis>
</taxonomicName>
clade.
</paragraph>
</subSubSection>
</treatment>
</document>
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