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<document ID-DOI="10.11646/zootaxa.5250.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="7705657" ID-ZooBank="4729EAF3-8B78-4AAD-B373-9179D008F239" approvalRequired="130" approvalRequired_for_document="1" approvalRequired_for_originalDoi="1" approvalRequired_for_textStreams="123" approvalRequired_for_treatments="5" checkinTime="1678178160677" checkinUser="plazi" docAuthor="WANG, YA-LIAN, LI, BING-XUE, CUI, MENG-DI, ZHANG, YUAN, WANG, LE, ZHANG, CAI-HONG, TSAUR, SHUN-CHERN, CHEN, HONG-WEI &amp; HUANG, JIA" docDate="2023" docId="0393914C2A728729C282FA876808FA26" docLanguage="en" docName="zootaxa.5250.1.1.pdf" docOrigin="Zootaxa 5250 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5250.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.5:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="5" docTitle="Stegana (Steganina) liusanjieae Li &amp; Chen, sp. nov." docType="treatment" docVersion="2" lastPageNumber="82" masterDocId="FFAAE9342A228778C215FF896958FB43" masterDocTitle="Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="81" updateTime="1678205790159" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
<mods:title>Revision of the subgenus Stegana (Steganina) from China, with assessment of species delimitation using DNA barcodes (Diptera, Drosophilidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>WANG, YA-LIAN</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>LI, BING-XUE</mods:namePart>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CUI, MENG-DI</mods:namePart>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ZHANG, YUAN</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>WANG, LE</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>ZHANG, CAI-HONG</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>TSAUR, SHUN-CHERN</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>CHEN, HONG-WEI</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>HUANG, JIA</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-07</mods:number>
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<mods:detail type="volume">
<mods:number>5250</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>1</mods:start>
<mods:end>109</mods:end>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5250.1.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.5250.1.1</mods:identifier>
<mods:identifier type="ISSN">1175­5334</mods:identifier>
<mods:identifier type="Zenodo-Dep">7705657</mods:identifier>
<mods:identifier type="ZooBank">4729EAF3-8B78-4AAD-B373-9179D008F239</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:0393914C2A728729C282FA876808FA26" httpUri="http://treatment.plazi.org/id/0393914C2A728729C282FA876808FA26" lastPageId="81" lastPageNumber="82" pageId="80" pageNumber="81">
<subSubSection box="[151,787,1293,1320]" pageId="80" pageNumber="81" type="nomenclature">
<paragraph blockId="80.[151,787,1293,1356]" box="[151,787,1293,1320]" pageId="80" pageNumber="81">
<heading bold="true" box="[151,787,1293,1320]" fontSize="11" level="1" pageId="80" pageNumber="81" reason="1">
<emphasis bold="true" box="[151,787,1293,1320]" pageId="80" pageNumber="81">
<taxonomicName authority="Li &amp; Chen" authorityName="Li &amp; Chen" box="[151,680,1293,1320]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="liusanjieae" status="sp. nov." subGenus="Steganina">
<emphasis bold="true" box="[151,249,1294,1320]" italics="true" pageId="80" pageNumber="81">Stegana</emphasis>
(
<emphasis bold="true" box="[266,388,1294,1320]" italics="true" pageId="80" pageNumber="81">Steganina</emphasis>
)
<emphasis bold="true" box="[405,537,1293,1320]" italics="true" pageId="80" pageNumber="81">liusanjieae</emphasis>
Li &amp; Chen
</taxonomicName>
,
<taxonomicNameLabel box="[693,787,1294,1320]" pageId="80" pageNumber="81" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,364,1330,1356]" pageId="80" pageNumber="81" type="description">
<paragraph blockId="80.[151,787,1293,1356]" box="[151,364,1330,1356]" pageId="80" pageNumber="81">
(
<figureCitation box="[159,315,1330,1356]" captionStart="FIGURE 10" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-19@33.[173,1415,181,1919]" captionTargetPageId="33" captionText="FIGURE 10. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) kanda sp. nov.; C, D. S. (S.) labao Cui &amp; Chen, sp. nov.; E, F. S. (S.) lancang Li &amp; Chen, sp. nov.; G, H. S. (S.) latifoliacea Wang &amp; Chen, sp. nov.; I, J. S. (S.) liusanjieae Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705687" httpUri="https://zenodo.org/record/7705687/files/figure.png" pageId="80" pageNumber="81">Figs. 10I, 10J</figureCitation>
,
<figureCitation box="[327,356,1330,1356]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">48</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="80" pageNumber="81" type="diagnosis">
<paragraph blockId="80.[151,1437,1402,1933]" pageId="80" pageNumber="81">
<emphasis bold="true" box="[151,274,1402,1428]" pageId="80" pageNumber="81">Diagnosis.</emphasis>
This species is similar to
<emphasis box="[565,579,1402,1428]" italics="true" pageId="80" pageNumber="81">S</emphasis>
.
<taxonomicName authority="Wang, Gao, &amp; Chen, 2013" authorityName="Wang, Gao, &amp; Chen" authorityYear="2013" box="[593,1042,1402,1428]" class="Insecta" family="Drosophilidae" genus="Stegana" kingdom="Animalia" order="Diptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="hirticeps" subGenus="Steganina">
(
<emphasis box="[602,616,1402,1428]" italics="true" pageId="80" pageNumber="81">S</emphasis>
.)
<emphasis box="[640,738,1402,1428]" italics="true" pageId="80" pageNumber="81">hirticeps</emphasis>
Wang, Gao, &amp; Chen,
<quantity box="[986,1042,1402,1428]" metricMagnitude="1" metricUnit="m" metricValue="5.11302" pageId="80" pageNumber="81" unit="in" value="2013.0">2013</quantity>
</taxonomicName>
in the shape of male terminalia (as fig.
<quantity box="[198,243,1438,1465]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="80" pageNumber="81" unit="in" value="5.0">5 in</quantity>
<bibRefCitation author="Wang, L. &amp; Gao, J. J. &amp; Chen, H. W." box="[251,451,1438,1464]" pageId="80" pageNumber="81" pagination="1993 - 2013" refId="ref74574" refString="Wang, L., Gao, J. J. &amp; Chen, H. W. (2013) Ten new species of the subgenus Steganina (Diptera: Drosophilidae: Stegana) from southern China. Journal of Natural History, 47, 1993 - 2013. https: // doi. org / 10.1080 / 00222933.2012.763055" type="journal article" year="2013">
Wang
<emphasis box="[325,383,1438,1464]" italics="true" pageId="80" pageNumber="81">et al.</emphasis>
2013
</bibRefCitation>
,
<figureCitation box="[464,599,1438,1464]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48A–F</figureCitation>
), but can be distinguished from the latter by having the oval cercus (
<figureCitation captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48A</figureCitation>
); the aedeagus with numerous minute, serrated processes laterally (
<figureCitation box="[957,1092,1474,1500]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48B, C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="80" pageNumber="81" type="description">
<paragraph blockId="80.[151,1437,1402,1933]" pageId="80" pageNumber="81">
<emphasis bold="true" box="[199,339,1510,1536]" pageId="80" pageNumber="81">Description.</emphasis>
Male and female: First flagellomere yellow. Clypeus yellow. Mesonotum slightly brownish medially, yellow laterally (
<figureCitation box="[332,421,1546,1572]" captionStart="FIGURE 10" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-19@33.[173,1415,181,1919]" captionTargetPageId="33" captionText="FIGURE 10. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) kanda sp. nov.; C, D. S. (S.) labao Cui &amp; Chen, sp. nov.; E, F. S. (S.) lancang Li &amp; Chen, sp. nov.; G, H. S. (S.) latifoliacea Wang &amp; Chen, sp. nov.; I, J. S. (S.) liusanjieae Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705687" httpUri="https://zenodo.org/record/7705687/files/figure.png" pageId="80" pageNumber="81">Fig. 10I</figureCitation>
). Katepisternum yellowish (
<figureCitation box="[722,812,1546,1572]" captionStart="FIGURE 10" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-19@33.[173,1415,181,1919]" captionTargetPageId="33" captionText="FIGURE 10. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) kanda sp. nov.; C, D. S. (S.) labao Cui &amp; Chen, sp. nov.; E, F. S. (S.) lancang Li &amp; Chen, sp. nov.; G, H. S. (S.) latifoliacea Wang &amp; Chen, sp. nov.; I, J. S. (S.) liusanjieae Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705687" httpUri="https://zenodo.org/record/7705687/files/figure.png" pageId="80" pageNumber="81">Fig. 10J</figureCitation>
). Scutellum brownish (
<figureCitation box="[1060,1148,1546,1572]" captionStart="FIGURE 10" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-19@33.[173,1415,181,1919]" captionTargetPageId="33" captionText="FIGURE 10. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) kanda sp. nov.; C, D. S. (S.) labao Cui &amp; Chen, sp. nov.; E, F. S. (S.) lancang Li &amp; Chen, sp. nov.; G, H. S. (S.) latifoliacea Wang &amp; Chen, sp. nov.; I, J. S. (S.) liusanjieae Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705687" httpUri="https://zenodo.org/record/7705687/files/figure.png" pageId="80" pageNumber="81">Fig. 10I</figureCitation>
). Halter mostly white (
<figureCitation captionStart="FIGURE 10" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-19@33.[173,1415,181,1919]" captionTargetPageId="33" captionText="FIGURE 10. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) kanda sp. nov.; C, D. S. (S.) labao Cui &amp; Chen, sp. nov.; E, F. S. (S.) lancang Li &amp; Chen, sp. nov.; G, H. S. (S.) latifoliacea Wang &amp; Chen, sp. nov.; I, J. S. (S.) liusanjieae Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705687" httpUri="https://zenodo.org/record/7705687/files/figure.png" pageId="80" pageNumber="81">Fig. 10J</figureCitation>
). Fore femur with a row of 3 long setae on distal part of posteroventral surface. Abdominal tergites mostly yellow (
<figureCitation box="[159,269,1618,1644]" captionStart="FIGURE 10" captionStartId="33.[151,250,1943,1967]" captionTargetBox="[186,1415,181,1919]" captionTargetId="figure-19@33.[173,1415,181,1919]" captionTargetPageId="33" captionText="FIGURE 10. Head, thorax, and abdomen in dorsal and lateral views: A, B. Stegana (Steganina) kanda sp. nov.; C, D. S. (S.) labao Cui &amp; Chen, sp. nov.; E, F. S. (S.) lancang Li &amp; Chen, sp. nov.; G, H. S. (S.) latifoliacea Wang &amp; Chen, sp. nov.; I, J. S. (S.) liusanjieae Li &amp; Chen, sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7705687" httpUri="https://zenodo.org/record/7705687/files/figure.png" pageId="80" pageNumber="81">Fig. 10I, J</figureCitation>
); sternites yellow, about 1.6 times as wide as long. Male terminalia (
<figureCitation box="[990,1119,1618,1644]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48A–F</figureCitation>
): Epandrium with
<emphasis box="[1313,1338,1619,1644]" italics="true" pageId="80" pageNumber="81">ca</emphasis>
. 15 setae on dorsal to posterior portions per side (
<figureCitation box="[583,683,1654,1680]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48A</figureCitation>
). Surstylus lacking peg-like tooth and pubescence, with some minute, serrated processes distally and setulae near distal margin (
<figureCitation box="[765,864,1690,1716]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48A</figureCitation>
). Cercus lacking pubescence (
<figureCitation box="[1184,1283,1690,1716]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48A</figureCitation>
). Hypandrium roundly expanded anteromedially (
<figureCitation box="[534,667,1726,1752]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48B, C</figureCitation>
). Pregonite small, with 2 sensilla (
<figureCitation box="[1051,1185,1726,1752]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48B, C</figureCitation>
). Aedeagus with much pubescence distally (
<figureCitation box="[373,500,1762,1788]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48B, C</figureCitation>
). Aedeagal sheath distally dorsad curved and with some minute, serrated processes (
<figureCitation captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48D, E</figureCitation>
). Subepandrial sclerite vaulted (
<figureCitation box="[571,667,1798,1824]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48F</figureCitation>
). Female terminalia (
<figureCitation box="[891,1014,1798,1824]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48G–I</figureCitation>
): Epiproct with some setae on distal 1/2 (
<figureCitation box="[159,259,1834,1860]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48G</figureCitation>
). Cercus and hypoproct with numerous setae and much pubescence (
<figureCitation box="[992,1112,1834,1860]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48G, I</figureCitation>
). Eight sternite slightly longer than wide, glabrous on basal 3/4, with some setae and much pubescence on distal 1/4 (
<figureCitation box="[1081,1182,1870,1896]" captionStart="FIGURE 48" captionStartId="81.[151,250,1518,1542]" captionTargetBox="[160,1427,391,1495]" captionTargetId="figure-74@81.[151,1436,381,1495]" captionTargetPageId="81" captionText="FIGURE 48. Stegana (Steganina) liusanjie Li &amp; Chen, sp. nov., male (A–F) and female terminalia (G–I). A. Epandrium, surstylus, and cercus in lateral view; B, C. Hypandrium, pregonite, aedeagus, and phallapodeme in ventral and lateral views; D, E. Aedeagal sheath in ventral and lateral views; F. Subepandrial sclerite in ventral view; G. Epiproct and cercus in dorsal view; H. Eighth sternite in ventral view; I. Hypoproct in ventral view." figureDoi="http://doi.org/10.5281/zenodo.7705782" httpUri="https://zenodo.org/record/7705782/files/figure.png" pageId="80" pageNumber="81">Fig. 48H</figureCitation>
).
</paragraph>
<paragraph blockId="80.[151,1437,1402,1933]" box="[199,1437,1906,1933]" pageId="80" pageNumber="81">
<emphasis bold="true" box="[199,522,1906,1932]" pageId="80" pageNumber="81">Measurements and indices.</emphasis>
BL =
<quantity box="[592,691,1906,1932]" metricMagnitude="-3" metricUnit="m" metricValue="2.88" pageId="80" pageNumber="81" unit="mm" value="2.88">2.88 mm</quantity>
in
<typeStatus box="[725,826,1906,1932]" pageId="80" pageNumber="81">holotype</typeStatus>
, ThL =
<quantity box="[912,1014,1906,1933]" metricMagnitude="-3" metricUnit="m" metricValue="1.59" pageId="80" pageNumber="81" unit="mm" value="1.59">1.59 mm</quantity>
, WL =
<quantity box="[1095,1197,1906,1932]" metricMagnitude="-3" metricUnit="m" metricValue="3.01" pageId="80" pageNumber="81" unit="mm" value="3.01">3.01 mm</quantity>
, WW =
<quantity box="[1288,1390,1906,1932]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="80" pageNumber="81" unit="mm" value="1.17">1.17 mm</quantity>
, arb
</paragraph>
<paragraph blockId="80.[151,1437,1942,2041]" pageId="80" pageNumber="81">= 6/4, avd = 0.95, adf = 1.05, flw = 2.85, FW/HW = 0.35, ch/o = 0.27, prorb = 1.13, rcorb = 0.66, vb = 0.65, dcl = 0.56, presctl = 0.51, sctl = 1.78, sterno = 1.20, orbito = 1.65, dcp = 0.30, sctlp = 1.79, C = 1.74, 4c = 1.32, 4v = 1.97, 5x = 1.42, ac = 10.08, M = 0.55, C3F = 0.64.</paragraph>
</subSubSection>
<subSubSection pageId="81" pageNumber="82" type="materials_examined">
<paragraph blockId="81.[151,1436,150,357]" pageId="81" pageNumber="82">
<emphasis bold="true" box="[199,386,150,177]" pageId="81" pageNumber="82">Type materials.</emphasis>
<materialsCitation collectingDate="2015-09-14" collectionCode="SCAU" collectorName="Y. Q. Liu" country="China" elevation="1500" latitude="25.313334" location="Huanjiang" longLatPrecision="21" longitude="108.63389" municipality="Jiuwanshan" pageId="81" pageNumber="82" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<typeStatus box="[394,498,151,177]" pageId="81" pageNumber="82">Holotype</typeStatus>
♁ (
<collectionCode box="[545,624,151,177]" pageId="81" pageNumber="82">SCAU</collectionCode>
, No. 125669, DNA #1951),
<collectingCountry box="[949,1042,151,177]" name="China" pageId="81" pageNumber="82">CHINA</collectingCountry>
:
<collectingMunicipality box="[1054,1188,151,177]" pageId="81" pageNumber="82">Jiuwanshan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:0393914C2A728729C282FA876808FA26:8EE576812A738729C6A5FF1E6C70FBF2" box="[1200,1320,151,177]" country="China" latitude="25.313334" longLatPrecision="21" longitude="108.63389" municipality="Jiuwanshan" name="Huanjiang" pageId="81" pageNumber="82" stateProvince="Guangxi">Huanjiang</location>
,
<collectingRegion box="[1333,1432,151,177]" country="China" name="Guangxi" pageId="81" pageNumber="82">Guangxi</collectingRegion>
,
<geoCoordinate box="[151,284,186,213]" degrees="25" direction="north" minutes="18" orientation="latitude" pageId="81" pageNumber="82" precision="15" seconds="48" value="25.313334">25°18'48''N</geoCoordinate>
,
<geoCoordinate box="[299,443,187,213]" degrees="108" direction="east" minutes="38" orientation="longitude" pageId="81" pageNumber="82" precision="15" seconds="02" value="108.63389">108°38'02''E</geoCoordinate>
, alt.
<quantity box="[502,582,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="81" pageNumber="82" unit="m" value="1500.0">
<elevation box="[502,582,186,213]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="81" pageNumber="82" unit="m" value="1500.0">1500m</elevation>
</quantity>
,
<date box="[596,719,186,213]" pageId="81" pageNumber="82" value="2015-09-14">
<collectingDate box="[596,719,186,213]" pageId="81" pageNumber="82" value="2015-09-14">14.ix.2015</collectingDate>
</date>
, ex tree trunk,
<collectorName box="[896,999,187,213]" pageId="81" pageNumber="82">Y.Q. Liu</collectorName>
</materialsCitation>
.
<materialsCitation collectingDate="2015-09-14" collectionCode="SCAU" collectorName="Y. Q. Liu" country="China" elevation="1500" latitude="25.313334" location="China" longLatPrecision="21" longitude="108.63389" municipality="Zhuang Nationality" pageId="81" pageNumber="82" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="paratype">
<typeStatus box="[1013,1126,187,213]" pageId="81" pageNumber="82">Paratypes</typeStatus>
:
<specimenCount box="[1139,1174,187,213]" pageId="81" pageNumber="82" type="female">1♀</specimenCount>
(
<collectionCode box="[1192,1271,187,213]" pageId="81" pageNumber="82">SCAU</collectionCode>
, No. 125670), other data same as holotype.
<emphasis bold="true" box="[199,353,258,284]" pageId="81" pageNumber="82">Distribution.</emphasis>
China (
<collectingRegion box="[441,544,259,285]" country="China" name="Guangxi" pageId="81" pageNumber="82">Guangxi</collectingRegion>
).
<emphasis bold="true" box="[199,334,295,321]" pageId="81" pageNumber="82">Etymology.</emphasis>
Patronym, in honour of &quot;Liu Sanjie&quot;, a famous girl singer from the
<collectingMunicipality box="[1112,1332,295,321]" pageId="81" pageNumber="82">Zhuang Nationality</collectingMunicipality>
living in
<collectingRegion box="[151,250,331,357]" country="China" name="Guangxi" pageId="81" pageNumber="82">Guangxi</collectingRegion>
,
<location LSID="urn:lsid:plazi:treatment:0393914C2A728729C282FA876808FA26:8EE576812A738729C313FEC26815FA26" box="[262,333,331,357]" municipality="Zhuang Nationality" name="China" pageId="81" pageNumber="82" stateProvince="Guangxi">China</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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