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<document id="CB9E4A2FD9233ADCE8AB1E6D5369FCE2" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589942712" checkinUser="felipe" docAuthor="Vaz-De-Mello, Fernando Z." docDate="2008" docId="03D287CEFF93AF3DD1DBDFE3FE1A15B4" docLanguage="en" docName="zt01955p075.pdf" docOrigin="Zootaxa 1955" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Pedaridium Harold 1868" docType="treatment" docVersion="3" lastPageNumber="40" masterDocId="FFEBFFB6FFB4AF1AD14CDF74FFA61358" masterDocTitle="Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus" masterLastPageNumber="75" masterPageNumber="1" pageNumber="40" updateTime="1699081972826" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DB0E0F39D8B56BA61658931D97F85DA3">Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus</mods:title>
</mods:titleInfo>
<mods:name id="84879AFA2539BD87FFFC54D15A27C702" type="personal">
<mods:role id="BDADB5B577DBA17F5B2BDD2DE3B20B66">
<mods:roleTerm id="E3773088C896892D5A88A9648D6E2F3A">Author</mods:roleTerm>
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<mods:namePart id="A1623EB77B5AF7BBC47EB009A349FAA5">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="33DBED30F44F35E8E2431C30379D8B43">text</mods:typeOfResource>
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<mods:title id="CB63C3FC49BEE159052EB137651C84C6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="38D16A829F8FF16F946E74344DEC1607">
<mods:date id="C68E177F6C3AC96D06448DA461CE316F">2008</mods:date>
<mods:detail id="2AC3A613B70657B6B43EB1FB855A2EAE" type="pubDate">
<mods:number id="B2DF1C1ADF8F6CC6D6056848A412B794">2008-12-05</mods:number>
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<mods:detail id="0CB04103409D4291236D3CF4D8EF32B6" type="volume">
<mods:number id="CB8B88C00C41392ABDEE2D2B9B5A2012">1955</mods:number>
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<mods:start id="A7B00149CFBC0CF7A916132E28877C1E">1</mods:start>
<mods:end id="CF15DAA81FE959206B1C249FA836772C">75</mods:end>
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<mods:classification id="AC310DF9DFA90E93F39D4A053554FB0E">journal article</mods:classification>
<mods:identifier id="8505F90C12240EC9B9840CCABAAA6AD7" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="03D287CEFF93AF3DD1DBDFE3FE1A15B4" ID-DOI="http://doi.org/10.5281/zenodo.5242041" ID-Zenodo-Dep="5242041" LSID="urn:lsid:plazi:treatment:03D287CEFF93AF3DD1DBDFE3FE1A15B4" httpUri="http://treatment.plazi.org/id/03D287CEFF93AF3DD1DBDFE3FE1A15B4" lastPageNumber="40" pageId="39" pageNumber="40">
<subSubSection id="C3616553FF93AF3DD1DBDFE3FE5E13E9" box="[151,504,151,177]" pageId="39" pageNumber="40" type="nomenclature">
<paragraph id="8BC436D8FF93AF3DD1DBDFE3FE5E13E9" blockId="39.[151,504,151,177]" box="[151,504,151,177]" pageId="39" pageNumber="40">
<heading id="D08C81B4FF93AF3DD1DBDFE3FE5E13E9" bold="true" box="[151,504,151,177]" fontSize="11" level="1" pageId="39" pageNumber="40" reason="1">
<emphasis id="B90FEACAFF93AF3DD1DBDFE3FE5E13E9" bold="true" box="[151,504,151,177]" pageId="39" pageNumber="40">
11.
<taxonomicName id="4C7B4D5BFF93AF3DD18DDFE3FE5E13E9" ID-CoL="6JXZ" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[193,504,151,177]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF93AF3DD18DDFE3FEE813E9" bold="true" box="[193,334,151,177]" italics="true" pageId="39" pageNumber="40">Pedaridium</emphasis>
<bibRefCitation id="EFEA4B29FF93AF3DD01ADFE3FE5E13E9" author="Harold, E." box="[342,504,151,177]" pageId="39" pageNumber="40" pagination="33 - 55" refId="ref48745" refString="Harold, E. (1868) Die Choeridiiden-Gattungen Uroxys und Trichillum. Coleopterologische Hefte, 3, 33 - 55." type="journal article" year="1868">Harold, 1868</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF93AF3DD1DBDF96FEA312C4" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="8BC436D8FF93AF3DD1DBDF96FEA312C4" blockId="39.[151,1436,226,1772]" pageId="39" pageNumber="40">
<emphasis id="B90FEACAFF93AF3DD1DBDF96FEBF13A4" bold="true" box="[151,281,226,252]" pageId="39" pageNumber="40">Diagnosis:</emphasis>
Body very large (
<quantity id="4C839B3DFF93AF3DD0BCDF96FD2713A4" box="[496,641,226,252]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="8.6" metricValueMin="5.2" pageId="39" pageNumber="40" unit="mm" value="6.9" valueMax="8.6" valueMin="5.2">5.2–8.6 mm</quantity>
) and elongated (
<figureCitation id="13402A5DFF93AF3DD204DF96FC0713A4" box="[840,929,226,252]" captionStart="FIGURES 23–30" captionStartId="57.[151,269,1934,1958]" captionTargetBox="[222,1365,194,1910]" captionTargetId="figure-54@57.[222,1365,194,1911]" captionTargetPageId="57" captionText="FIGURES 23–30. Dorsal views. 23. Leotrichillum louzadaorum; 24–25: Martinezidium: 24. M. fulgens; 25. M. maya; 26. Nunoidium argentinum; 27. Onoreidium cristatum; 28. Onoreidium bottimeri; 29. Pedaridium hirsutum; 30. Pereiraidium almeidai, ♂." pageId="39" pageNumber="40">Fig. 29</figureCitation>
). Pronotum clearly flat to concave posteriorly (
<figureCitation id="13402A5DFF93AF3DD196DE7EFE93127C" box="[218,309,266,292]" captionStart="FIGURES 87–94" captionStartId="65.[151,269,1961,1985]" captionTargetBox="[210,1372,194,1934]" captionTargetId="figure-48@65.[210,1372,194,1938]" captionTargetPageId="65" captionText="FIGURES 87–94. 87. Pedaridium hirsutum, pronotum; 88–90: Pereiraidium almeidai: 88. ♂ head; 89. parameres; 90. ♀ head; 91–92: Silvinha unica: 91. head; 92. parameres; 93–94: Trichillidium quadridens: 93. head; 94. parameres." pageId="39" pageNumber="40">Fig. 87</figureCitation>
). Elytra tectiform (suture elevated in relation to disc, each elytron separately flattened), disc forming a weak fold at apex; interstriae with biseriate setose punctures (
<figureCitation id="13402A5DFF93AF3DD56FDE46FB271214" box="[1059,1153,306,332]" captionStart="FIGURES 23–30" captionStartId="57.[151,269,1934,1958]" captionTargetBox="[222,1365,194,1910]" captionTargetId="figure-54@57.[222,1365,194,1911]" captionTargetPageId="57" captionText="FIGURES 23–30. Dorsal views. 23. Leotrichillum louzadaorum; 24–25: Martinezidium: 24. M. fulgens; 25. M. maya; 26. Nunoidium argentinum; 27. Onoreidium cristatum; 28. Onoreidium bottimeri; 29. Pedaridium hirsutum; 30. Pereiraidium almeidai, ♂." pageId="39" pageNumber="40">Fig. 29</figureCitation>
). Males with proclaws strongly bent (Fig. 86). Scattered longer setae present in sides and anterior part of pronotum and elytral apex (
<figureCitation id="13402A5DFF93AF3DD1D3DEF6FF5112C4" box="[159,247,386,412]" captionStart="FIGURES 23–30" captionStartId="57.[151,269,1934,1958]" captionTargetBox="[222,1365,194,1910]" captionTargetId="figure-54@57.[222,1365,194,1911]" captionTargetPageId="57" captionText="FIGURES 23–30. Dorsal views. 23. Leotrichillum louzadaorum; 24–25: Martinezidium: 24. M. fulgens; 25. M. maya; 26. Nunoidium argentinum; 27. Onoreidium cristatum; 28. Onoreidium bottimeri; 29. Pedaridium hirsutum; 30. Pereiraidium almeidai, ♂." pageId="39" pageNumber="40">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF93AF3DD18ADEDEFB591604" pageId="39" pageNumber="40" type="description">
<paragraph id="8BC436D8FF93AF3DD18ADEDEFC0A17BC" blockId="39.[151,1436,226,1772]" pageId="39" pageNumber="40">
<emphasis id="B90FEACAFF93AF3DD18ADEDEFEC6129C" bold="true" box="[198,352,426,452]" pageId="39" pageNumber="40">Description:</emphasis>
Length
<quantity id="4C839B3DFF93AF3DD089DEDEFDF1129C" box="[453,599,426,452]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="8.6" metricValueMin="5.2" pageId="39" pageNumber="40" unit="mm" value="6.9" valueMax="8.6" valueMin="5.2">5.2–8.6 mm</quantity>
, body very elongated and parallel-sided (
<figureCitation id="13402A5DFF93AF3DD50DDEDEFB3F129C" box="[1089,1177,426,452]" captionStart="FIGURES 23–30" captionStartId="57.[151,269,1934,1958]" captionTargetBox="[222,1365,194,1910]" captionTargetId="figure-54@57.[222,1365,194,1911]" captionTargetPageId="57" captionText="FIGURES 23–30. Dorsal views. 23. Leotrichillum louzadaorum; 24–25: Martinezidium: 24. M. fulgens; 25. M. maya; 26. Nunoidium argentinum; 27. Onoreidium cristatum; 28. Onoreidium bottimeri; 29. Pedaridium hirsutum; 30. Pereiraidium almeidai, ♂." pageId="39" pageNumber="40">Fig. 29</figureCitation>
), color brown to dark grey, completely opaque and lacking metallic sheen, with yellow to orange setae. Clypeus with two teeth, separated by wide shallow emargination rounded at bottom; laterally unequally rounded, mesally strongly so and laterally straight, continuous to genal margin. Head flat, feebly depressed near eyes. Clypeofrontal and clypeogenal sutures indistinct. Eyes, in dorsal view, as wide as half their length; interocular width ten to 15 times eye width (Fig. 84). Pronotum lacking anterior and posterior beads, separated from hypomeron by distinct longitudinal carina; disc covered by setose puntures, setae much longer anteriorly and laterally; disc surface with posterior shallow to deep concavity, projected anteriorly along midline; and slightly expanded (shallowly concave) on anterior angles. Pronotal lateral outline sinuate, wider at anterior angles than at middle (
<figureCitation id="13402A5DFF93AF3DD47BDD9EFA29105C" box="[1335,1423,746,772]" captionStart="FIGURES 87–94" captionStartId="65.[151,269,1961,1985]" captionTargetBox="[210,1372,194,1934]" captionTargetId="figure-48@65.[210,1372,194,1938]" captionTargetPageId="65" captionText="FIGURES 87–94. 87. Pedaridium hirsutum, pronotum; 88–90: Pereiraidium almeidai: 88. ♂ head; 89. parameres; 90. ♀ head; 91–92: Silvinha unica: 91. head; 92. parameres; 93–94: Trichillidium quadridens: 93. head; 94. parameres." pageId="39" pageNumber="40">Fig. 87</figureCitation>
). Hypomeron with distinct longitudinal carina; mesoepimeron with distinct transverse anterior carina. Elytral discal interstriae with two rows of setiferous punctures, flat. Elytra tectiform for entire length, with sutural interstria clearly elevated and each elytron flat on disc; apically with lateral fold between declivity and pseudoepipleuron. Striae clearly deeper and wider at apex. Pseudoepipleura gradually narrowed to apex (
<figureCitation id="13402A5DFF93AF3DD408DCFEFA2A10FC" box="[1348,1420,906,932]" captionStart="FIGURES 5–11" captionStartId="54.[151,269,1929,1953]" captionTargetBox="[223,1320,197,1895]" captionTargetId="figure-60@54.[203,1372,197,1917]" captionTargetPageId="54" captionText="FIGURES 5–11. Left epipleuron and pseudoepipleuron, ventral view. 5. Trichillum externepunctatum; 6. Onoreidium cristatum; 7. Scatrichus bicarinatus; 8. Pedaridium hirsutum; 9. Silvinha unica; 10. Besourenga minutus; 11. Degallieridium lilliputanum. Scale = 1.0 mm." pageId="39" pageNumber="40">Fig. 8</figureCitation>
), with row of large setiferous punctures along entire length and smaller irregular setae in the anterior half. Protibiae with three teeth distributed along apical half of lateral margin, basally denticulate and with small scale-like setae on the ventral face of teeth only. Mesofemora and metafemora elongate, mesotibiae and metatibiae four times as long as wide apically. Pygidium transverse, vertical, last abdominal sternite long. Phallobase very thin, parameres as long as third of phallobase length, apically flattened (Fig. 85). Internal sac with very thin canaliculated straight pseudoflagellum, with basal curved auxiliar elongated lamella. Spermatheca C-shaped on apical half, gradually narrowed apically, basally strongly narrowed, pointed (
<figureCitation id="13402A5DFF93AF3DD587DBD6FA8217E4" box="[1227,1316,1186,1212]" captionStart="FIGURES 12–14" captionStartId="55.[151,269,1373,1397]" captionTargetBox="[173,1369,194,1301]" captionTargetId="figure-46@55.[169,1419,194,1350]" captionTargetPageId="55" captionText="FIGURES 12–14. Spermatheca (receptaculum seminis), and attached part of spermathecal duct, and spermathecal gland; 12. Pedaridium hirsutum; 13. Trichillum externepunctatum; 14. Eutrichillum hirsutum. Scale = 0.2 mm." pageId="39" pageNumber="40">Fig. 12</figureCitation>
), duct not sclerotized. Coxites minute, symmetrical and multidentate apically.
</paragraph>
<paragraph id="8BC436D8FF93AF3DD18ADB86FB591604" blockId="39.[151,1436,226,1772]" pageId="39" pageNumber="40">
<emphasis id="B90FEACAFF93AF3DD18ADB86FE611654" bold="true" box="[198,455,1266,1292]" pageId="39" pageNumber="40">Sexual dimorphism:</emphasis>
Males have proclaws strongly bent (Fig. 86), a very strong mesoapical protibial tooth; small mesotibial and metatibial mesoapical teeth (
<figureCitation id="13402A5DFF93AF3DD27CDA6EFC07166C" box="[816,929,1306,1332]" captionStart="FIGURES 1–4" captionStartId="53.[151,269,1431,1455]" captionTargetBox="[175,1415,182,1420]" captionTargetId="figure-36@53.[175,1415,182,1420]" captionTargetPageId="53" captionText="FIGURES 1–4. Pedaridium hirsutum, apex of right anterior tibia and tarsus, respectively dorsal and ventral views: 1–2. ♂; 3–4. ♀." pageId="39" pageNumber="40">Figs. 1–2</figureCitation>
); metasternal disc strongly concave (flat in females), abdominal disc flat (convex in females), and less transverse pygidium than in females.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF93AF3DD18ADA1EFA3316DC" box="[198,1429,1386,1412]" pageId="39" pageNumber="40" type="type_taxon">
<paragraph id="8BC436D8FF93AF3DD18ADA1EFA3316DC" blockId="39.[151,1436,226,1772]" box="[198,1429,1386,1412]" pageId="39" pageNumber="40">
<emphasis id="B90FEACAFF93AF3DD18ADA1EFECC16DC" bold="true" box="[198,362,1386,1412]" pageId="39" pageNumber="40">
<typeStatus id="54C0887AFF93AF3DD18ADA1EFEA516DC" box="[198,259,1386,1412]" pageId="39" pageNumber="40">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C7B4D5BFF93AF3DD03EDA1FFD6A16DC" ID-CoL="75X3D" authority="Harold, 1859" authorityName=": Harold" authorityYear="1859" box="[370,716,1386,1412]" class="Insecta" family="Scarabaeidae" genus="Pedaria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hirsuta">
<emphasis id="B90FEACAFF93AF3DD03EDA1FFD8C16DC" box="[370,554,1387,1412]" italics="true" pageId="39" pageNumber="40">Pedaria hirsuta</emphasis>
<bibRefCitation id="EFEA4B29FF93AF3DD37EDA1EFD6A16DC" author="Harold, E." box="[562,716,1386,1412]" pageId="39" pageNumber="40" pagination="193 - 224" refId="ref48671" refString="Harold, E. (1859) Beitrage zur Kenntnis einiger coprophagen Lamellicornien. Berliner Entomologische Zeitschrift, 3, 193 - 224." type="journal article" year="1859">Harold, 1859</bibRefCitation>
</taxonomicName>
(implicit monotypy) =
<taxonomicName id="4C7B4D5BFF93AF3DD293DA1FFA3716DC" ID-CoL="6TX3Q" authority="(Harold, 1859)" baseAuthorityName="Harold" baseAuthorityYear="1859" box="[991,1425,1386,1412]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="hirsutum">
<emphasis id="B90FEACAFF93AF3DD293DA1FFB7E16DC" box="[991,1240,1387,1412]" italics="true" pageId="39" pageNumber="40">Pedaridium hirsutum</emphasis>
(
<bibRefCitation id="EFEA4B29FF93AF3DD5ABDA1EFA2E16DC" author="Harold, E." box="[1255,1416,1386,1412]" pageId="39" pageNumber="40" pagination="193 - 224" refId="ref48671" refString="Harold, E. (1859) Beitrage zur Kenntnis einiger coprophagen Lamellicornien. Berliner Entomologische Zeitschrift, 3, 193 - 224." type="journal article" year="1859">Harold, 1859</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF93AF3DD18ADAE6FB69168C" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8BC436D8FF93AF3DD18ADAE6FB69168C" blockId="39.[151,1436,226,1772]" pageId="39" pageNumber="40">
<emphasis id="B90FEACAFF93AF3DD18ADAE6FECF16F4" bold="true" box="[198,361,1426,1452]" pageId="39" pageNumber="40">Distribution:</emphasis>
<collectingCountry id="F36C7648FF93AF3DD038DAE6FE1D16F4" box="[372,443,1426,1452]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
(
<collectingRegion id="49BFF83AFF93AF3DD081DAE6FDD416F4" box="[461,626,1426,1452]" country="Brazil" name="Minas Gerais" pageId="39" pageNumber="40">Minas Gerais</collectingRegion>
,
<taxonomicName id="4C7B4D5BFF93AF3DD3CDDAE6FD5B16F4" authority="Paulo" authorityName="Paulo" box="[641,765,1426,1452]" class="Gastropoda" family="Retusidae" genus="Sao" higherTaxonomySource="CoL" kingdom="Animalia" order="Cephalaspidea" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">São Paulo</taxonomicName>
,
<collectingRegion id="49BFF83AFF93AF3DD240DAE6FC1B16F4" box="[780,957,1426,1452]" country="Brazil" name="Rio de Janeiro" pageId="39" pageNumber="40">Rio de Janeiro</collectingRegion>
, and
<collectingRegion id="49BFF83AFF93AF3DD54DDAE6FBF616F4" box="[1025,1104,1426,1452]" country="Brazil" name="Parana" pageId="39" pageNumber="40">Paraná</collectingRegion>
states). Present only in the Eastern
<collectingRegion id="49BFF83AFF93AF3DD1BADACEFEE3168C" box="[246,325,1466,1492]" country="Brazil" name="Parana" pageId="39" pageNumber="40">Parana</collectingRegion>
subregion, in the
<collectingRegion id="49BFF83AFF93AF3DD354DACEFDCE168C" box="[536,616,1466,1492]" country="Brazil" name="Parana" pageId="39" pageNumber="40">Parana</collectingRegion>
Forest and
<taxonomicName id="4C7B4D5BFF93AF3DD3BEDACFFBE8168C" authority="Forest" authorityName="Forest" box="[754,1102,1466,1492]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="39" pageNumber="40" phylum="Tracheophyta" rank="species" species="angustifolia">
<emphasis id="B90FEACAFF93AF3DD3BEDACFFC58168C" box="[754,1022,1467,1492]" italics="true" pageId="39" pageNumber="40">Araucaria angustifolia</emphasis>
Forest
</taxonomicName>
provinces.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF93AF3DD18ADA96FC52159C" pageId="39" pageNumber="40" type="discussion">
<paragraph id="8BC436D8FF93AF3DD18ADA96FB03152C" blockId="39.[151,1436,226,1772]" pageId="39" pageNumber="40">
<emphasis id="B90FEACAFF93AF3DD18ADA96FEE716A4" bold="true" box="[198,321,1506,1532]" pageId="39" pageNumber="40">Remarks:</emphasis>
<taxonomicName id="4C7B4D5BFF93AF3DD005DA97FD8816A4" box="[329,558,1506,1532]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90FEACAFF93AF3DD005DA97FE7516A4" box="[329,467,1507,1532]" italics="true" pageId="39" pageNumber="40">Pedaridium</emphasis>
species
</taxonomicName>
are the only representatives of the group that really have a body shape similar to that of species in the African genus
<taxonomicName id="4C7B4D5BFF93AF3DD3D9D97FFD55157C" authorityName="Laporte" authorityYear="1832" box="[661,755,1547,1572]" class="Insecta" family="Scarabaeidae" genus="Pedaria" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF93AF3DD3D9D97FFD55157C" box="[661,755,1547,1572]" italics="true" pageId="39" pageNumber="40">Pedaria</emphasis>
</taxonomicName>
. Synapomorphies shared by species of this genus are the unique form of the pronotum and elytra; dual-length dorsal pilosity; reduction of number of internal sac sclerites; reduction of coxite size; and elongation of phallobase (see also under
<taxonomicName id="4C7B4D5BFF93AF3DD54DD92FFB33152C" authorityName="Vaz-De-Mello" authorityYear="2008" box="[1025,1173,1627,1652]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF93AF3DD54DD92FFB33152C" box="[1025,1173,1627,1652]" italics="true" pageId="39" pageNumber="40">Genieridium</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC436D8FF93AF3DD18AD9F6FC52159C" blockId="39.[151,1436,226,1772]" pageId="39" pageNumber="40">
<emphasis id="B90FEACAFF93AF3DD18AD9F6FEC915C4" bold="true" box="[198,367,1666,1692]" pageId="39" pageNumber="40">Composition:</emphasis>
In the present restricted sense,
<taxonomicName id="4C7B4D5BFF93AF3DD3AFD9F7FCCB15C4" authorityName="Harold" authorityYear="1868" box="[739,877,1667,1692]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF93AF3DD3AFD9F7FCCB15C4" box="[739,877,1667,1692]" italics="true" pageId="39" pageNumber="40">Pedaridium</emphasis>
</taxonomicName>
is monotypic. However, it contains at least one undescribed species currently under study, in addition to the
<typeStatus id="54C0887AFF93AF3DD212D9DEFC36159C" box="[862,912,1706,1732]" pageId="39" pageNumber="40">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF93AF3DD18AD9A6FE1A15B4" box="[198,444,1746,1772]" pageId="39" pageNumber="40" type="materials_examined">
<paragraph id="8BC436D8FF93AF3DD18AD9A6FE1A15B4" blockId="39.[151,1436,226,1772]" box="[198,444,1746,1772]" pageId="39" pageNumber="40">
<emphasis id="B90FEACAFF93AF3DD18AD9A6FE1A15B4" bold="true" box="[198,444,1746,1772]" pageId="39" pageNumber="40">Material examined:</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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