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286 lines
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<mods:title id="D441BC08249ADE39EF98411E963B409D">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<mods:namePart id="B3B39F7D1A31AA99C2BB6F01EA89504B">Chamorro, William</mods:namePart>
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<mods:namePart id="DFC5C462FA62FEA006F9BC5E412083C6">Marin-Armijos, Diego</mods:namePart>
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<mods:namePart id="4B60E7B82F100855C5678EAFA5A1BFCC">senjo, Angelico</mods:namePart>
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<mods:namePart id="B39C43C98231AD019C28704728F11C44">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:title id="0FF64170164C2483BFB74C419FDA9B2F">ZooKeys</mods:title>
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<mods:date id="0F4452C5D30369A03BA999424FFE9ACD">2019</mods:date>
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<mods:number id="03839666C80149B00D40E7665AAEE4A9">826</mods:number>
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<mods:classification id="CB5CEA4A902C0489648F38E51CB81462">journal article</mods:classification>
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<treatment id="58071E6D2CBB4638F57CB9C8B5D2211E" ID-GBIF-Taxon="154127625" LSID="urn:lsid:plazi:treatment:58071E6D2CBB4638F57CB9C8B5D2211E" httpUri="http://treatment.plazi.org/id/58071E6D2CBB4638F57CB9C8B5D2211E" lastPageId="216" lastPageNumber="217" pageId="213" pageNumber="214">
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<subSubSection id="3863EC045AB779B6765B3598FFF26189" pageId="213" pageNumber="214" type="nomenclature">
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<paragraph id="A15540C606A3F0BDE0818BDAA5093B78" pageId="213" pageNumber="214">
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<taxonomicName id="B6C20948CB30151C86B4DCD7B5CA171A" ID-CoL="4FSJX" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus (Notiophanaeus) pyrois" order="Coleoptera" pageId="213" pageNumber="214" phylum="Arthropoda" rank="species" species="pyrois" subGenus="Notiophanaeus">
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<pageBreakToken id="359EDEC47D1ABC261C29D217591665D8" pageId="213" pageNumber="214" start="start">Phanaeus</pageBreakToken>
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(Notiophanaeus) pyrois Bates, 1887
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</taxonomicName>
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Plate 47C
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</paragraph>
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</subSubSection>
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<subSubSection id="BDF4D5A77D657688D471B13C279D5641" pageId="214" pageNumber="215" type="reference_group">
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<paragraph id="67DEA76C54B08A7D720478E4EF73AC2F" pageId="214" pageNumber="215">
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<taxonomicName id="51CB807C2F544DF99AB7F287A615C4DA" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois">
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<pageBreakToken id="FE75AAB5A3C97F298580057F631BAEA7" pageId="214" pageNumber="215" start="start">Phanaeus</pageBreakToken>
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pyrois
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</taxonomicName>
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Bates, 1887: 58 (original description. Type locality: Nicaragua, Chontales; Costa Rica: Panama, Bugaba, Chiriqui Volcano, 2000-3000 feet [= 610-915 m]; South America, Colombia).
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</paragraph>
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<paragraph id="29AAA10A38E49F11F6A0949A6AB0C4A3" pageId="214" pageNumber="215">
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<taxonomicName id="7EBCB52E8803B6D36DBD530AC4E9ACB4" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois">Phanaeus pyrois</taxonomicName>
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:
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<bibRefCitation id="32190A134B15C00815C2C9AF72397906" author="Nevinson, BG" journalOrPublisher="Published privately, London" pageId="274" pageNumber="275" title="Revised synonymic list of species in the genera Phanaeus Macleay and Oxysternon Castelnau." url="https://doi.org/10.5962/bhl.title.9416" year="1892">Nevinson 1892</bibRefCitation>
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: 6 (catalog, distribution);
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<bibRefCitation id="B601407058977FC5C3A15F773598CE6D" author="Gillet, JJE" journalOrPublisher="IW Junk, Berlin, Germany" pageId="268" pageNumber="269" title="Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae." year="1911 a">Gillet 1911a</bibRefCitation>
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: 85 (complete list of species);
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<normalizedToken id="B78EE2B65D55D3D5C70BB9AF6E806626" originalValue="d’">d'</normalizedToken>
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<bibRefCitation id="AD94125FEC287508B90575F6A4C27FEC" author="Olsoufieff, Gd'" journalOrPublisher="Revue Illustree d'Entomologie" pageId="274" pageNumber="275" pagination="4 - 172" title="Les Phanaeides, Famille Scarabaeidae. Tr. CopriniInsecta." volume="13" year="1924">Olsoufieff 1924</bibRefCitation>
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: 37 (characters in key); 93 (comment, distribution), 152 (distribution);
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<bibRefCitation id="52F09D513794A8206DD34A5789121B95" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
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: 210 (list of species from Latin America);
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<bibRefCitation id="B74DBA49484FDA3552D389CD33FBFC15" author="Medina, CA" journalOrPublisher="Biota Colombiana" pageId="274" pageNumber="275" pagination="131 - 144" title="Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia." volume="2" year="2001">Medina et al. 2001</bibRefCitation>
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: 140 (cited for Colombia);
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<bibRefCitation id="8FC4BF8927DCB457C2823F5ABA1A0B7E" pageId="214" pageNumber="215">Krajcik 2012</bibRefCitation>
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: 204 (complete list of species);
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<bibRefCitation id="016943AD1CC41B4407B4FD884D530D41" author="Moctezuma, V" journalOrPublisher="The Coleopterists Bulletin" pageId="274" pageNumber="275" pagination="47 - 56" title="A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico." url="https://doi.org/10.1649/0010-065X-71.1.47" volume="71" year="2017">Moctezuma and Halffter 2017</bibRefCitation>
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: 55 (characters in key, cited for Ecuador);
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<bibRefCitation id="E9BC274459213CDE97122915A30CA468" author="Kohlmann, B" journalOrPublisher="ZooKeys" pageId="271" pageNumber="272" pagination="67 - 93" title="Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae)." url="https://doi.org/10.3897/zookeys.743.23029" volume="743" year="2018">Kohlmann et al. 2018</bibRefCitation>
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: 79 (figures 8b and d), 83 (characters in key, cited for Ecuador).
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</paragraph>
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<paragraph id="606B69D1A3C12C56D861899793708AF4" pageId="214" pageNumber="215">
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<taxonomicName id="6F698F4DC8D522131CA6850F5ED6AF7E" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus (Notiophanaeus) pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois" subGenus="Notiophanaeus">Phanaeus (Notiophanaeus) pyrois</taxonomicName>
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:
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<bibRefCitation id="75559E9212567D7CE83CDA1C616F9CA7" author="Edmonds, WD" journalOrPublisher="Contributions in Science" pageId="266" pageNumber="267" pagination="1 - 105" title="Revision of Phanaeus Macleay, a new world genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)." volume="443" year="1994">Edmonds 1994</bibRefCitation>
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: 41 (characters in key), 45 (redescription), 103 (distribution);
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<bibRefCitation id="5009FCB0E57DBA2C85A8481E1597823D" author="Vitolo, AL" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Exactas Fisicas y Naturales" pageId="278" pageNumber="279" pagination="591 - 601" title="Clave para la identificacion de los generos y especies Phanaeinas de Colombia." volume="24" year="2000">Vitolo 2000</bibRefCitation>
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: 597 (characters in key);
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<bibRefCitation id="6CB0EB01FBC3143821E8CCB6684800F9" author="Vitolo, A" editor="Fernandez, F" journalOrPublisher="Facultad de Ciencias, Universidad Nacional de Colombia, Bogota" pageId="278" pageNumber="279" pagination="277 - 318" title="Escarabajos estercoleros de la tribu Phanaeini de Colombia (Coleoptera: Scarabaeoidea: Scarabaeidae)." volume="3" volumeTitle="Insectos de Colombia." year="2004">Vitolo 2004</bibRefCitation>
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: 284 (diagnosis);
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<bibRefCitation id="752E262CC15B35F859E4193D71B93777" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
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: 322-323 (cited for Ecuador);
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<bibRefCitation id="822E4375D64D4B4ACA9D3C72D0051D0D" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 108" title="Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini)." volume="0274" year="2012">
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Edmonds and
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<normalizedToken id="EB41F1D9AE22F69DE701EB1F91EB218F" originalValue="Zídek">Zidek</normalizedToken>
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2012
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</bibRefCitation>
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: 13 (characters in key), 53 (distribution figure 143);
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<bibRefCitation id="75330B4CCC5AF3C373BDE1A32C41C835" author="Bezdek, A" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="262" pageNumber="263" pagination="387 - 442" title="Catalogue of type specimens of beetles deposited in the National Museum, Prague, Czech Republic. Scarabaeidae: Scarabaeinae: Onitini, Onthophagini, Phanaeini, Scarabaeini and Sisyphini." volume="53" year="2013">Bezdek and Hajek 2013</bibRefCitation>
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: 432 (cited as junior subjective synonym of
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<taxonomicName id="2AF5D6506E7E6A972900ECCC679D6D34" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus (Notiophanaeus) pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois" subGenus="Notiophanaeus">Phanaeus (Notiophanaeus) pyrois</taxonomicName>
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Bates, 1887. Catalog of the types of the NMPC);
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<bibRefCitation id="0C3683FE7DB3AE968570E2DAAEE627CE" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
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: 84 (figure 7D), 98 (cited for Ecuador);
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<bibRefCitation id="1FE204CF2039FE787734D1531FE21452" author="Arnaud, P" journalOrPublisher="Besoiro" pageId="261" pageNumber="262" pagination="3 - 7" title="Description d'une nouvelle espe ̀ ce de Phanaeus d'E ́ quateur et revalidation de l'espe ̀ ce Coprophanaeus (C.) edmondsi Arnaud (Coleoptera: Scarabaeidae, Scarabaeinae, Phanaeini)." volume="26" year="2018">Arnaud 2018</bibRefCitation>
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: 4 (comment), 5 (figures 2e-d).
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</paragraph>
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<paragraph id="C8026A4A88309991F870F657B79FAECB" pageId="214" pageNumber="215">
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<taxonomicName id="47AC12CAB130B2F73872AFE3E0CE40C9" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus blanchardi" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="blanchardi">Phanaeus blanchardi</taxonomicName>
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<normalizedToken id="028E1FACF286367CE0C4C9711C977383" originalValue="d’Olsoufieff">d'Olsoufieff</normalizedToken>
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, 1924: 37 (characters in key), 92 (original description, cited for Ecuador);
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<bibRefCitation id="20CF9C02329EE7D15FA09722789736ED" author="Vulcano, MA" journalOrPublisher="Atas do Simposio sobre a Biota Amazonica" pageId="279" pageNumber="280" pagination="533 - 603" title="Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera)." volume="5" year="1967">Vulcano and Pereira 1967</bibRefCitation>
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: 575 (characters in key);
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<bibRefCitation id="7F005948808847EBAAFA54C1DCB8B211" author="Martinez, A" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="273" pageNumber="274" pagination="53 - 69" title="Notas Escarabeidologicas III." volume="29" year="1967">
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<normalizedToken id="917C175DBA932044AF376DFE72502F3C" originalValue="Martínez">Martinez</normalizedToken>
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and Pereira 1967
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</bibRefCitation>
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: 68 (synonym of
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<taxonomicName id="694ED50373620EF6AEC45782FB7E3952" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus (Phanaeus) funereus" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="funereus" subGenus="Phanaeus">Phanaeus (Phanaeus) funereus</taxonomicName>
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Balthasar, 1939. Cited for Ecuador); Edmonds, 1994: 45 (synonym of
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<taxonomicName id="3A4B4CF9B52D39D43B71DDA789C700B3" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois">Phanaeus pyrois</taxonomicName>
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Bates, 1887);
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<bibRefCitation id="168BA8B5B4DAE0777896B6F482C67154" author="Vitolo, A" editor="Fernandez, F" journalOrPublisher="Facultad de Ciencias, Universidad Nacional de Colombia, Bogota" pageId="278" pageNumber="279" pagination="277 - 318" title="Escarabajos estercoleros de la tribu Phanaeini de Colombia (Coleoptera: Scarabaeoidea: Scarabaeidae)." volume="3" volumeTitle="Insectos de Colombia." year="2004">Vitolo 2004</bibRefCitation>
|
|||
|
|
: 284 (cited as synonym for
|
|||
|
|
<taxonomicName id="5F105B0CB895C1CB738E435F307D19E1" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois">Phanaeus pyrois</taxonomicName>
|
|||
|
|
Bates, 1887);
|
|||
|
|
<bibRefCitation id="FAF0247230548851B72A2C918480F4D7" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
|
|||
|
|
: 320-321 (cited for Ecuador); Edmonds and Zídek, 2012: 5 (cited as jr. prim. hom. of
|
|||
|
|
<taxonomicName id="1D217682AE459E73D135744B66A5D298" family="Scarabaeidae" genus="Gromphina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Gromphina blanchardi" order="Coleoptera" pageId="214" pageNumber="215" rank="species" species="blanchardi">blanchardi</taxonomicName>
|
|||
|
|
Harold. Permanently unavailable, valid name
|
|||
|
|
<taxonomicName id="E765F226A503BAB037CF7D7CB4D4E0BE" family="Scarabaeidae" genus="Gromphina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Gromphina pyrois" order="Coleoptera" pageId="214" pageNumber="215" rank="species" species="pyrois">pyrois</taxonomicName>
|
|||
|
|
Bates), 13 (comment).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6AA6D548586204FCFA0E93D9762599CF" pageId="214" pageNumber="215">
|
|||
|
|
<taxonomicName id="60E1A31DF8BDD14B9294D4AED02824EC" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus funereus" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="funereus">Phanaeus (s. str.) funereus</taxonomicName>
|
|||
|
|
Balthasar, 1939: 241 (original description, cited for Ecuador).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="6B58BAC407F39CCBEFF24750974146DF" pageId="214" pageNumber="215">
|
|||
|
|
<taxonomicName id="CD255AD3CE1C2A7353319C0C46DED4A8" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus funereus" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="funereus">Phanaeus funereus</taxonomicName>
|
|||
|
|
:
|
|||
|
|
<bibRefCitation id="43F6AB99069F3DAAE7A17F848A8107E4" author="Blackwelder, RE" journalOrPublisher="National Museum Bulletin" pageId="262" pageNumber="263" pagination="189 - 341" title="Checklist of the coleopterous insects of Mexico, Central America, West Indies and South America. Part 2." volume="185" year="1944">Blackwelder 1944</bibRefCitation>
|
|||
|
|
: 209 (list of species from Latin America);
|
|||
|
|
<bibRefCitation id="2EAD2F114A3ED946F2B2300669C431FA" author="Martinez, A" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="273" pageNumber="274" pagination="53 - 69" title="Notas Escarabeidologicas III." volume="29" year="1967">
|
|||
|
|
<normalizedToken id="D6AEB262EF47656C2054BFD917980B11" originalValue="Martínez">Martinez</normalizedToken>
|
|||
|
|
and Pereira 1967
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 68 (distribution, comment);
|
|||
|
|
<bibRefCitation id="49E575D35893C88E762FF685CFF68748" author="Edmonds, WD" journalOrPublisher="Contributions in Science" pageId="266" pageNumber="267" pagination="1 - 105" title="Revision of Phanaeus Macleay, a new world genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)." volume="443" year="1994">Edmonds 1994</bibRefCitation>
|
|||
|
|
: 45 (synonym of
|
|||
|
|
<taxonomicName id="CB262A71D4BF25D4A98217C5EACC9929" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois">Phanaeus pyrois</taxonomicName>
|
|||
|
|
Bates, 1887); 103 (distribution);
|
|||
|
|
<bibRefCitation id="8347B7CB19E1108A3DD587B703763905" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 108" title="Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini)." volume="0274" year="2012">
|
|||
|
|
Edmonds and
|
|||
|
|
<normalizedToken id="640D3D45E1A1B3FC39F67D5158C30582" originalValue="Zídek">Zidek</normalizedToken>
|
|||
|
|
2012
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 5 (cited as junior synonym for
|
|||
|
|
<taxonomicName id="525BDE2130D49B4EE8132FC7A19844D0" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois">Phanaeus pyrois</taxonomicName>
|
|||
|
|
Bates, 1887).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="AF058F7C3687D51271AF153E985380EA" pageId="214" pageNumber="215">
|
|||
|
|
<taxonomicName id="A2FBD607FFA2C43444DC2C3A63E7584A" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus (Notiophanaeus) pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="subSpecies" species="pyrois" subGenus="Notiophanaeus" subSpecies="funereus">Phanaeus (Notiophanaeus) pyrois funereus</taxonomicName>
|
|||
|
|
:
|
|||
|
|
<bibRefCitation id="8D02C56D88564CAD9FFF3076D40D23BF" author="Arnaud, P" journalOrPublisher="Hillside Books, Canterbury, England" pageId="261" pageNumber="262" title="Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus." year="2002 a">Arnaud 2002a</bibRefCitation>
|
|||
|
|
: 97 (diagnosis);
|
|||
|
|
<bibRefCitation id="9C573A4A6A622558B0165B5141259E6A" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
|
|||
|
|
: 322-323 (cited for Ecuador).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="378A3FF90B1C8F31CAAA9E05AF266477" lastPageId="215" lastPageNumber="216" pageId="214" pageNumber="215" type="type specimens">
|
|||
|
|
<paragraph id="0097F5D2362A1288AFD7B8BF85F53388" pageId="214" pageNumber="215">Type specimens.</paragraph>
|
|||
|
|
<paragraph id="EB2C89200CA17A4D5C07AB66B52221F1" pageId="214" pageNumber="215">
|
|||
|
|
<taxonomicName id="E5154B691B0701BEA861EBFBF8999420" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus pyrois" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="pyrois">Phanaeus pyrois</taxonomicName>
|
|||
|
|
Bates, 1887. The lectotype (♂) is deposited at the NHML (ex coll. H. Bates). Locality: Chontales, Nicaragua, examined.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="65FFBFBEC51C4648CD9FD61A9DA4B48F" pageId="214" pageNumber="215">Lectotype (♂): "Chontales / Nicaragua / T. Belt. [p]", "P. pyrois ♂ [hw]", "B.C.A. / p. 58, sp 8. [p]", "Sp. figured. [p]", "Phanaeus / pyrois. Bates / Lectotype ♂ / P. ARNAUD DET 1980 [hw and p, red margin]".</paragraph>
|
|||
|
|
<paragraph id="BDC8EB86E422B578F6E97DFF5CA0E91A" pageId="214" pageNumber="215">
|
|||
|
|
<taxonomicName id="9086F76DF848517C1CB90A238001B8DE" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus blanchardi" order="Coleoptera" pageId="214" pageNumber="215" phylum="Arthropoda" rank="species" species="blanchardi">Phanaeus blanchardi</taxonomicName>
|
|||
|
|
<normalizedToken id="F05E208B25A2FBC8DEFD3E2ECC8405D4" originalValue="d’Olsoufieff">d'Olsoufieff</normalizedToken>
|
|||
|
|
, 1924. The lectotype (♂) is deposited at the MNHN (see
|
|||
|
|
<bibRefCitation id="FCA8FF754949402CDC0385ADC0EB1C81" author="Arnaud, P" journalOrPublisher="Revue Francaise d'Entomologie" pageId="260" pageNumber="261" pagination="113 - 118" title="Liste des types de Phanaeini du Museum national d'Historie naturelle de Paris." volume="4" year="1982 a">Arnaud 1982a</bibRefCitation>
|
|||
|
|
: 116). Locality: Colombie
|
|||
|
|
<normalizedToken id="CCAFCDBCF71BF479DCF025B14466723B" originalValue="Vallée">Vallee</normalizedToken>
|
|||
|
|
de Cauca [= Colombia, Valle del Cauca], not examined.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="BB5ADD7B5788AA38AFD1D455BF090A04" pageId="215" pageNumber="216">
|
|||
|
|
<taxonomicName id="8A46754A6BC4768D676759CD95381FE1" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus funereus" order="Coleoptera" pageId="215" pageNumber="216" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="funereus">
|
|||
|
|
<pageBreakToken id="32B391C8EBA69836F1AEDA423C298BE0" pageId="215" pageNumber="216" start="start">Phanaeus</pageBreakToken>
|
|||
|
|
(s. str.) funereus
|
|||
|
|
</taxonomicName>
|
|||
|
|
Balthasar, 1939. The holotype (♂) is deposited at the NMPC (ex coll. V. Balthasar). Locality: Pucay [= Bucay], examined.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="A6E6D29B78F2BEB75F0A5EAC801F041E" pageId="215" pageNumber="216">Holotype (♂): "ECUADOR / Pucay / F.O., 10.6.05 [p]", "♂ [hw]", "H. Blut Determ. / Phanaeus / blanchardi Olsuf. [hw and p]", "Felsche det. / Phanaeus / pyrois Bts. [hw and p]", "Ph. funereus / m. n.sp. / Dr. v. Balthasar det. [hw and p]", "Typus [p, red label, black margin]", "Mus. Nat. Pragae / Inv. [p], 26347 [hw, orange label]", "ex coll. V. Balthasar / National Museum Prague / Czech Republic [p]".</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="6EE76819413853E816D1B3240747045F" pageId="215" pageNumber="216" type="distribution">
|
|||
|
|
<paragraph id="995639B78A12420588BC696D3A9E3C14" pageId="215" pageNumber="216">Distribution.</paragraph>
|
|||
|
|
<paragraph id="01B1830E9E8FB12E6BC6B6D00291C01B" pageId="215" pageNumber="216">Colombia and Ecuador.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="118A13A4D94ECF7E8364035291A3F6EA" pageId="215" pageNumber="216" type="records examined">
|
|||
|
|
<paragraph id="4182D8448C1392CD9A80E5251CAE9405" pageId="215" pageNumber="216">Records examined.</paragraph>
|
|||
|
|
<paragraph id="058FB1CE33EEAB24805024155E35A6B1" pageId="215" pageNumber="216">
|
|||
|
|
BOLIVAR: Altamira de Echeandia, 520-720 m (1 specimen MUTPL). CARCHI: Tobar Donoso, 300 m (5 specimens MECN). ESMERALDAS:
|
|||
|
|
<normalizedToken id="C0A084E521362AEE8DCBD1DD57018450" originalValue="Colón">Colon</normalizedToken>
|
|||
|
|
del Onzole (1 specimen MQCAZ; 8 specimens MECN); Chisprero (5 specimens MECN); E C
|
|||
|
|
<normalizedToken id="F8B45093C9FCA7309ECFC7532972974A" originalValue="Río">Rio</normalizedToken>
|
|||
|
|
<normalizedToken id="C99AED2E023F7F8EE6F459E32AE59CEE" originalValue="Canandé">Canande</normalizedToken>
|
|||
|
|
, 390 m (2 specimens MQCAZ); Jeyambi PMFC (1 specimen MUTPL; 4 specimens MECN); Kumanii Lodge, 40 m (1 specimen MQCAZ); Majua (1 specimen CEMT; 7 specimens MECN); Gualpi El Pajonal (7 specimens MECN); Palma Real (1 specimen CEMT; 5 specimens MECN); Playa de Oro Estero Pote, 200 m (6 specimens CEMT; 11 specimens MECN); Playa de Oro La Tabla (7 specimens MECN; 2 specimens MQCAZ); Playa de Oro Playa Rica (4 specimens MECN); Playa de Oro Padre Santo (1 specimen MQCAZ; 9 specimens MECN); Salto del Bravo (5 specimens MECN); Tjespi (7 specimens MECN); Tjespi
|
|||
|
|
<normalizedToken id="E063925984C92CE75AEB25CFDBC32C98" originalValue="río">rio</normalizedToken>
|
|||
|
|
Zapallo (1 specimen MQCAZ). GUAYAS: Guayaquil, 50 m (1 specimen MQCAZ); Pucay [= Bucay] (1 specimen NMPC). IMBABURA: El Chontal, El Cauchero, 900 m (1 specimen MUTPL); Lita, 680 (3 specimens MECN). LOS
|
|||
|
|
<normalizedToken id="C2536B32DA7EC463C3F431439F61F36D" originalValue="RÍOS">RIOS</normalizedToken>
|
|||
|
|
: Quevedo Pichilingue (1 specimen CEMT).
|
|||
|
|
<normalizedToken id="86599F717AFC6D827C6C9896C63CA522" originalValue="MANABÍ">MANABI</normalizedToken>
|
|||
|
|
: Puerto
|
|||
|
|
<normalizedToken id="2BE42C038BC8AAD4A02C047AF55DE1ED" originalValue="López">Lopez</normalizedToken>
|
|||
|
|
, Guale, 310 m (1 specimen MUTPL); Puerto
|
|||
|
|
<normalizedToken id="8C0A50130B118A6D4DB24F2057BFC6B0" originalValue="López">Lopez</normalizedToken>
|
|||
|
|
, Las Tunas, 200 m (1 specimen MUTPL); Puerto
|
|||
|
|
<normalizedToken id="2204938C730D1304D52EFD5346D61C5D" originalValue="López">Lopez</normalizedToken>
|
|||
|
|
, Puerto Rico, 110 m (1 specimen MUTPL). PICHINCHA: Guayabilla, 515 m,
|
|||
|
|
<normalizedToken id="27F8AC0822E082D0F257708557235B80" originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Guayllabamba, Manduriacus (1 specimen CEMT); El Encuentro, 620 m, San Miguel de los Bancos (3 specimens MUTPL); El Tigre
|
|||
|
|
<normalizedToken id="C525C53C1A1B35156E17F6CEF5DAF90C" originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Guayllabamba, Pedro Vicente Maldonado (2 specimens MUTPL); Llurimaguas, 290 m,
|
|||
|
|
<normalizedToken id="4A4EF1D0D3D5435EC50E1DE704A7B151" originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Guayllabamba, Pedro Vicente Maldonado (1 specimen MUTPL); Mangaloma, 820 m, San Miguel de los Bancos (1 specimen MUTPL); Mindo, 1500 m (2 specimens MECN); San Roque, 580 m,
|
|||
|
|
<normalizedToken id="B98FC9AF4E4D3778AFE6EE0B207CC62E" originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Guayllabamba, Pedro Vicente Maldonado (2 specimens MUTPL); Tortugo
|
|||
|
|
<normalizedToken id="16777ACC8C2C6317AA7BF1DD80756570" originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Guayllabamba, 450 m, Pedro Vicente Maldonado (1 specimen MUTPL). SANTA ELENA: La Rinconada, 10 m (1 specimen MQCAZ);
|
|||
|
|
<normalizedToken id="3B7BD1581C4DB234D6D8303399D4AC6A" originalValue="Olón">Olon</normalizedToken>
|
|||
|
|
50 m (2 specimens CEMT). SANTO DOMINGO DE LOS
|
|||
|
|
<normalizedToken id="31BCFC6C752A0D1AD90A7CDEF611FDBF" originalValue="TSÁCHILAS">TSACHILAS</normalizedToken>
|
|||
|
|
: Otongachi, 960 m (3 specimens MQCAZ); Santo Domingo, 500 m (2 specimens MECN).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="A19EDE7C46F93AE92400A9E2D1C7F0D5" pageId="215" pageNumber="216" type="literature records">
|
|||
|
|
<paragraph id="A3847A9828A529B42C2C8496D47EFB0D" pageId="215" pageNumber="216">Literature records.</paragraph>
|
|||
|
|
<paragraph id="FF44DDA611A126E6A7335FC517869AF9" pageId="215" pageNumber="216">
|
|||
|
|
ESMERALDAS: 11 km SE San Lorenzo (
|
|||
|
|
<bibRefCitation id="0AAED907F84EB0E9B15B19CA11C5E0BA" author="Edmonds, WD" journalOrPublisher="Contributions in Science" pageId="266" pageNumber="267" pagination="1 - 105" title="Revision of Phanaeus Macleay, a new world genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)." volume="443" year="1994">Edmonds 1994</bibRefCitation>
|
|||
|
|
: 103); San Mateo (
|
|||
|
|
<bibRefCitation id="D5481D07C6CFB62A2DC959F45D8DA0A0" author="Martinez, A" journalOrPublisher="Revista de la Sociedad Entomologica Argentina" pageId="273" pageNumber="274" pagination="53 - 69" title="Notas Escarabeidologicas III." volume="29" year="1967">
|
|||
|
|
<normalizedToken id="4827142B30CC9F3088C1C563E7F89E60" originalValue="Martínez">Martinez</normalizedToken>
|
|||
|
|
and Pereira 1967
|
|||
|
|
</bibRefCitation>
|
|||
|
|
: 68). LOS
|
|||
|
|
<normalizedToken id="66DCA654765A976A95CD3EA90CDECEBF" originalValue="RÍOS">RIOS</normalizedToken>
|
|||
|
|
: Quevedo, 45 m (
|
|||
|
|
<bibRefCitation id="9996496A6BA3A4C7EF504CA62F670E19" author="Edmonds, WD" journalOrPublisher="Contributions in Science" pageId="266" pageNumber="267" pagination="1 - 105" title="Revision of Phanaeus Macleay, a new world genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)." volume="443" year="1994">Edmonds 1994</bibRefCitation>
|
|||
|
|
: 103).
|
|||
|
|
<normalizedToken id="5BF8CE481E39377D7C689897C6F1D536" originalValue="MANABÍ">MANABI</normalizedToken>
|
|||
|
|
: 78 km NE de Chone, 450 m (
|
|||
|
|
<bibRefCitation id="44AF6E6391E2FDECEE10EA6A52F21CC3" author="Edmonds, WD" journalOrPublisher="Contributions in Science" pageId="266" pageNumber="267" pagination="1 - 105" title="Revision of Phanaeus Macleay, a new world genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)." volume="443" year="1994">Edmonds 1994</bibRefCitation>
|
|||
|
|
: 103). PICHINCHA [= SANTO DOMINGO DE LOS
|
|||
|
|
<normalizedToken id="1EA59D885DC046B966569CAE1D8FAE9E" originalValue="TSÁCHILAS">TSACHILAS</normalizedToken>
|
|||
|
|
]: 4 km SE de Santo Domingo, 500 m (
|
|||
|
|
<bibRefCitation id="512503060DF9C1EA66EB6F025AD39CBD" author="Edmonds, WD" journalOrPublisher="Contributions in Science" pageId="266" pageNumber="267" pagination="1 - 105" title="Revision of Phanaeus Macleay, a new world genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae)." volume="443" year="1994">Edmonds 1994</bibRefCitation>
|
|||
|
|
: 103). UNDETERMINED PROVINCE: without specific locality (
|
|||
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<bibRefCitation id="7247962A4C1249FD01D0E55EF0BB472C" author="Balthasar, V" journalOrPublisher="Folia Zoologica et Hydrobiologica" pageId="261" pageNumber="262" pagination="238 - 247" title="Neue Phanaeus-Arten." volume="9" year="1939 e">Balthasar 1939e</bibRefCitation>
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: 242).
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</paragraph>
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</subSubSection>
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<subSubSection id="CCB6BEC04CFA64CB5BD3ED8D72822FEE" pageId="215" pageNumber="216" type="temporal data">
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<paragraph id="B944109FFEA9CA90922321AB49545186" pageId="215" pageNumber="216">Temporal data.</paragraph>
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<paragraph id="4F391AC4FD5B3E0AAACCBEAFA7F3283A" pageId="215" pageNumber="216">Collected every month of the year.</paragraph>
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</subSubSection>
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<subSubSection id="32D7B4040A3AEF07F85076409D0C9163" pageId="216" pageNumber="217" type="remarks">
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<paragraph id="65471B33C49EF3BAD5148E74A6A263FA" pageId="216" pageNumber="217">
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<pageBreakToken id="12C9AFC272E0EDB953A3B485338B7748" pageId="216" pageNumber="217" start="start">Remarks</pageBreakToken>
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.
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</paragraph>
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<paragraph id="9C4147CBE7C42FADC0BF277BC0040CDF" pageId="216" pageNumber="217">Inhabits coastal lowland evergreen forests and coastal evergreen foothill forests from 45-820 m a.s.l. Species was collected with pitfall traps baited with fungus, carrion, and human feces.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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