treatments-xml/data/89/15/87/891587DDFFC59B25D2A2F9C5FA2A67D8.xml

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<mods:title id="D2DB9E974FF6024604A3366D2533712D">Revalidation of Platystigma Kennedy, 1920, with a synopsis of the quadratum species group and the description of three new species (Odonata: Pseudostigmatidae)</mods:title>
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<mods:namePart id="EEEB590BC127F8A586B5237A7644317E">Machado, Angelo B. M.</mods:namePart>
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<heading id="5A4B81A7FFC59B22D2A2F9C5FE5C6130" bold="true" box="[151,493,1544,1570]" fontSize="11" level="1" pageId="3" pageNumber="496" reason="1">
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Key to males of 
<taxonomicName id="C6BC4D48FFC59B22D36AF9C5FE5C6130" box="[351,493,1544,1570]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="genus">
<emphasis id="33C8EAD9FFC59B22D36AF9C5FE5C6130" bold="true" box="[351,493,1544,1570]" italics="true" pageId="3" pageNumber="496">Platystigma</emphasis>
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(male of 
<taxonomicName id="C6BC4D48FFC59B22D2C9F9E1FED96151" box="[252,360,1580,1603]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="pronoti">
<emphasis id="33C8EAD9FFC59B22D2C9F9E1FED96151" box="[252,360,1580,1603]" italics="true" pageId="3" pageNumber="496">P. pronoti</emphasis>
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unknown)
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1 Hw with black or dark brown band. Species known only from Colombia, Peru and Venezuela............... 
<taxonomicName id="C6BC4D48FFC59B22D72FF9BEFAEF619A" box="[1306,1374,1651,1672]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="jocaste">
<emphasis id="33C8EAD9FFC59B22D72FF9BEFAEF619A" box="[1306,1374,1651,1672]" italics="true" pageId="3" pageNumber="496">jocaste</emphasis>
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group
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<paragraph id="010336CBFFC59B22D2A2F95DFA2D61D0" blockId="3.[151,1436,1650,2028]" pageId="3" pageNumber="496">
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- Hw without black or dark brown band. Species of Northern and Southeastern Brazil, Bolivia, Ecuador and Peru (
<figureCitation id="99872A4EFFC59B22D717F942FA2661B7" box="[1314,1431,1679,1701]" captionStart="FIGURES 25 – 28" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[236,1360,265,1872]" captionTargetId="figure@13.[151,1436,193,1940]" captionTargetPageId="13" captionText="FIGURES 25 – 28. Wings in P. astictum male, Espírito Santo, Hw non saturated (25), saturated (26); female, Itatiaia, Rio de Janeiro, Hw non saturated (27) saturated (28)." figureDoi="http://doi.org/10.5281/zenodo.376932" httpUri="https://zenodo.org/record/376932/files/figure.png" pageId="3" pageNumber="496">Figs. 25, 27</figureCitation>
, 
<figureCitation id="99872A4EFFC59B22D2D4F961FEAE61D0" box="[225,287,1708,1730]" captionStart="FIGURES 29 – 32" captionStartId="14.[151,264,1951,1974]" captionTargetBox="[207,1393,228,1899]" captionTargetId="figure@14.[151,1436,193,1930]" captionTargetPageId="14" captionText="FIGURES 29 – 32. P. minimum, sp. nov. male, holotype, Fw non saturated (29), saturated (30). P. humaita sp. nov. male, holotype, Hw non saturated (31). P. quadratum sp. nov. male, holotype, Hw non saturated (32)." figureDoi="http://doi.org/10.5281/zenodo.376933" httpUri="https://zenodo.org/record/376933/files/figure.png" pageId="3" pageNumber="496">31, 32</figureCitation>
, 
<figureCitation id="99872A4EFFC59B22D31CF961FEF361D0" box="[297,322,1708,1730]" captionStart="FIGURES 33 – 36" captionStartId="15.[151,264,1974,1997]" captionTargetBox="[194,1404,194,1933]" captionTargetId="figure@15.[156,1430,151,1965]" captionTargetPageId="15" captionText="FIGURES 33 – 36. P. pronoti reproduced from Machado &amp; Lacerda (2016), female holotype, Fw non saturated (33), saturated (34), Hw non saturated (35), saturated (36)." figureDoi="http://doi.org/10.5281/zenodo.376934" httpUri="https://zenodo.org/record/376934/files/figure.png" pageId="3" pageNumber="496">35</figureCitation>
)........................................................................... 
<taxonomicName id="C6BC4D48FFC59B22D6E5F961FA8861D0" box="[1232,1337,1708,1730]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="quadratum">
<emphasis id="33C8EAD9FFC59B22D6E5F961FA8861D0" box="[1232,1337,1708,1730]" italics="true" pageId="3" pageNumber="496">quadratum</emphasis>
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group ...2
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<paragraph id="010336CBFFC59B22D2A2F907FA2A61F2" blockId="3.[151,1436,1650,2028]" box="[151,1435,1738,1760]" pageId="3" pageNumber="496">
<keyLead id="BA4D95FEFFC59B22D2A2F907FA2A61F2" box="[151,1435,1738,1760]" pageId="3" pageNumber="496">
2 Border of Hw with subapical elevation (
<figureCitation id="99872A4EFFC59B22D057F907FD1861F2" box="[610,681,1738,1760]" captionStart="FIGURES 25 – 28" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[236,1360,265,1872]" captionTargetId="figure@13.[151,1436,193,1940]" captionTargetPageId="13" captionText="FIGURES 25 – 28. Wings in P. astictum male, Espírito Santo, Hw non saturated (25), saturated (26); female, Itatiaia, Rio de Janeiro, Hw non saturated (27) saturated (28)." figureDoi="http://doi.org/10.5281/zenodo.376932" httpUri="https://zenodo.org/record/376932/files/figure.png" pageId="3" pageNumber="496">Fig. 25</figureCitation>
).............................................................3
</keyLead>
</paragraph>
<paragraph id="010336CBFFC59B22D2A2F925FA2A61EF" blockId="3.[151,1436,1650,2028]" box="[151,1435,1767,1789]" pageId="3" pageNumber="496">
<keyLead id="BA4D95FEFFC59B22D2A2F925FA2A61EF" box="[151,1435,1767,1789]" pageId="3" pageNumber="496">
- Border of Hw without subapical elevation (
<figureCitation id="99872A4EFFC59B22D0B4F92AFD6061EF" box="[641,721,1767,1789]" captionStart="FIGURES 25 – 28" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[236,1360,265,1872]" captionTargetId="figure@13.[151,1436,193,1940]" captionTargetPageId="13" captionText="FIGURES 25 – 28. Wings in P. astictum male, Espírito Santo, Hw non saturated (25), saturated (26); female, Itatiaia, Rio de Janeiro, Hw non saturated (27) saturated (28)." figureDoi="http://doi.org/10.5281/zenodo.376932" httpUri="https://zenodo.org/record/376932/files/figure.png" pageId="3" pageNumber="496">Figs. 27</figureCitation>
, 
<figureCitation id="99872A4EFFC59B22D0EFF92AFCA861EF" box="[730,793,1767,1789]" captionStart="FIGURES 29 – 32" captionStartId="14.[151,264,1951,1974]" captionTargetBox="[207,1393,228,1899]" captionTargetId="figure@14.[151,1436,193,1930]" captionTargetPageId="14" captionText="FIGURES 29 – 32. P. minimum, sp. nov. male, holotype, Fw non saturated (29), saturated (30). P. humaita sp. nov. male, holotype, Hw non saturated (31). P. quadratum sp. nov. male, holotype, Hw non saturated (32)." figureDoi="http://doi.org/10.5281/zenodo.376933" httpUri="https://zenodo.org/record/376933/files/figure.png" pageId="3" pageNumber="496">31, 32</figureCitation>
, 
<figureCitation id="99872A4EFFC59B22D117F92AFC8A61EF" box="[802,827,1767,1789]" captionStart="FIGURES 33 – 36" captionStartId="15.[151,264,1974,1997]" captionTargetBox="[194,1404,194,1933]" captionTargetId="figure@15.[156,1430,151,1965]" captionTargetPageId="15" captionText="FIGURES 33 – 36. P. pronoti reproduced from Machado &amp; Lacerda (2016), female holotype, Fw non saturated (33), saturated (34), Hw non saturated (35), saturated (36)." figureDoi="http://doi.org/10.5281/zenodo.376934" httpUri="https://zenodo.org/record/376934/files/figure.png" pageId="3" pageNumber="496">35</figureCitation>
)............................................... 5
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<keyStep id="BA482E6EFFC59B22D2A2F8C9FA2A602A" pageId="3" pageNumber="496">
<paragraph id="010336CBFFC59B22D2A2F8C9FA2D6008" blockId="3.[151,1436,1650,2028]" box="[151,1436,1796,1818]" pageId="3" pageNumber="496">
<keyLead id="BA4D95FEFFC59B22D2A2F8C9FA2D6008" box="[151,1436,1796,1818]" pageId="3" pageNumber="496">
3 Fw with pale apical area....................................................................... 
<taxonomicName id="C6BC4D48FFC59B22D71FF8C8FA2D6008" box="[1322,1436,1797,1818]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="33C8EAD9FFC59B22D71FF8C8FA2D6008" box="[1322,1436,1797,1818]" italics="true" pageId="3" pageNumber="496">P. martinezi</emphasis>
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<paragraph id="010336CBFFC59B22D2A2F8EEFA2A602A" blockId="3.[151,1436,1650,2028]" box="[151,1435,1826,1848]" pageId="3" pageNumber="496">
<keyLead id="BA4D95FEFFC59B22D2A2F8EEFA2A602A" box="[151,1435,1826,1848]" pageId="3" pageNumber="496">- Fw without pale apical area............................................................................ 4</keyLead>
</paragraph>
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<paragraph id="010336CBFFC59B22D2A2F8F2FA2D6047" blockId="3.[151,1436,1650,2028]" box="[151,1436,1855,1877]" pageId="3" pageNumber="496">
<keyLead id="BA4D95FEFFC59B22D2A2F8F2FA2D6047" box="[151,1436,1855,1877]" pageId="3" pageNumber="496">
4 Hw with pale apical area........................................................................ 
<taxonomicName id="C6BC4D48FFC59B22D701F88DFA2D6047" box="[1332,1436,1855,1877]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="buckleyi">
<emphasis id="33C8EAD9FFC59B22D701F88DFA2D6047" box="[1332,1436,1855,1877]" italics="true" pageId="3" pageNumber="496">P. buckleyi</emphasis>
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<paragraph id="010336CBFFC59B22D2A2F890FA2D6060" blockId="3.[151,1436,1650,2028]" box="[151,1436,1884,1906]" pageId="3" pageNumber="496">
<keyLead id="BA4D95FEFFC59B22D2A2F890FA2D6060" box="[151,1436,1884,1906]" pageId="3" pageNumber="496">
- Hw without pale apical area (
<figureCitation id="99872A4EFFC59B22D3CFF891FDF16060" box="[506,576,1884,1906]" captionStart="FIGURES 25 – 28" captionStartId="13.[151,264,1961,1984]" captionTargetBox="[236,1360,265,1872]" captionTargetId="figure@13.[151,1436,193,1940]" captionTargetPageId="13" captionText="FIGURES 25 – 28. Wings in P. astictum male, Espírito Santo, Hw non saturated (25), saturated (26); female, Itatiaia, Rio de Janeiro, Hw non saturated (27) saturated (28)." figureDoi="http://doi.org/10.5281/zenodo.376932" httpUri="https://zenodo.org/record/376932/files/figure.png" pageId="3" pageNumber="496">Fig. 25</figureCitation>
).............................................................. 
<taxonomicName id="C6BC4D48FFC59B22D707F890FA2D6060" box="[1330,1436,1885,1906]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="astictum">
<emphasis id="33C8EAD9FFC59B22D707F890FA2D6060" box="[1330,1436,1885,1906]" italics="true" pageId="3" pageNumber="496">P. astictum</emphasis>
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<paragraph id="010336CBFFC59B22D2A2F8B1FA2D60A3" blockId="3.[151,1436,1650,2028]" pageId="3" pageNumber="496">
<keyLead id="BA4D95FEFFC59B22D2A2F8B1FA2D60A3" pageId="3" pageNumber="496">
5 Small species (Hw 32, abdomen 48). Cleft of anterior lamina with elongated medial border sclerotized apically (
<figureCitation id="99872A4EFFC59B22D725F8B1FAE66080" box="[1296,1367,1916,1938]" captionStart="FIGURES 37 – 41" captionStartId="16.[151,264,1976,1999]" captionTargetBox="[254,1339,240,1915]" captionTargetId="figure@16.[165,1422,141,1955]" captionTargetPageId="16" captionText="FIGURES 37 – 41. Genitalia of second segment in P. astictum, Tinguá, Rio de Janeiro (37); P. martinezi, Bolívia, Santa Cruz (38); P. humaita sp. nov., holotype (39); P. minimum sp. nov., holotype (40); P. quadratum sp. nov., holotype (41)." figureDoi="http://doi.org/10.5281/zenodo.376935" httpUri="https://zenodo.org/record/376935/files/figure.png" pageId="3" pageNumber="496">Fig. 40</figureCitation>
). Penis segment 3 subrectangular (
<figureCitation id="99872A4EFFC59B22D3D6F856FD9F60A3" box="[483,558,1947,1969]" captionStart="FIGURES 42 – 51" captionStartId="17.[151,264,1928,1951]" captionTargetBox="[264,1319,197,1865]" captionTargetId="figure@17.[204,1372,153,1898]" captionTargetPageId="17" captionText="FIGURES 42 – 51. Penis of P. astictum, Tinguá, Rio de Janeiro, in ectal (42) and lateral (43) views; P. martinezi, Bolívia, Santa Cruz, in ectal (44) and lateral (45) views. P. humaita sp. nov. in ectal (46) and lateral (47) views; P. minimum sp. nov. in ectal (48) and lateral (49) views; P. quadratum sp. nov. in ectal (50) and lateral views (51)." figureDoi="http://doi.org/10.5281/zenodo.376936" httpUri="https://zenodo.org/record/376936/files/figure.png" pageId="3" pageNumber="496">Fig. 48</figureCitation>
)............................................................... 
<taxonomicName id="C6BC4D48FFC59B22D71EF851FA2D60A3" box="[1323,1436,1948,1969]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="496" phylum="Arthropoda" rank="species" species="minimum">
<emphasis id="33C8EAD9FFC59B22D71EF851FA2D60A3" box="[1323,1436,1948,1969]" italics="true" pageId="3" pageNumber="496">P. minimum</emphasis>
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<paragraph id="010336CBFFC59B22D2A2F874FA2D60FE" blockId="3.[151,1436,1650,2028]" pageId="3" pageNumber="496">
<keyLead id="BA4D95FEFFC59B22D2A2F874FA2D60FE" pageId="3" pageNumber="496">
- Larger species (Hw 40–48, abdomen 52–65). Cleft of anterior lamina without elongated medial border. Penis segment 3 sub- quadrate (
<figureCitation id="99872A4EFFC59B22D375F81BFE0560FE" box="[320,436,2006,2028]" captionStart="FIGURES 42 – 51" captionStartId="17.[151,264,1928,1951]" captionTargetBox="[264,1319,197,1865]" captionTargetId="figure@17.[204,1372,153,1898]" captionTargetPageId="17" captionText="FIGURES 42 – 51. Penis of P. astictum, Tinguá, Rio de Janeiro, in ectal (42) and lateral (43) views; P. martinezi, Bolívia, Santa Cruz, in ectal (44) and lateral (45) views. P. humaita sp. nov. in ectal (46) and lateral (47) views; P. minimum sp. nov. in ectal (48) and lateral (49) views; P. quadratum sp. nov. in ectal (50) and lateral views (51)." figureDoi="http://doi.org/10.5281/zenodo.376936" httpUri="https://zenodo.org/record/376936/files/figure.png" pageId="3" pageNumber="496">Figs. 46, 50</figureCitation>
)................................................................................. 6
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</paragraph>
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<keyStep id="BA482E6EFFC29B25D2A2FF5AFA2A67D8" pageId="4" pageNumber="497">
<paragraph id="010336CBFFC29B25D2A2FF5AFA2D67BF" blockId="4.[151,1436,151,202]" box="[151,1436,151,173]" pageId="4" pageNumber="497">
<keyLead id="BA4D95FEFFC29B25D2A2FF5AFA2D67BF" box="[151,1436,151,173]" pageId="4" pageNumber="497">
6 Pale apical area present in Fw and Hw (
<figureCitation id="99872A4EFFC29B25D06BFF5AFD1567BF" box="[606,676,151,173]" captionStart="FIGURES 29 – 32" captionStartId="14.[151,264,1951,1974]" captionTargetBox="[207,1393,228,1899]" captionTargetId="figure@14.[151,1436,193,1930]" captionTargetPageId="14" captionText="FIGURES 29 – 32. P. minimum, sp. nov. male, holotype, Fw non saturated (29), saturated (30). P. humaita sp. nov. male, holotype, Hw non saturated (31). P. quadratum sp. nov. male, holotype, Hw non saturated (32)." figureDoi="http://doi.org/10.5281/zenodo.376933" httpUri="https://zenodo.org/record/376933/files/figure.png" pageId="4" pageNumber="497">Fig. 31</figureCitation>
)..................................................... 
<taxonomicName id="C6BC4D48FFC29B25D700FF55FA2D67BF" box="[1333,1436,151,173]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="humaita">
<emphasis id="33C8EAD9FFC29B25D700FF55FA2D67BF" box="[1333,1436,151,173]" italics="true" pageId="4" pageNumber="497">P. humaita</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="010336CBFFC29B25D2A2FF78FA2A67D8" blockId="4.[151,1436,151,202]" box="[151,1435,180,202]" pageId="4" pageNumber="497">
<keyLead id="BA4D95FEFFC29B25D2A2FF78FA2A67D8" box="[151,1435,180,202]" pageId="4" pageNumber="497">
- Pale apical area present only in Hw (
<figureCitation id="99872A4EFFC29B25D074FF79FD3667D8" box="[577,647,180,202]" captionStart="FIGURES 29 – 32" captionStartId="14.[151,264,1951,1974]" captionTargetBox="[207,1393,228,1899]" captionTargetId="figure@14.[151,1436,193,1930]" captionTargetPageId="14" captionText="FIGURES 29 – 32. P. minimum, sp. nov. male, holotype, Fw non saturated (29), saturated (30). P. humaita sp. nov. male, holotype, Hw non saturated (31). P. quadratum sp. nov. male, holotype, Hw non saturated (32)." figureDoi="http://doi.org/10.5281/zenodo.376933" httpUri="https://zenodo.org/record/376933/files/figure.png" pageId="4" pageNumber="497">Fig. 32</figureCitation>
)...................................................... 
<taxonomicName id="C6BC4D48FFC29B25D72FFF78FA2A67D8" box="[1306,1435,180,202]" class="Insecta" family="Pseudostigmatidae" genus="Platystigma" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="497" phylum="Arthropoda" rank="species" species="quadratum">
<emphasis id="33C8EAD9FFC29B25D72FFF78FA2A67D8" box="[1306,1435,180,202]" italics="true" pageId="4" pageNumber="497">P. quadratum</emphasis>
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