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<document id="C12EB428BF836DB9D0924A649687A6A2" ID-DOI="10.1080/00222933.2017.1326640" ID-ISSN="1464-5262" ID-Zenodo-Dep="5181243" ID-ZooBank="BD40ACCF-3EEB-43C3-BC54-0D2BB88F679F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628689236996" checkinUser="felipe" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. &amp; Vaz-de-Mello, Fernando Z." docDate="2017" docId="03E7473FFFDBFFACFE66FB8776E825FA" docLanguage="en" docName="JNatHist.51.17-18.995-1014.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (17 - 18)" docSource="http://dx.doi.org/10.1080/00222933.2017.1326640" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Paraegidium Vulcano 1966" docType="treatment" docVersion="3" lastPageNumber="998" masterDocId="FFDE3F47FFD9FFA8FFAFFFE0726C265C" masterDocTitle="Revision of the South American genus Paraegidium Vulcano et al. (Coleoptera: Scarabaeidae: Orphninae)" masterLastPageNumber="1014" masterPageNumber="995" pageNumber="996" updateTime="1699049738498" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="32EACACB1EFE8016FD278D0472B57EFA">Revision of the South American genus Paraegidium Vulcano et al. (Coleoptera: Scarabaeidae: Orphninae)</mods:title>
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<mods:namePart id="6194F0A9739310D19141A5A54DB8D916">Akhmetova, Lilia A.</mods:namePart>
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<mods:namePart id="C0B243A6D11A4CC6C71A548F47BFF22D">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:number id="881F0750A1336813B29E9ECB153BC3E3">2017-05-25</mods:number>
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<subSubSection id="C354A5A2FFDBFFAAFE66FB87713522DD" box="[457,857,1127,1153]" pageId="2" pageNumber="996" type="nomenclature">
<paragraph id="8BF1F629FFDBFFAAFE66FB87713522DD" blockId="2.[457,857,1127,1153]" box="[457,857,1127,1153]" pageId="2" pageNumber="996">
<heading id="D0B94145FFDBFFAAFE66FB87713522DD" box="[457,857,1127,1153]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="996" reason="5">
<taxonomicName id="4C4E8DAAFFDBFFAAFE66FB87713522DD" authority="Vulcano et al., 1966" authorityName="Vulcano" authorityYear="1966" box="[457,857,1127,1153]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="B93A2A3BFFDBFFAAFE66FB87700922DD" bold="true" box="[457,613,1127,1153]" italics="true" pageId="2" pageNumber="996">Paraegidium</emphasis>
<bibRefCitation id="EFDF8BD8FFDBFFAAFDC0FB88713522DD" author="Vulcano MA &amp; Pereira FS &amp; Martinez A" box="[623,857,1128,1153]" pageId="2" pageNumber="996" pagination="251 - 260" refId="ref8214" refString="Vulcano MA, Pereira FS, Martinez A. 1966. Notas sobre Orphninae Neotropicos com descricao de um genero e una especie novos (Coleoptera). Papeis Dep Zool S Paulo. 18: 251 - 260." type="journal article" year="1966">Vulcano et al., 1966</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C354A5A2FFDBFFAAFF0FFB7D767A22A7" pageId="2" pageNumber="996" type="materials_examined">
<paragraph id="8BF1F629FFDBFFAAFF0FFB7D76E82284" blockId="2.[160,1156,1181,1275]" pageId="2" pageNumber="996">
<typeStatus id="54F5488BFFDBFFAAFF0FFB7D72BA22E9" box="[160,214,1181,1205]" pageId="2" pageNumber="996">Type</typeStatus>
species:
<taxonomicName id="4C4E8DAAFFDBFFAAFE90FB7D714522E9" authority="Vulcano et al., 1966" authorityName="Vulcano" authorityYear="1966" box="[319,809,1181,1205]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="996" phylum="Arthropoda" rank="species" species="costalimai">
<emphasis id="B93A2A3BFFDBFFAAFE90FB7D702C22E9" box="[319,576,1181,1205]" italics="true" pageId="2" pageNumber="996">Paraegidium costalimai</emphasis>
<bibRefCitation id="EFDF8BD8FFDBFFAAFDE6FB7D714522E9" author="Vulcano MA &amp; Pereira FS &amp; Martinez A" box="[585,809,1181,1205]" pageId="2" pageNumber="996" pagination="251 - 260" refId="ref8214" refString="Vulcano MA, Pereira FS, Martinez A. 1966. Notas sobre Orphninae Neotropicos com descricao de um genero e una especie novos (Coleoptera). Papeis Dep Zool S Paulo. 18: 251 - 260." type="journal article" year="1966">Vulcano et al., 1966</bibRefCitation>
</taxonomicName>
, by original designation.
<taxonomicName id="4C4E8DAAFFDBFFAAFF0FFB2070282284" authority=": Vulcano et al. 1966: 252" authorityName="Vulcano" authorityPageNumber="252" authorityYear="1966" box="[160,580,1216,1240]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="996" phylum="Arthropoda" rank="genus">
<emphasis id="B93A2A3BFFDBFFAAFF0FFB2073462284" box="[160,298,1216,1240]" italics="true" pageId="2" pageNumber="996">Paraegidium</emphasis>
:
<bibRefCitation id="EFDF8BD8FFDBFFAAFE99FB2070282284" author="Vulcano MA &amp; Pereira FS &amp; Martinez A" box="[310,580,1216,1240]" pageId="2" pageNumber="996" pagination="251 - 260" refId="ref8214" refString="Vulcano MA, Pereira FS, Martinez A. 1966. Notas sobre Orphninae Neotropicos com descricao de um genero e una especie novos (Coleoptera). Papeis Dep Zool S Paulo. 18: 251 - 260." type="journal article" year="1966">Vulcano et al. 1966: 252</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFDF8BD8FFDBFFAAFDE2FB20716B2284" author="Paulian R" box="[589,775,1216,1240]" pageId="2" pageNumber="996" pagination="65 - 92" refId="ref8141" refString="Paulian R. 1984. Les Orphnidae Americains (Coleopteres, Scarabaeoidea). Ann Soc Ent Fr (NS). 20: 65 - 92." type="journal article" year="1984">Paulian 1984: 91</bibRefCitation>
,
<bibRefCitation id="EFDF8BD8FFDBFFAAFCBDFB2071D62284" author="Colby J" box="[786,954,1216,1240]" pageId="2" pageNumber="996" pagination="1 - 41" refId="ref7888" refString="Colby J. 2009. Monographic revision of the genus Aegidinus Arrow (1904) and generic phylogeny of the world Orphninae (Coleoptera: Scarabaeidae: Orphninae). Insecta Mundi. 76: 1 - 41." type="journal article" year="2009">Colby 2009: 16</bibRefCitation>
,
<bibRefCitation id="EFDF8BD8FFDBFFAAFC6BFB2076ED2284" author="Frolov AV" box="[964,1153,1216,1240]" pageId="2" pageNumber="996" pagination="782 - 797" refId="ref7927" refString="Frolov AV. 2012. Diagnosis, classification, and phylogenetic relationships of the orphnine scarab beetles (Coleoptera, Scarabaeidae: Orphninae). Ent Rev. 92: 782 - 797." type="journal article" year="2012">Frolov 2012: 783</bibRefCitation>
,
</paragraph>
<paragraph id="8BF1F629FFDBFFAAFF6FFB03767A22A7" blockId="2.[160,1156,1181,1275]" box="[192,1046,1250,1275]" pageId="2" pageNumber="996">
<bibRefCitation id="EFDF8BD8FFDBFFAAFF6FFB03734F22A7" author="Frolov AV" box="[192,291,1251,1275]" pageId="2" pageNumber="996" pagination="33 - 46" refId="ref7961" refString="Frolov AV. 2013. Stenosternus Karsch, a possible link between Neotropical and Afrotropical Orphninae (Coleoptera, Scarabaeidae). Zookeys. 335: 33 - 46." type="journal article" year="2013">2013: 37</bibRefCitation>
,
<bibRefCitation id="EFDF8BD8FFDBFFAAFE80FB0370D122A7" author="Frolov AV &amp; Vaz-de-Mello FZ" box="[303,701,1251,1275]" pageId="2" pageNumber="996" pagination="433 - 436" refId="ref8102" refString="Frolov AV, Vaz-de-Mello FZ. 2015. A new genus and species of Orphninae (Coleoptera: Scarabaeidae) associated with epiphytes in an Andean cloud forest in Ecuador. Zootaxa. 4007: 433 - 436." type="journal article" year="2015">Frolov and Vaz-de-Mello 2015: 435</bibRefCitation>
,
<bibRefCitation id="EFDF8BD8FFDBFFAAFD67FB03767F22A7" author="Rojkoff S &amp; Frolov AV" box="[712,1043,1250,1275]" pageId="2" pageNumber="996" pagination="354 - 368" refId="ref8169" refString="Rojkoff S, Frolov AV. 2017. Revision des especes du genre Aegidium Westwood, 1845, des Petites Antilles (Coleoptera: Scarabaeidae, Orphninae, Aegidiini). Ann Soc Ent Fr (NS). 52: 354 - 368." type="journal article" year="2017">Rojkoff and Frolov 2017: 355</bibRefCitation>
.
</paragraph>
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<subSubSection id="C354A5A2FFDBFFABFF0FFAC9763D2478" lastPageId="3" lastPageNumber="997" pageId="2" pageNumber="996" type="description">
<paragraph id="8BF1F629FFDBFFAAFF0FFAC9734A231E" blockId="2.[160,1156,1321,1727]" box="[160,294,1321,1346]" pageId="2" pageNumber="996">
<emphasis id="B93A2A3BFFDBFFAAFF0FFAC9734A231E" bold="true" box="[160,294,1321,1346]" italics="true" pageId="2" pageNumber="996">Description</emphasis>
</paragraph>
<paragraph id="8BF1F629FFDBFFABFF0FFAAD763D2478" blockId="2.[160,1156,1321,1727]" lastBlockId="3.[160,1157,142,548]" lastPageId="3" lastPageNumber="997" pageId="2" pageNumber="996">
<taxonomicName id="4C4E8DAAFFDBFFAAFF0FFAAD73122339" box="[160,382,1357,1381]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="996" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93A2A3BFFDBFFAAFF0FFAAD734B2339" box="[160,295,1357,1381]" italics="true" pageId="2" pageNumber="996">Paraegidium</emphasis>
species
</taxonomicName>
are relatively small beetles (body length 5.5–9.0 mm), brown to black coloured, without pattern, and densely punctate. Mandibles subsymmetrical, without lateral processes, protruding past anterior margin of frontoclypeus in dorsal view. Frontoclypeus in males with variably shaped bilobate anterior pronotal process, which is species-specific but subject to reasonable allometric variation. Frontoclypeus of females with slightly upturned anterior margin. Propleurae with carinae separating anterolateral areas from basal area. Pronotum of ‘major’ males with deep excavation in the middle and two horn-shaped lateral pronotal processes directed somewhat anteriorly. In ‘minor’ males, pronotal and head armature less developed: lateral pronotal processes are short, tubercle-shaped and anterior frontoclypeal process is carina-shaped. In females, pronotum is slightly depressed medially, without horns or tubercles. Scutellum narrowly rounded apically, about 1/10 length of elytra. Elytra convex, with marked humeral and apical umbones, without distinct striae or carinae, densely punctate with U-shaped punctures. Wings fully developed in all species. Metepisternon elongated, its posterior angle subtriangular and situated in distinct concavity of epipleuron. Mesocoxal cavities connected by a hole. Protibiae with 3 strong outer teeth in both sexes, without a smaller medial tooth. Procoxae without cavities. Mesotibiae without a tuft of setae ventroapically. Phallobase tube-shaped with strongly sclerotized ventral side but without differentiation of ventral and dorsal sclerites; ventroapical plate absent. Parameres with relatively simple apices, apices curved downwards and tapering, without setae or processes. Extreme apex of parameres with minute teeth laterally, visible in some species, almost indistinguishable in others. Endophallus with feebly developed armature. Subcoxites with sparse, long setae medioapically. Coxites triangular, with row of short spinules mediobasally and long, sparse setae apically. Stili not separated from coxites.
</paragraph>
</subSubSection>
<subSubSection id="C354A5A2FFDAFFABFF0FFDB173B223F7" pageId="3" pageNumber="997" type="diagnosis">
<paragraph id="8BF1F629FFDAFFABFF0FFDB173782436" blockId="3.[160,1155,593,826]" box="[160,276,593,618]" pageId="3" pageNumber="997">
<emphasis id="B93A2A3BFFDAFFABFF0FFDB173782436" bold="true" box="[160,276,593,618]" italics="true" pageId="3" pageNumber="997">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BF1F629FFDAFFABFF0FFD9476002566" blockId="3.[160,1155,593,826]" pageId="3" pageNumber="997">
<taxonomicName id="4C4E8DAAFFDAFFABFF0FFD9473EE24D0" box="[160,386,628,652]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="997" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93A2A3BFFDAFFABFF0FFD94734624D0" box="[160,298,628,652]" italics="true" pageId="3" pageNumber="997">Paraegidium</emphasis>
species
</taxonomicName>
can easily be distinguished from other members of the Aegidiini by the dense, irregular punctation and dense pubescence of the dorsal side of the body and by a bilobate or bifurcated anterior clypeal process in males. The peculiar U-shaped elytral punctures in
<taxonomicName id="4C4E8DAAFFDAFFABFE2CFD3C706124A8" box="[387,525,732,756]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="997" phylum="Arthropoda" rank="genus">
<emphasis id="B93A2A3BFFDAFFABFE2CFD3C706124A8" box="[387,525,732,756]" italics="true" pageId="3" pageNumber="997">Paraegidium</emphasis>
</taxonomicName>
are similar to those in
<taxonomicName id="4C4E8DAAFFDAFFABFCBCFD3C763F24A8" authority="Karsch" authorityName="Karsch" authorityYear="1881" box="[787,1107,732,756]" class="Insecta" family="Orphnidae" genus="Stenosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="997" phylum="Arthropoda" rank="species" species="costatus">
<emphasis id="B93A2A3BFFDAFFABFCBCFD3C766C24A8" box="[787,1024,732,756]" italics="true" pageId="3" pageNumber="997">Stenosternus costatus</emphasis>
Karsch
</taxonomicName>
, the only member of the West African orphnine genus
<taxonomicName id="4C4E8DAAFFDAFFABFD4AFD1F71A9254B" authority="Karsch" authorityName="Karsch" authorityYear="1881" box="[741,965,767,791]" class="Insecta" family="Orphnidae" genus="Stenosternus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="997" phylum="Arthropoda" rank="genus">
<emphasis id="B93A2A3BFFDAFFABFD4AFD1F7103254B" box="[741,879,767,791]" italics="true" pageId="3" pageNumber="997">Stenosternus</emphasis>
Karsch
</taxonomicName>
, although in the latter the punctures are directed cranially (
<bibRefCitation id="EFDF8BD8FFDAFFABFD28FCC271632566" author="Frolov AV" box="[647,783,802,826]" pageId="3" pageNumber="997" pagination="33 - 46" refId="ref7961" refString="Frolov AV. 2013. Stenosternus Karsch, a possible link between Neotropical and Afrotropical Orphninae (Coleoptera, Scarabaeidae). Zookeys. 335: 33 - 46." type="journal article" year="2013">Frolov 2013</bibRefCitation>
;
<bibRefCitation id="EFDF8BD8FFDAFFABFCB2FCC2760C2566" author="Frolov AV &amp; Akhmetova LA" box="[797,1120,802,826]" pageId="3" pageNumber="997" pagination="440 - 444" refId="ref7991" refString="Frolov AV, Akhmetova LA. 2015. Rediscovery of the enigmatic Stenosternus costatus Karsch (Coleoptera: Scarabaeidae: Orphninae) from Sao Tome island. Zootaxa. 4007: 440 - 444." type="journal article" year="2015">Frolov and Akhmetova 2015</bibRefCitation>
).
</paragraph>
<paragraph id="8BF1F629FFDAFFABFF0FFC87707325DC" blockId="3.[160,1157,871,1208]" box="[160,543,871,896]" pageId="3" pageNumber="997">
<emphasis id="B93A2A3BFFDAFFABFF0FFC87707325DC" bold="true" box="[160,543,871,896]" italics="true" pageId="3" pageNumber="997">Diagnostic characters of species</emphasis>
</paragraph>
<paragraph id="8BF1F629FFDAFFABFF0FFC6A760B22E4" blockId="3.[160,1157,871,1208]" pageId="3" pageNumber="997">
Species of
<taxonomicName id="4C4E8DAAFFDAFFABFEBEFC6A73F925FE" box="[273,405,906,930]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="997" phylum="Arthropoda" rank="genus">
<emphasis id="B93A2A3BFFDAFFABFEBEFC6A73F925FE" box="[273,405,906,930]" italics="true" pageId="3" pageNumber="997">Paraegidium</emphasis>
</taxonomicName>
can be separated by the combination of the shape and relative length of the parameres and the shape of the anterior frontoclypeal process in males. The endophallic armature is also specific in some species although it may be not reliable in most cases. Characters of the females are not diagnostic; therefore identification of single females is not possible at this stage. It should be noted that the photos taken from the dry aedeagi do not perfectly represent the shape and specifically the relative position of the phallobase and parameres. When the aedeagus dries out, the parameres bend downwards to various degree depending on the sclerotization of the base of the parameres and content of the phallobase (whether it has the endophallus). The shape of the aedeagi is more consistent in glycerol.
</paragraph>
<paragraph id="8BF1F629FFDAFFABFF0FFB05703622A2" blockId="3.[160,1155,1253,1451]" box="[160,602,1253,1278]" pageId="3" pageNumber="997">
<emphasis id="B93A2A3BFFDAFFABFF0FFB05703622A2" bold="true" box="[160,602,1253,1278]" italics="true" pageId="3" pageNumber="997">Species composition and distribution</emphasis>
</paragraph>
<paragraph id="8BF1F629FFDAFFABFF0FFAE973B223F7" blockId="3.[160,1155,1253,1451]" pageId="3" pageNumber="997">
The genus
<taxonomicName id="4C4E8DAAFFDAFFABFE8BFAE973C2237D" box="[292,430,1289,1313]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="997" phylum="Arthropoda" rank="genus">
<emphasis id="B93A2A3BFFDAFFABFE8BFAE973C2237D" box="[292,430,1289,1313]" italics="true" pageId="3" pageNumber="997">Paraegidium</emphasis>
</taxonomicName>
was previously known from the Atlantic Forest and southern Amazon regions. Judging from the material available to us, the genus range covers most of the central part of South America from Guiana Shield in the north to the Peruvian Yungas in the west and to extreme southeastern
<collectingCountry id="F359B6B9FFDAFFABFD7CFA91717E23D5" box="[723,786,1393,1417]" name="Brazil" pageId="3" pageNumber="997">Brazil</collectingCountry>
. The majority of the localities lie within the Brazilian Plateau.
</paragraph>
</subSubSection>
<subSubSection id="C354A5A2FFDAFFACFF0FFA0B76E825FA" lastPageId="4" lastPageNumber="998" pageId="3" pageNumber="997" type="key">
<paragraph id="8BF1F629FFDAFFABFF0FFA0B7027205F" blockId="3.[160,587,1514,1539]" box="[160,587,1514,1539]" pageId="3" pageNumber="997">
<emphasis id="B93A2A3BFFDAFFABFF0FFA0B7027205F" bold="true" box="[160,587,1514,1539]" pageId="3" pageNumber="997">
Key to
<taxonomicName id="4C4E8DAAFFDAFFABFF58FA0A7380205F" box="[247,492,1514,1539]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="997" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93A2A3BFFDAFFABFF58FA0A73E0205F" bold="true" box="[247,396,1514,1539]" italics="true" pageId="3" pageNumber="997">Paraegidium</emphasis>
species
</taxonomicName>
(males)
</emphasis>
</paragraph>
<paragraph id="8BF1F629FFDAFFABFF0FF9FF76EF2021" blockId="3.[160,1156,1567,1727]" pageId="3" pageNumber="997">
<keyLead id="30BF551CFFDAFFABFF0FF9FF76EF2021" pageId="3" pageNumber="997">
<keyStep id="30BAEE8CFFDAFFABFF0FF9FF76EF2021" pageId="3" pageNumber="997">
<key id="FFDFA118FFDAFFABFF0FF9FF76EF2021" pageId="3" pageNumber="997">
1. Parameres longer: ratio phallobase length/paramere length 2.2–1.9 (
<figureCitation id="1375EAACFFDAFFABFC58F9FF7632206B" box="[1015,1118,1567,1591]" captionStart="Figure 1" captionStartId="5.[160,225,1553,1576]" captionTargetBox="[168,1150,144,1522]" captionTargetId="figure-1@5.[163,1153,142,1522]" captionTargetPageId="5" captionText="Figure 1. Paraegidium monneorum, holotype male. Habitus in dorsal view (a); endophallus (b); aedeagus in lateral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.5181245" httpUri="https://zenodo.org/record/5181245/files/figure.png" pageId="3" pageNumber="997">Figures 1</figureCitation>
(c), 2(f), 3(f), 4(e)). Lobes of anterior frontoclypeal process angulate, rounded or truncated but not bimodal or bifurcated (
<figureCitation id="1375EAACFFDAFFABFD2FF985708F2021" box="[640,739,1637,1661]" captionStart="Figure 1" captionStartId="5.[160,225,1553,1576]" captionTargetBox="[168,1150,144,1522]" captionTargetId="figure-1@5.[163,1153,142,1522]" captionTargetPageId="5" captionText="Figure 1. Paraegidium monneorum, holotype male. Habitus in dorsal view (a); endophallus (b); aedeagus in lateral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.5181245" httpUri="https://zenodo.org/record/5181245/files/figure.png" pageId="3" pageNumber="997">Figures 1</figureCitation>
(a), 2(a, d, e), 3(a, c, d), 4(a, d)........ 2
</key>
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BF1F629FFDAFFACFF0FF96476EF2695" blockId="3.[160,1156,1567,1727]" lastBlockId="4.[210,1155,142,201]" lastPageId="4" lastPageNumber="998" pageId="3" pageNumber="997">
<keyLead id="30BF551CFFDAFFABFF0FF96476EF20E3" pageId="3" pageNumber="997">
– Parameres shorter: ratio phallobase length/paramere length 2.5–2.7 (
<figureCitation id="1375EAACFFDAFFABFC48F965763D20C1" box="[999,1105,1669,1693]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (c–e, h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="3" pageNumber="997">Figures 5</figureCitation>
(f), 6 (d)). Lobes of anterior frontoclypeal process bimodal or bifurcated (
<figureCitation id="1375EAACFFDAFFABFC68F947764120E3" box="[967,1069,1703,1727]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (c–e, h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="3" pageNumber="997">Figures 5</figureCitation>
(a, c, d),
</keyLead>
6(a–c)) (may appear truncated in larger males depending on angle of view,
<figureCitation id="1375EAACFFDDFFACFB88FF6E76EF26FA" box="[1063,1155,142,166]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (c–e, h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="4" pageNumber="998">Figure 5</figureCitation>
(a)). .............................................................................................................................................................. 5
</paragraph>
<key id="FFDFA118FFDDFFACFF0FFF0976E825FA" pageId="4" pageNumber="998">
<keyStep id="30BAEE8CFFDDFFACFF0FFF0976EF27D5" pageId="4" pageNumber="998">
<paragraph id="8BF1F629FFDDFFACFF0FFF0976E8277F" blockId="4.[160,1156,233,393]" pageId="4" pageNumber="998">
<keyLead id="30BF551CFFDDFFACFF0FFF0976E8277F" pageId="4" pageNumber="998">
2. Parameres longer: ratio phallobase length/paramere length 1.9 (
<figureCitation id="1375EAACFFDDFFACFC1DFF097667275D" box="[946,1035,233,257]" captionStart="Figure 1" captionStartId="5.[160,225,1553,1576]" captionTargetBox="[168,1150,144,1522]" captionTargetId="figure-1@5.[163,1153,142,1522]" captionTargetPageId="5" captionText="Figure 1. Paraegidium monneorum, holotype male. Habitus in dorsal view (a); endophallus (b); aedeagus in lateral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.5181245" httpUri="https://zenodo.org/record/5181245/files/figure.png" pageId="4" pageNumber="998">Figure 1</figureCitation>
(c)). Lobes of anterior frontoclypeal process truncated (
<figureCitation id="1375EAACFFDDFFACFD65FEEB714F277F" box="[714,803,267,291]" captionStart="Figure 1" captionStartId="5.[160,225,1553,1576]" captionTargetBox="[168,1150,144,1522]" captionTargetId="figure-1@5.[163,1153,142,1522]" captionTargetPageId="5" captionText="Figure 1. Paraegidium monneorum, holotype male. Habitus in dorsal view (a); endophallus (b); aedeagus in lateral view (c); parameres in dorsal view (d)." figureDoi="http://doi.org/10.5281/zenodo.5181245" httpUri="https://zenodo.org/record/5181245/files/figure.png" pageId="4" pageNumber="998">Figure 1</figureCitation>
(a)).......
<taxonomicName id="4C4E8DAAFFDDFFACFC2EFEEB764E277F" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[897,1058,267,291]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="998" phylum="Arthropoda" rank="species" species="monneorum" status="sp. nov.">
<emphasis id="B93A2A3BFFDDFFACFC2EFEEB764E277F" box="[897,1058,267,291]" italics="true" pageId="4" pageNumber="998">P. monneorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFDDFFACFB84FEEB76E8277F" box="[1067,1156,267,291]" pageId="4" pageNumber="998" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF1F629FFDDFFACFF0FFECB76EF27D5" blockId="4.[160,1156,233,393]" pageId="4" pageNumber="998">
<keyLead id="30BF551CFFDDFFACFF0FFECB76EF27D5" pageId="4" pageNumber="998">
– Parameres shorter: ratio phallobase length/paramere length 2.1–2.2 (
<figureCitation id="1375EAACFFDDFFACFC48FECB763D271F" box="[999,1105,299,323]" captionStart="Figure 2" captionStartId="6.[160,225,1492,1515]" captionTargetBox="[165,1152,143,1460]" captionTargetId="figure-1@6.[163,1153,142,1461]" captionTargetPageId="6" captionText="Figure 2. Paraegidium barretoi holotype male (a); paratype female (b); paratype males (c–h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181247" httpUri="https://zenodo.org/record/5181247/files/figure.png" pageId="4" pageNumber="998">Figures 2</figureCitation>
(f), 3 (f), 4(e)). Lobes of anterior frontoclypeal process rounded or angulate (
<figureCitation id="1375EAACFFDDFFACFC48FEAE7621273A" box="[999,1101,334,358]" captionStart="Figure 2" captionStartId="6.[160,225,1492,1515]" captionTargetBox="[165,1152,143,1460]" captionTargetId="figure-1@6.[163,1153,142,1461]" captionTargetPageId="6" captionText="Figure 2. Paraegidium barretoi holotype male (a); paratype female (b); paratype males (c–h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181247" httpUri="https://zenodo.org/record/5181247/files/figure.png" pageId="4" pageNumber="998">Figures 2</figureCitation>
(a), 3 (a), 4(a))..................................................................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAEE8CFFDDFFACFF0FFE4976EF247A" pageId="4" pageNumber="998">
<paragraph id="8BF1F629FFDDFFACFF0FFE4976E827BF" blockId="4.[160,1156,425,550]" pageId="4" pageNumber="998">
<keyLead id="30BF551CFFDDFFACFF0FFE4976E827BF" pageId="4" pageNumber="998">
3. Base of parameres wider in lateral view (
<figureCitation id="1375EAACFFDDFFACFD07FE49716B279D" box="[680,775,425,449]" captionStart="Figure 2" captionStartId="6.[160,225,1492,1515]" captionTargetBox="[165,1152,143,1460]" captionTargetId="figure-1@6.[163,1153,142,1461]" captionTargetPageId="6" captionText="Figure 2. Paraegidium barretoi holotype male (a); paratype female (b); paratype males (c–h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181247" httpUri="https://zenodo.org/record/5181247/files/figure.png" pageId="4" pageNumber="998">Figure 2</figureCitation>
(f)). Apices of parameres wider in dorsal view (
<figureCitation id="1375EAACFFDDFFACFECCFE2B73AE27BF" box="[355,450,459,483]" captionStart="Figure 2" captionStartId="6.[160,225,1492,1515]" captionTargetBox="[165,1152,143,1460]" captionTargetId="figure-1@6.[163,1153,142,1461]" captionTargetPageId="6" captionText="Figure 2. Paraegidium barretoi holotype male (a); paratype female (b); paratype males (c–h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181247" httpUri="https://zenodo.org/record/5181247/files/figure.png" pageId="4" pageNumber="998">Figure 2</figureCitation>
(g)).................................................................................
<taxonomicName id="4C4E8DAAFFDDFFACFC1EFE2B764E27BF" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[945,1058,459,483]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="998" phylum="Arthropoda" rank="species" species="barretoi" status="sp. nov.">
<emphasis id="B93A2A3BFFDDFFACFC1EFE2B764E27BF" box="[945,1058,459,483]" italics="true" pageId="4" pageNumber="998">P. barretoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFDDFFACFB84FE2B76E827BF" box="[1067,1156,459,483]" pageId="4" pageNumber="998" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF1F629FFDDFFACFF0FFE0B76EF247A" blockId="4.[160,1156,425,550]" pageId="4" pageNumber="998">
<keyLead id="30BF551CFFDDFFACFF0FFE0B76EF247A" pageId="4" pageNumber="998">
– Base of parameres narrower in lateral view (
<figureCitation id="1375EAACFFDDFFACFD62FE0B715A245F" box="[717,822,491,515]" captionStart="Figure 3" captionStartId="7.[160,225,1308,1331]" captionTargetBox="[166,1153,142,1264]" captionTargetId="figure-74@7.[163,1153,142,1277]" captionTargetPageId="7" captionText="Figure 3. Paraegidium costalimai male (a, c–h); female (b). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181249" httpUri="https://zenodo.org/record/5181249/files/figure.png" pageId="4" pageNumber="998">Figures 3</figureCitation>
(f), 4(e)). Apices of parameres narrower in dorsal view (
<figureCitation id="1375EAACFFDDFFACFE40FDEE7035247A" box="[495,601,526,550]" captionStart="Figure 3" captionStartId="7.[160,225,1308,1331]" captionTargetBox="[166,1153,142,1264]" captionTargetId="figure-74@7.[163,1153,142,1277]" captionTargetPageId="7" captionText="Figure 3. Paraegidium costalimai male (a, c–h); female (b). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181249" httpUri="https://zenodo.org/record/5181249/files/figure.png" pageId="4" pageNumber="998">Figures 3</figureCitation>
(g), 4(f))............................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAEE8CFFDDFFACFF0FFDA676E8249F" pageId="4" pageNumber="998">
<paragraph id="8BF1F629FFDDFFACFF0FFDA676EF24DD" blockId="4.[160,1156,582,707]" pageId="4" pageNumber="998">
<keyLead id="30BF551CFFDDFFACFF0FFDA676EF24DD" pageId="4" pageNumber="998">
4. Lobes of anterior frontoclypeal process angulate (
<figureCitation id="1375EAACFFDDFFACFD51FDA671362402" box="[766,858,582,606]" captionStart="Figure 3" captionStartId="7.[160,225,1308,1331]" captionTargetBox="[166,1153,142,1264]" captionTargetId="figure-74@7.[163,1153,142,1277]" captionTargetPageId="7" captionText="Figure 3. Paraegidium costalimai male (a, c–h); female (b). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181249" httpUri="https://zenodo.org/record/5181249/files/figure.png" pageId="4" pageNumber="998">Figure 3</figureCitation>
(a, c, d)). Group of spinules in endophallus larger and elongated (
<figureCitation id="1375EAACFFDDFFACFD2CFD89708D24DD" box="[643,737,617,641]" captionStart="Figure 3" captionStartId="7.[160,225,1308,1331]" captionTargetBox="[166,1153,142,1264]" captionTargetId="figure-74@7.[163,1153,142,1277]" captionTargetPageId="7" captionText="Figure 3. Paraegidium costalimai male (a, c–h); female (b). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181249" httpUri="https://zenodo.org/record/5181249/files/figure.png" pageId="4" pageNumber="998">Figure 3</figureCitation>
(h)).........................................
<taxonomicName id="4C4E8DAAFFDDFFACFC57FD8976EF24DD" box="[1016,1155,617,641]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="998" phylum="Arthropoda" rank="species" species="costalimai">
<emphasis id="B93A2A3BFFDDFFACFC57FD8976EF24DD" box="[1016,1155,617,641]" italics="true" pageId="4" pageNumber="998">P. costalimai</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF1F629FFDDFFACFF0FFD6876E8249F" blockId="4.[160,1156,582,707]" pageId="4" pageNumber="998">
<keyLead id="30BF551CFFDDFFACFF0FFD6876E8249F" pageId="4" pageNumber="998">
– Lobes of anterior frontoclypeal process rounded (
<figureCitation id="1375EAACFFDDFFACFD54FD68713B24FC" box="[763,855,648,672]" captionStart="Figure 4" captionStartId="8.[160,225,1414,1437]" captionTargetBox="[163,1153,143,1379]" captionTargetId="figure-39@8.[163,1153,142,1383]" captionTargetPageId="8" captionText="Figure 4. Paraegidium pokornyi, holotype male (a); paratype female (b); paratype males (c–g). Habitus in dorsal view (a, b); pronotum and head (c, d); aedeagus in lateral view (e); parameres in dorsal view (f); endophallus (g)." figureDoi="http://doi.org/10.5281/zenodo.5181251" httpUri="https://zenodo.org/record/5181251/files/figure.png" pageId="4" pageNumber="998">Figure 4</figureCitation>
(a, d)). Group of spinules in endophallus smaller (
<figureCitation id="1375EAACFFDDFFACFE69FD4B7049249F" box="[454,549,683,707]" captionStart="Figure 4" captionStartId="8.[160,225,1414,1437]" captionTargetBox="[163,1153,143,1379]" captionTargetId="figure-39@8.[163,1153,142,1383]" captionTargetPageId="8" captionText="Figure 4. Paraegidium pokornyi, holotype male (a); paratype female (b); paratype males (c–g). Habitus in dorsal view (a, b); pronotum and head (c, d); aedeagus in lateral view (e); parameres in dorsal view (f); endophallus (g)." figureDoi="http://doi.org/10.5281/zenodo.5181251" httpUri="https://zenodo.org/record/5181251/files/figure.png" pageId="4" pageNumber="998">Figure 4</figureCitation>
(g)).............................................................
<taxonomicName id="4C4E8DAAFFDDFFACFC07FD4B764F249F" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[936,1059,683,707]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="998" phylum="Arthropoda" rank="species" species="pokornyi" status="sp. nov.">
<emphasis id="B93A2A3BFFDDFFACFC07FD4B764F249F" box="[936,1059,683,707]" italics="true" pageId="4" pageNumber="998">P. pokornyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFDDFFACFB84FD4B76E8249F" box="[1067,1156,683,707]" pageId="4" pageNumber="998" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BAEE8CFFDDFFACFF0FFD0376E825FA" pageId="4" pageNumber="998">
<paragraph id="8BF1F629FFDDFFACFF0FFD0376E8251D" blockId="4.[160,1156,739,934]" pageId="4" pageNumber="998">
<keyLead id="30BF551CFFDDFFACFF0FFD0376E8251D" pageId="4" pageNumber="998">
5. Parameres longer: ratio phallobase length/paramere length 2.5 (
<figureCitation id="1375EAACFFDDFFACFC00FD03766224A7" box="[943,1038,739,763]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (c–e, h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="4" pageNumber="998">Figure 5</figureCitation>
(f)). Apices of parameres more or less acute in lateral view. Lobes of anterior frontoclypeal process bimodal (sinuate apically,
<figureCitation id="1375EAACFFDDFFACFDF6FCC970DB251D" box="[601,695,809,833]" captionStart="Figure 5" captionStartId="9.[160,225,1398,1421]" captionTargetBox="[163,1153,143,1367]" captionTargetId="figure-19@9.[163,1153,142,1367]" captionTargetPageId="9" captionText="Figure 5. Paraegidium howdeni, holotype male (a, f, g); paratype female (b); paratype males (c–e, h). Habitus in dorsal view (a, b); pronotum and head (c–e); aedeagus in lateral view (f); parameres in dorsal view (g); endophallus (h)." figureDoi="http://doi.org/10.5281/zenodo.5181253" httpUri="https://zenodo.org/record/5181253/files/figure.png" pageId="4" pageNumber="998">Figure 5</figureCitation>
(a, c, d)).........................
<taxonomicName id="4C4E8DAAFFDDFFACFC06FCC9764E251D" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[937,1058,809,833]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="998" phylum="Arthropoda" rank="species" species="howdeni" status="sp. nov.">
<emphasis id="B93A2A3BFFDDFFACFC06FCC9764E251D" box="[937,1058,809,833]" italics="true" pageId="4" pageNumber="998">P. howdeni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFDDFFACFB84FCC976E8251D" box="[1067,1156,809,833]" pageId="4" pageNumber="998" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF1F629FFDDFFACFF0FFCA876E825FA" blockId="4.[160,1156,739,934]" pageId="4" pageNumber="998">
<keyLead id="30BF551CFFDDFFACFF0FFCA876E825FA" pageId="4" pageNumber="998">
– Parameres shorter: ratio phallobase length/paramere length 2.7 (
<figureCitation id="1375EAACFFDDFFACFC04FCA87664253C" box="[939,1032,840,864]" captionStart="Figure 6" captionStartId="10.[160,225,1147,1170]" captionTargetBox="[165,1153,143,1115]" captionTargetId="figure-55@10.[163,1153,142,1116]" captionTargetPageId="10" captionText="Figure 6. Paraegidium oliveirai, holotype male (a, d–f); paratype males (b, c). Habitus in dorsal view (a); pronotum and head (b, c); aedeagus in lateral view (d); parameres in dorsal view (e); endophallus (f)." figureDoi="http://doi.org/10.5281/zenodo.5181255" httpUri="https://zenodo.org/record/5181255/files/figure.png" pageId="4" pageNumber="998">Figure 6</figureCitation>
(d)). Apices of parameres rounded in lateral view. Lobes of anterior frontoclypeal process bifurcated (
<figureCitation id="1375EAACFFDDFFACFEFAFC6E73DF25FA" box="[341,435,910,934]" captionStart="Figure 6" captionStartId="10.[160,225,1147,1170]" captionTargetBox="[165,1153,143,1115]" captionTargetId="figure-55@10.[163,1153,142,1116]" captionTargetPageId="10" captionText="Figure 6. Paraegidium oliveirai, holotype male (a, d–f); paratype males (b, c). Habitus in dorsal view (a); pronotum and head (b, c); aedeagus in lateral view (d); parameres in dorsal view (e); endophallus (f)." figureDoi="http://doi.org/10.5281/zenodo.5181255" httpUri="https://zenodo.org/record/5181255/files/figure.png" pageId="4" pageNumber="998">Figure 6</figureCitation>
(a–c))...............................................................................
<taxonomicName id="4C4E8DAAFFDDFFACFC1DFC6E764F25FA" authority="Frolov &amp; Akhmetova &amp; Vaz-de-Mello, 2017" authorityName="Frolov &amp; Akhmetova &amp; Vaz-de-Mello" authorityYear="2017" box="[946,1059,910,934]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="998" phylum="Arthropoda" rank="species" species="oliveirai" status="sp. nov.">
<emphasis id="B93A2A3BFFDDFFACFC1DFC6E764F25FA" box="[946,1059,910,934]" italics="true" pageId="4" pageNumber="998">P. oliveirai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2099740FFDDFFACFB84FC6E76E825FA" box="[1067,1156,910,934]" pageId="4" pageNumber="998" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>
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