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<document id="A068098EFC119ADBCE31951DB47AC2F0" ID-DOI="10.5281/zenodo.174029" ID-GBIF-Dataset="d9d8d147-e8fc-4014-8af2-aa2fd28c20df" ID-ISSN="1175­5326" ID-Zenodo-Dep="174029" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790304226" checkinUser="plazi" docAuthor="Schawaller, Wolfgang" docDate="2006" docId="7F54080DFFE8FF82D56AD80FFB71FD83" docLanguage="en" docName="zt01325p384.pdf" docOrigin="Zootaxa 1325" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Laena rolandi Schawaller, 2006, n. sp." docType="treatment" docVersion="9" lastPageNumber="375" masterDocId="836D7075FFEDFF84D462DD29FF82FFF5" masterDocTitle="New species of the genus Laena Latreille (Coleoptera: Tenebrionidae) from Southeastern Asia *" masterLastPageNumber="384" masterPageNumber="369" pageNumber="374" updateTime="1698219154779" updateUser="plazi">
<mods:mods id="CE6E15A70DA2358D03231037744C50B5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="116EC02B9C33F74E77EB1F2C47597214">
<mods:title id="295AF5378BFF14080AF588C97F58CF9C">New species of the genus Laena Latreille (Coleoptera: Tenebrionidae) from Southeastern Asia *</mods:title>
</mods:titleInfo>
<mods:name id="A3B9E221BBC4F82C4A67EB8F4B0D9A25" type="personal">
<mods:role id="8CBF0C4AAB2EBCB9906FEFB3FD5E8119">
<mods:roleTerm id="8EABC82EB7540EDDDBF48F681C9D19EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D6DD1DEF454335C29893C5F5E67E2B5D">Schawaller, Wolfgang</mods:namePart>
</mods:name>
<mods:typeOfResource id="E932C0FC85BED3732E32D2423C9225A9">text</mods:typeOfResource>
<mods:relatedItem id="438720A1F88FF7F93D975DCDDC476ECB" type="host">
<mods:titleInfo id="5474BF92792A2012E4BE453E5CBD106C">
<mods:title id="D1A2F0C16979C32342F67406ABF6C334">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EB51D01383012649B9EA40C3AC314980">
<mods:date id="33EACDCCF1D144A2161DFC4E216E89C2">2006</mods:date>
<mods:detail id="53B66BB69FD7BF45DE94F1B65675DF09" type="volume">
<mods:number id="CA13DB5F0292F17288A0E98BBFDCA47C">1325</mods:number>
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<mods:classification id="AAB5B36688CD531F9F5AD8C19CA5DFB9">journal article</mods:classification>
<mods:identifier id="8A4CB4E69A63B2702AAB4534B06890C2" type="DOI">10.5281/zenodo.174029</mods:identifier>
<mods:identifier id="D01A2B4CAA49A5E651C303F1E7987F82" type="GBIF-Dataset">d9d8d147-e8fc-4014-8af2-aa2fd28c20df</mods:identifier>
<mods:identifier id="DA214539628E582A02CFCDBD2CA41CF4" type="ISSN">1175­5326</mods:identifier>
<mods:identifier id="5BC25AF420E10BA497716B4DFDECD896" type="Zenodo-Dep">174029</mods:identifier>
</mods:mods>
<treatment id="7F54080DFFE8FF82D56AD80FFB71FD83" ID-DOI="http://doi.org/10.5281/zenodo.6258203" ID-GBIF-Taxon="119350251" ID-Zenodo-Dep="6258203" LSID="urn:lsid:plazi:treatment:7F54080DFFE8FF82D56AD80FFB71FD83" httpUri="http://treatment.plazi.org/id/7F54080DFFE8FF82D56AD80FFB71FD83" lastPageId="6" lastPageNumber="375" pageId="5" pageNumber="374">
<subSubSection id="BFE7EA90FFE8FF81D56AD80FFD1EFAB5" box="[264,668,1318,1344]" pageId="5" pageNumber="374" type="nomenclature">
<paragraph id="F742B91BFFE8FF81D56AD80FFD1EFAB5" blockId="5.[264,668,1318,1344]" box="[264,668,1318,1344]" pageId="5" pageNumber="374">
<heading id="AC0A0E77FFE8FF81D56AD80FFD1EFAB5" bold="true" box="[264,668,1318,1344]" fontSize="11" level="1" pageId="5" pageNumber="374" reason="1">
<emphasis id="C5896509FFE8FF81D56AD80FFD1EFAB5" bold="true" box="[264,668,1318,1344]" pageId="5" pageNumber="374">
<taxonomicName id="30FDC298FFE8FF81D56AD80FFE31FAB5" box="[264,435,1318,1344]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="rolandi" status="sp. nov.">
<emphasis id="C5896509FFE8FF81D56AD80FFE31FAB5" bold="true" box="[264,435,1318,1344]" italics="true" pageId="5" pageNumber="374">Laena rolandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE8FF81D5D9D80FFE7CFAB5" box="[443,510,1318,1344]" pageId="5" pageNumber="374" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="6FC6A59EFFE8FF81D66ED80FFDEBFAB5" box="[524,617,1318,1344]" captionStart="FIGURES 9 – 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 – 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="5" pageNumber="374">Figs 10</figureCitation>
,
<figureCitation id="6FC6A59EFFE8FF81D614D80FFD17FAB5" box="[630,661,1318,1344]" captionStart="FIGURES 17 – 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 – 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="5" pageNumber="374">24</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFE8FF81D56AD858FD19F9DE" pageId="5" pageNumber="374" type="materials_examined">
<paragraph id="F742B91BFFE8FF81D56AD858FE3EFA46" blockId="5.[264,1325,1393,1899]" pageId="5" pageNumber="374">
<typeStatus id="284607B9FFE8FF81D56AD858FEF8FA7E" box="[264,378,1393,1419]" pageId="5" pageNumber="374" type="holotype">
<emphasis id="C5896509FFE8FF81D56AD858FEF8FA7E" bold="true" box="[264,378,1393,1419]" pageId="5" pageNumber="374">Holotype</emphasis>
</typeStatus>
(♂): N
<collectingCountry id="8FEAF98BFFE8FF81D586D858FDCDFA7E" box="[484,591,1393,1419]" name="Thailand" pageId="5" pageNumber="374">Thailand</collectingCountry>
, Chiang Mai, Doi Pui,
<quantity id="300514FEFFE8FF81D718D858FBA6FA7E" box="[890,1060,1393,1419]" metricMagnitude="3" metricUnit="m" metricValue="1.6425" metricValueMax="1.685" metricValueMin="1.6" pageId="5" pageNumber="374" unit="m" value="1642.5" valueMax="1685.0" valueMin="1600.0">1600–1685 m</quantity>
,
<date id="83439FDBFFE8FF81D054D858FB44FA7E" box="[1078,1222,1393,1419]" pageId="5" pageNumber="374" value="2004-05-07" valueMax="2004-05-09" valueMin="2004-05-07">7.–9.V.2004</date>
, leg. R. Grimm,
<collectionCode id="91EC21DEFFE8FF81D509D8B0FE3AFA46" box="[363,440,1433,1459]" pageId="5" pageNumber="374">CRGT</collectionCode>
.
</paragraph>
<paragraph id="F742B91BFFE8FF81D55AD8E8FD19F9DE" blockId="5.[264,1325,1393,1899]" pageId="5" pageNumber="374">
<typeStatus id="284607B9FFE8FF81D55AD8E8FE2CFA2E" box="[312,430,1473,1499]" pageId="5" pageNumber="374" type="paratype">Paratypes</typeStatus>
: N
<collectingCountry id="8FEAF98BFFE8FF81D5B5D8E8FDC0FA2E" box="[471,578,1473,1499]" name="Thailand" pageId="5" pageNumber="374">Thailand</collectingCountry>
, Chiang Mai, Doi Suthep Pui NP,
<date id="83439FDBFFE8FF81D7B8D8E8FBD7FA2E" box="[986,1109,1473,1499]" pageId="5" pageNumber="374" value="1999-05-30">30.V.1999</date>
, leg. R. Grimm, 1 ex.
<collectionCode id="91EC21DEFFE8FF81D556D8C0FE03F9F6" box="[308,385,1513,1539]" pageId="5" pageNumber="374">CRGT</collectionCode>
. – N
<collectingCountry id="8FEAF98BFFE8FF81D5A7D8C0FDB2F9F6" box="[453,560,1513,1539]" name="Thailand" pageId="5" pageNumber="374">Thailand</collectingCountry>
, Chiang Mai, Doi Pui,
<quantity id="300514FEFFE8FF81D72AD8C0FC6DF9F6" box="[840,1007,1513,1539]" metricMagnitude="3" metricUnit="m" metricValue="1.6425" metricValueMax="1.685" metricValueMin="1.6" pageId="5" pageNumber="374" unit="m" value="1642.5" valueMax="1685.0" valueMin="1600.0">1600–1685 m</quantity>
,
<date id="83439FDBFFE8FF81D79FD8C0FB4FF9F6" box="[1021,1229,1513,1539]" pageId="5" pageNumber="374" value="2003-04-23" valueMax="2003-05-12" valueMin="2003-04-23">23.IV.–12.V.2003</date>
, leg. R. Grimm, 5 ex.
<collectionCode id="91EC21DEFFE8FF81D5CEDB38FE7BF9DE" box="[428,505,1553,1579]" pageId="5" pageNumber="374">CRGT</collectionCode>
, 2 ex.
<collectionCode id="91EC21DEFFE8FF81D621DB38FD15F9DE" LSID="urn:lsid:biocol.org:col:34840" box="[579,663,1553,1579]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="5" pageNumber="374">SMNS</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFE8FF81D55ADB10FD1BF98E" pageId="5" pageNumber="374" type="etymology">
<paragraph id="F742B91BFFE8FF81D55ADB10FD1BF98E" blockId="5.[264,1325,1393,1899]" pageId="5" pageNumber="374">
<emphasis id="C5896509FFE8FF81D55ADB10FE4AF9A6" bold="true" box="[312,456,1593,1619]" pageId="5" pageNumber="374">Etymology:</emphasis>
Dedicated to Dr. Roland Grimm (Tübingen), collector of the
<typeStatus id="284607B9FFE8FF81D0C7DB10FB55F9A6" box="[1189,1239,1593,1619]" pageId="5" pageNumber="374">type</typeStatus>
series, for fruitful long­term cooperation.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFE8FF82D55ADBA0FD48FEC3" lastPageId="6" lastPageNumber="375" pageId="5" pageNumber="374" type="diagnosis">
<paragraph id="F742B91BFFE8FF82D55ADBA0FD48FEC3" blockId="5.[264,1325,1393,1899]" lastBlockId="6.[264,1323,284,630]" lastPageId="6" lastPageNumber="375" pageId="5" pageNumber="374">
<emphasis id="C5896509FFE8FF81D55ADBA0FE38F956" bold="true" box="[312,442,1673,1699]" pageId="5" pageNumber="374">Diagnosis:</emphasis>
<taxonomicName id="30FDC298FFE8FF81D5A6DBA0FDEEF957" box="[452,620,1673,1698]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="rolandi" status="sp. nov.">
<emphasis id="C5896509FFE8FF81D5A6DBA0FDEEF957" box="[452,620,1673,1698]" italics="true" pageId="5" pageNumber="374">Laena rolandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="DEBAD872FFE8FF81D614DBA0FD3EF956" box="[630,700,1673,1699]" pageId="5" pageNumber="374" rank="species">
<emphasis id="C5896509FFE8FF81D614DBA0FD3EF956" bold="true" box="[630,700,1673,1699]" pageId="5" pageNumber="374">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="30FDC298FFE8FF81D698DBA0FB70F956" authority="Kaszab &amp; Chujo, 1966" authorityName="Kaszab &amp; Chujo" authorityYear="1966" box="[762,1266,1673,1699]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="C5896509FFE8FF81D698DBA0FC51F957" box="[762,979,1673,1698]" italics="true" pageId="5" pageNumber="374">Laena thailandica</emphasis>
<bibRefCitation id="936CC4EAFFE8FF81D7BEDBA0FB70F956" author="Kaszab" box="[988,1266,1673,1699]" pageId="5" pageNumber="384" refString="Kaszab, Z. &amp; Chujo, M. (1966) Coleoptera from Southeast Asia (V), 14. Familie Tenebrionidae (1). Memoirs of the Faculty of Education, Kagawa University, part II. no. 140, 51 - 56." type="journal article" year="1966">Kaszab &amp; Chûjô, 1966</bibRefCitation>
</taxonomicName>
(see photo
<quantity id="300514FEFFE8FF81D535DB98FE0BF93E" box="[343,393,1713,1739]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="5" pageNumber="374" unit="in" value="6.0">6 in</quantity>
<bibRefCitation id="936CC4EAFFE8FF81D5F7DB98FD29F93E" author="Kaszab" box="[405,683,1713,1739]" pageId="5" pageNumber="384" refString="Kaszab, Z. &amp; Chujo, M. (1966) Coleoptera from Southeast Asia (V), 14. Familie Tenebrionidae (1). Memoirs of the Faculty of Education, Kagawa University, part II. no. 140, 51 - 56." type="journal article" year="1966">Kaszab &amp; Chûjô 1966</bibRefCitation>
) from southern
<collectingCountry id="8FEAF98BFFE8FF81D714DB98FC5DF93E" box="[886,991,1713,1739]" name="Thailand" pageId="5" pageNumber="374">Thailand</collectingCountry>
share the small body size, unarmed legs and the glabrous surface, but can be separated by completely different shaped pronotum (round and widest in the middle in
<taxonomicName id="30FDC298FFE8FF81D7E2DA28FB87F8EF" box="[896,1029,1793,1818]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="C5896509FFE8FF81D7E2DA28FB87F8EF" box="[896,1029,1793,1818]" italics="true" pageId="5" pageNumber="374">thailandica</emphasis>
</taxonomicName>
), dense nearly confluent pronotal punctation (sparse in
<taxonomicName id="30FDC298FFE8FF81D611DA00FD7AF8B7" box="[627,760,1833,1858]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="C5896509FFE8FF81D611DA00FD7AF8B7" box="[627,760,1833,1858]" italics="true" pageId="5" pageNumber="374">thailandica</emphasis>
</taxonomicName>
), unbordered lateral pronotal margin (bordered in
<taxonomicName id="30FDC298FFE8FF81D54FDA78FE31F89F" box="[301,435,1873,1898]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="C5896509FFE8FF81D54FDA78FE31F89F" box="[301,435,1873,1898]" italics="true" pageId="5" pageNumber="374">thailandica</emphasis>
</taxonomicName>
) and narrower elytral intervals. The aedeagus of
<taxonomicName id="30FDC298FFE8FF81D04EDA78FA8AF89F" box="[1068,1288,1873,1898]" class="Insecta" family="Terebellidae" genus="Laena" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="374" phylum="Arthropoda" rank="species" species="thailandica">
<emphasis id="C5896509FFE8FF81D04EDA78FA8AF89F" box="[1068,1288,1873,1898]" italics="true" pageId="5" pageNumber="374">Laena thailandica</emphasis>
</taxonomicName>
is unknown and so can not be compared.
</paragraph>
</subSubSection>
<subSubSection id="BFE7EA90FFEBFF82D55ADC6AFB71FD83" pageId="6" pageNumber="375" type="description">
<paragraph id="F742B91BFFEBFF82D55ADC6AFB71FD83" blockId="6.[264,1323,284,630]" pageId="6" pageNumber="375">
<emphasis id="C5896509FFEBFF82D55ADC6AFE50FEA8" bold="true" box="[312,466,323,349]" pageId="6" pageNumber="375">Description:</emphasis>
Body length 3.8–5.0 mm. Eyes somewhat prominent. Pronotum (
<figureCitation id="6FC6A59EFFEBFF82D099DC6DFEA4FE73" captionStart="FIGURES 9 – 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 – 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="6" pageNumber="375">Fig. 10</figureCitation>
) with large punctures, distance between 0.5–3 diameters, most punctures bearing short adpressed seta; surface flat, shining; lateral margin unbordered; propleura with sparser punctation, nearly without setation. Elytra (
<figureCitation id="6FC6A59EFFEBFF82D740DC95FCFFFE23" box="[802,893,444,470]" captionStart="FIGURES 9 – 16" captionStartId="10.[264,382,1733,1757]" captionTargetBox="[283,1306,328,1706]" captionTargetId="figure@10.[283,1309,326,1718]" captionTargetPageId="10" captionText="FIGURES 9 – 16. Dorsal view of Laena species. 9 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 10 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 11 Laena siamica, Ψ paratype HNHM, Thailand. 12 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 13 Laena tonkinensis, non­type Ψ HNHM, Vietnam. 14 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 15 Laena vietnamica, non­type ♂ HNHM, Vietnam. 16 Laena thailandica, Ψ holotype HNHM, Thailand." httpUri="https://zenodo.org/record/174031/files/figure.png" pageId="6" pageNumber="375">Fig. 10</figureCitation>
) with complete rows of punctures without striae, punctures equal in size to pronotal punctures, all without setae; intervals with very few scattered punctures, without setae, interval 9 with 3 distinct setiferous pores; all intervals slightly convex, shining. Femora in both sexes without distinct tooth or other modification. Tibiae in males without distinct sexual character. Aedeagus see
<figureCitation id="6FC6A59EFFEBFF82D0FADF75FB6DFD83" box="[1176,1263,604,630]" captionStart="FIGURES 17 – 26" captionStartId="12.[264,382,1382,1406]" captionTargetBox="[346,1246,319,1358]" captionTargetId="figure@12.[342,1249,316,1361]" captionTargetPageId="12" captionText="FIGURES 17 – 26. Aedeagus of Laena species. 17 Laena inthanonica n. sp., ♂ holotype MHNG, Thailand. 18 Laena jocheni n. sp., ♂ holotype SMNS, Laos. 19 Laena kenyirica n. sp., ♂ holotype SMNS, Malaysia. 20 Laena khaolaka n. sp., ♂ holotype SMNS, Thailand. 21 Laena kiyoshii n. sp., ♂ holotype SMNS, Laos. 22 Laena moguntia n. sp., ♂ holotype CKAO, Vietnam. 23 Laena schulzi n. sp., ♂ holotype SMNS, Malaysia. 24 Laena rolandi n. sp., ♂ holotype CRGT, Thailand. 25 Laena tamdaoensis, ♂ holotype NSMT, Vietnam. 26 Laena vietnamica, non­type ♂ HNHM, Vietnam." httpUri="https://zenodo.org/record/174032/files/figure.png" pageId="6" pageNumber="375">Fig. 24</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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