treatments-xml/data/74/43/87/744387D5B112A133EBADFD65E08DFD2D.xml

<document id="A08BAE391BBA1FDEEC73322698D11E7B" ID-DOI="10.11646/zootaxa.3846.4.1" ID-GBIF-Dataset="317a43cd-c408-4fde-8374-357fbb9a0d53" ID-ISSN="1175-5326" ID-Zenodo-Dep="250495" ID-ZooBank="0DD5A603-D65F-4976-BBE9-94DA7110053F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467125281" checkinUser="plazi" docAuthor="Jacobsen, Niels H. G., Kuhn, Arianna L., Jackman, Todd R. &amp; Bauer, Aaron M." docDate="2014" docId="744387D5B112A133EBADFD65E08DFD2D" docLanguage="en" docName="zt03846p501.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afroedura pongola Jacobsen, Kuhn, Jackman &amp; Bauer, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="482" masterDocId="887AFFADB10CA12CEB3AFFD1E06AFF85" masterDocTitle="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa" masterLastPageNumber="501" masterPageNumber="451" pageNumber="481" updateTime="1698305158428" updateUser="plazi">
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<mods:title id="001899FA49E2EC1218FC702DA478DB07">A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa</mods:title>
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<mods:namePart id="FC24B6557B80FAFEC771503A2D40A977">Jacobsen, Niels H. G.</mods:namePart>
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<mods:namePart id="7CEEBBD8D31B22B36EB3651C80B9B5DC">Kuhn, Arianna L.</mods:namePart>
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<mods:namePart id="DBA8AE03F4E6229036EC5144D6C09EB1">Bauer, Aaron M.</mods:namePart>
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<subSubSection id="B4F06548B112A132EBADFD65E094FD6A" pageId="30" pageNumber="481" type="nomenclature">
<paragraph id="FC5536C3B112A132EBADFD65E188FD4B" blockId="30.[151,482,692,751]" box="[151,482,692,718]" pageId="30" pageNumber="481">
<heading id="A71D81AFB112A132EBADFD65E188FD4B" bold="true" box="[151,482,692,718]" fontSize="11" level="1" pageId="30" pageNumber="481" reason="1">
<emphasis id="CE9EEAD1B112A132EBADFD65E188FD4B" bold="true" box="[151,482,692,718]" pageId="30" pageNumber="481">
<taxonomicName id="3BEA4D40B112A132EBADFD65E114FD4B" box="[151,382,692,718]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="481" phylum="Chordata" rank="species" species="pongola" status="sp. nov.">
<emphasis id="CE9EEAD1B112A132EBADFD65E114FD4B" bold="true" box="[151,382,692,718]" italics="true" pageId="30" pageNumber="481">Afroedura pongola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5AD57AAB112A132EABEFD65E188FD4B" box="[388,482,692,718]" pageId="30" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="FC5536C3B112A132EBADFD07E094FD6A" blockId="30.[151,482,692,751]" box="[151,254,726,751]" pageId="30" pageNumber="481">
(
<figureCitation id="64D12A46B112A132EBA5FD07E088FD6A" box="[159,226,726,751]" captionStart="FIGURE 9" captionStartId="29.[151,250,1919,1941]" captionTargetBox="[218,1372,193,1889]" captionTargetId="figure@29.[218,1372,193,1891]" captionTargetPageId="29" captionText="FIGURE 9. New species in the Afroedura marleyi group. (A) Afroedura maripi sp. nov. from Mariepskop, Mpumalanga, South Africa (Photo Marius Burger). (B) Afroedura pongola sp. nov. from from west of Pongola, KwaZulu-Natal, South Africa (Photo Johan Marais). (C) Afroedura rondavelica sp. nov. from the Three Rondavels, Blyde River Nature Reserve, Mpumalanga Province, South Africa (24 ° 34 ' S, 30 ° 50 ' E) (Photo by N. H. G. Jacobsen)." httpUri="https://zenodo.org/record/250504/files/figure.png" pageId="30" pageNumber="481">Fig. 9</figureCitation>
B)
</paragraph>
</subSubSection>
<subSubSection id="B4F06548B112A132EBADFCCFE2A6FCD6" pageId="30" pageNumber="481" type="reference_group">
<paragraph id="FC5536C3B112A132EBADFCCFE2A6FCD6" blockId="30.[151,764,798,851]" pageId="30" pageNumber="481">
<taxonomicName id="3BEA4D40B112A132EBADFCCFE134FCB1" box="[151,350,798,820]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="481" phylum="Chordata" rank="species" species="pondolia">
<emphasis id="CE9EEAD1B112A132EBADFCCFE134FCB1" box="[151,350,798,820]" italics="true" pageId="30" pageNumber="481">Afroedura pondolia</emphasis>
</taxonomicName>
'Godlwayo' 
<bibRefCitation id="987B4B32B112A132EAE7FCCFE21EFCB1" author="Jacobsen" box="[477,628,798,820]" pageId="30" pageNumber="481" refString="Jacobsen, N. H. G. (1990) A Herpetological Survey of the Transvaal. Unpublished Ph. D. Dissertation, University of Natal, Durban, South Africa, 1621 pp." type="book" year="1990">Jacobsen 1990</bibRefCitation>
, 
<bibRefCitation id="987B4B32B112A132E944FCCFE2D4FCB1" author="Jacobsen" box="[638,702,798,820]" pageId="30" pageNumber="481" refString="Jacobsen, N. H. G. (1992 a) Flat geckos (genus Afroedura) in the Transvaal. Journal of the Herpetological Association of Africa, 40, 22 - 25. http: // dx. doi. org / 10.1080 / 04416651.1992.9650313" type="journal article" year="1992" yearSuffix="a">1992a</bibRefCitation>
, 
<bibRefCitation id="987B4B32B112A132E9F3FCCFE296FCB1" author="Jacobsen" box="[713,764,798,820]" pageId="30" pageNumber="481" refString="Jacobsen, N. H. G. (1997) Family Gekkonidae, Genus Afroedura. In: Van Wyk, J. H. (Ed.), Proceedings of the FitzSimons Commemorative Symposium, South African Lizards: 50 years of Progress and Third H. A. A. Symposium on African Herpetology, Transvaal Museum, Pretoria, South Africa, 11 - 15 October 1993. Herpetological Association of Africa, Stellenbosch, pp. 34 - 39." type="journal article" year="1997">1997</bibRefCitation>
<taxonomicName id="3BEA4D40B112A132EBADFCECE095FCD6" box="[151,255,829,851]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="481" phylum="Chordata" rank="genus" status="sp. nov.">
<emphasis id="CE9EEAD1B112A132EBADFCECE095FCD6" box="[151,255,829,851]" italics="true" pageId="30" pageNumber="481">Afroedura</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5AD57AAB112A132EA3EFCEFE139FCD6" box="[260,339,830,851]" pageId="30" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
“ 
<taxonomicName id="3BEA4D40B112A132EA58FCECE1DDFCD6" box="[354,439,829,851]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="483" phylum="Chordata" rank="species" species="pongola" status="sp. nov.">Pongola</taxonomicName>
flat gecko” 
<bibRefCitation id="987B4B32B112A132E909FCECE2A6FCD6" author="Bourquin" box="[563,716,829,851]" pageId="30" pageNumber="481" refString="Bourquin, O. (2004) Reptiles (Reptilia) in KwaZulu - Natal: 1 - Diversity and distribution. Durban Museum Novitates, 29, 57 - 103." type="journal article" year="2004">Bourquin 2004</bibRefCitation>
</paragraph>
</subSubSection>
<subSubSection id="B4F06548B112A132EBADFCAEE41DFBAD" pageId="30" pageNumber="481" type="materials_examined">
<paragraph id="FC5536C3B112A132EBADFCAEE31DFC39" blockId="30.[151,1437,895,2037]" pageId="30" pageNumber="481">
<emphasis id="CE9EEAD1B112A132EBADFCAEE161FC1D" bold="true" box="[151,267,895,920]" pageId="30" pageNumber="481">
<typeStatus id="23518861B112A132EBADFCAEE16CFC1D" box="[151,262,895,920]" pageId="30" pageNumber="481" type="holotype">Holotype</typeStatus>
.
</emphasis>
TM 81210, adult male, Godlwayo, 
<geoCoordinate id="99DE5004B112A132E99DFC51E363FC1D" box="[679,777,896,920]" direction="south" orientation="latitude" pageId="30" pageNumber="481" precision="925" value="-27.333334">27°20' S</geoCoordinate>
, 
<geoCoordinate id="99DE5004B112A132E82DFCAEE316FC12" box="[791,892,895,920]" direction="east" orientation="longitude" pageId="30" pageNumber="481" precision="925" value="31.416666">31°25' E</geoCoordinate>
, Piet Retief District (2731AD), Kwazulu-Natal, 
<collectingCountry id="84FD7653B112A132EBADFC75E1DEFC38" box="[151,436,932,957]" name="South Africa" pageId="30" pageNumber="481">Republic of South Africa</collectingCountry>
, collector R. E. Newbery, 
<date id="88541003B112A132E9E1FC75E319FC39" box="[731,883,932,957]" pageId="30" pageNumber="481" value="1981-05-24">24 May 1981</date>
.
</paragraph>
<paragraph id="FC5536C3B112A132EBFDFC16E41DFBAD" blockId="30.[151,1437,895,2037]" pageId="30" pageNumber="481">
<emphasis id="CE9EEAD1B112A132EBFDFC16E12CFC65" bold="true" box="[199,326,967,992]" pageId="30" pageNumber="481">
<typeStatus id="23518861B112A132EBFDFC16E12BFC65" box="[199,321,967,992]" pageId="30" pageNumber="481" type="paratype">Paratypes</typeStatus>
.
</emphasis>
TM 81211–81213, 81216 same data as 
<typeStatus id="23518861B112A132E829FC16E312FC65" box="[787,888,967,992]" pageId="30" pageNumber="481" type="holotype">holotype</typeStatus>
; TM 81208–81209, 81214, 81217–81218, same locality as 
<typeStatus id="23518861B112A132EA2AFC3DE11FFB80" box="[272,373,1004,1029]" pageId="30" pageNumber="481" type="holotype">holotype</typeStatus>
, collectors R. E. Newbery &amp; W. Petersen, 
<date id="88541003B112A132E875FC3DE443FB80" box="[847,1065,1004,1029]" pageId="30" pageNumber="481" value="1985-11-19">19 November 1985</date>
; TM 81215, 81219–81220, Farm Zwartkloof 60HU, Piet Retief district (2731BC), collector R. E. Newbery, 
<date id="88541003B112A132E8E1FBC1E419FBAD" box="[987,1139,1040,1064]" pageId="30" pageNumber="481" value="1981-05-21">21 May 1981</date>
.
</paragraph>
</subSubSection>
<subSubSection id="B4F06548B112A132EBFDFBE5E15AFBF5" pageId="30" pageNumber="481" type="etymology">
<paragraph id="FC5536C3B112A132EBFDFBE5E15AFBF5" blockId="30.[151,1437,895,2037]" pageId="30" pageNumber="481">
<emphasis id="CE9EEAD1B112A132EBFDFBE5E124FBC8" bold="true" box="[199,334,1076,1101]" pageId="30" pageNumber="481">Etymology.</emphasis>
The name refers to the 
<taxonomicName id="3BEA4D40B112A132E962FBE5E2DEFBC8" box="[600,692,1076,1101]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="483" phylum="Chordata" rank="species" species="pongola" status="sp. nov.">Pongola</taxonomicName>
River, along part of which this species occurs. It is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="B4F06548B112A132EBFDFBADE3F2FA85" pageId="30" pageNumber="481" type="diagnosis">
<paragraph id="FC5536C3B112A132EBFDFBADE3F2FA85" blockId="30.[151,1437,895,2037]" pageId="30" pageNumber="481">
<emphasis id="CE9EEAD1B112A132EBFDFBADE12BFB10" bold="true" box="[199,321,1148,1173]" pageId="30" pageNumber="481">Diagnosis.</emphasis>
A small 
<taxonomicName id="3BEA4D40B112A132EAFAFBADE258FB10" box="[448,562,1148,1173]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="481" phylum="Chordata" rank="genus">
<emphasis id="CE9EEAD1B112A132EAFAFBADE258FB10" box="[448,562,1148,1173]" italics="true" pageId="30" pageNumber="481">Afroedura</emphasis>
</taxonomicName>
(to 
<quantity id="3B129B26B112A132E94BFBADE2A2FB11" box="[625,712,1148,1172]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="30" pageNumber="481" unit="mm" value="40.0">40 mm</quantity>
SVL) differing from all other congeners by the following combination of characters: two pairs of enlarged subdigital lamellae per digit; tail not obviously verticillate, with 4–5 subcaudal rows and 6–7 supracaudal rows per verticil; dorsal scales smooth, 95–103 scale rows at midbody; a single (rarely two) internasal scale; 21–23 precloacal pores in males.
</paragraph>
</subSubSection>
<subSubSection id="B4F06548B112A133EBFDFADDE2A5FE98" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481" type="description">
<paragraph id="FC5536C3B112A132EBFDFADDE4A8FA5D" blockId="30.[151,1437,895,2037]" pageId="30" pageNumber="481">
<emphasis id="CE9EEAD1B112A132EBFDFADDE133FAA0" bold="true" box="[199,345,1292,1317]" pageId="30" pageNumber="481">Description.</emphasis>
(based on 
<typeStatus id="23518861B112A132EAEFFADDE25DFAA0" box="[469,567,1292,1317]" pageId="30" pageNumber="481" type="holotype">holotype</typeStatus>
TM 81210) Adult male; SVL 35.0 mm; TailL 32.0 mm (partly regenerated); mass in life 1.0 g. Head oval, wider than neck. Rostral approximately 1.5 times wider than high. Nostril pierced between rostral, first upper labial and three nasal scales. Nasorostrals separated by a single granular scale. Scales on snout round and flattened, decreasing in size posteriorly, smallest on crown. Ten scales between nasals and anterior margin of eye; 21 scales between eye and ear. No enlarged supraciliary spines. Supralabials eight. Mental broadly wedge-shaped, wider than deep and in contact with two postmentals; infralabials eight.
</paragraph>
<paragraph id="FC5536C3B112A132EBFDFA35E246F950" blockId="30.[151,1437,895,2037]" pageId="30" pageNumber="481">Dorsals relatively uniform, smooth, rounded and flattened, becoming larger laterally. Scales at midbody 101. Ventral scales rounded and imbricate, largest mesially becoming smaller ventrolaterally. Limbs relatively short and feet small; digits with two pairs of enlarged scansors; three enlarged inferomedian scales under the fourth toe. Precloacal pores 21, arranged in a continuous shallow ‘V’-shape. Tail flattened at base, becoming more cylindrical distally. Distal half regenerated. Caudal scales in whorls (verticils) with 6–7 scales rows dorsally and 4–5 ventrally. Supracaudals almost square with a rounded posterior margin and subimbricate. Subcaudals large and imbricate. 1–2 postcloacal spurs at base of tail.</paragraph>
<paragraph id="FC5536C3B112A132EBFDF90EE49FF80D" blockId="30.[151,1437,895,2037]" pageId="30" pageNumber="481">
<emphasis id="CE9EEAD1B112A132EBFDF90EE17AF97D" bold="true" box="[199,272,1759,1784]" pageId="30" pageNumber="481">Color.</emphasis>
Pale to mid-brown with 6–8 dark brown, irregular, wavy crossbands extending from occiput to sacrum. A median white spot or spots adjacent to each crossbar. Crown of head pale with brown to dark brown patches. A dark brown stripe extends from the nostrils through the middle of the eye to link with the occipital bar. In life the eyeballs appear dark gray to blackish through the skin. Limbs spotted and mottled with dark brown. Original tail with nine dark brown crossbands. Venter pinkish white with slightly darker pigmentation under tail.
</paragraph>
<paragraph id="FC5536C3B112A133EBFDF845E2A5FE98" blockId="30.[151,1437,895,2037]" lastBlockId="31.[151,1436,151,680]" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481">
<emphasis id="CE9EEAD1B112A132EBFDF845E151F828" bold="true" box="[199,315,1940,1965]" pageId="30" pageNumber="481">Variation.</emphasis>
<typeStatus id="23518861B112A132EA73F844E1DCF828" box="[329,438,1941,1965]" pageId="30" pageNumber="481" type="paratype">Paratypes</typeStatus>
and other specimens agree with the 
<typeStatus id="23518861B112A132E853F845E3A1F828" box="[873,971,1940,1965]" pageId="30" pageNumber="481" type="holotype">holotype</typeStatus>
in most features of scalation (
<tableCitation id="B1680378B112A132EE09F845E5E5F829" box="[1331,1423,1940,1965]" captionStart="TABLE 4" captionStartId="27.[125,205,605,626]" captionText="TABLE 4. variatiοn in key mοrphοlοgical characters in Limpοpο and Mpumalanga Prοvinces Afroedura including, fοr cοmparative purpοses, values fοr A. pondolia and A. major. Fοr size" pageId="30" pageNumber="481">Table 4</tableCitation>
). Nasorostrals separated by two granules in TM 81209. Scales between nasals and eye 10–13, from eye to ear 20–23. Supraciliary spines 2–3 and mental nearly as deep as wide in some 
<typeStatus id="23518861B112A132E8B1F80CE390F870" box="[907,1018,2013,2037]" pageId="30" pageNumber="481" type="paratype">paratypes</typeStatus>
. Postmentals 2–3. Supralabials 8–10, infralabials 8–9. Midbody scale rows 95–103. Enlarged inferomedian scales under fourth toe mostly 0, rarely 3–5. Precloacal pores in male 
<typeStatus id="23518861B113A133EA8BFF6CE276FF50" box="[433,540,189,213]" pageId="31" pageNumber="482" type="paratype">paratypes</typeStatus>
21–23, females lacking pores. Postcloacal spurs 1–2, rarely three. Original tails 50.5–52. 8% of total length. Tail with 6–7 dorsal and 4—5 ventral scale rows per indistinct verticil. 66.7% (8/12) of individuals have regenerated tails.
</paragraph>
</subSubSection>
<subSubSection id="B4F06548B113A133EBFDFEF6E226FE28" pageId="31" pageNumber="482" type="distribution">
<paragraph id="FC5536C3B113A133EBFDFEF6E507FEC5" blockId="31.[151,1436,151,680]" box="[199,1389,295,320]" pageId="31" pageNumber="482">
<emphasis id="CE9EEAD1B113A133EBFDFEF6E10AFEC5" bold="true" box="[199,352,295,320]" pageId="31" pageNumber="482">Distribution.</emphasis>
Known only from north of the 
<taxonomicName id="3BEA4D40B113A133E9FAFEF6E376FEC5" box="[704,796,295,320]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="483" phylum="Chordata" rank="species" species="pongola" status="sp. nov.">Pongola</taxonomicName>
River in the far northeast of Kwazulu-Natal (
<figureCitation id="64D12A46B113A133EE21FEF6E535FEC5" box="[1307,1375,295,320]" captionStart="FIGURE 6" captionStartId="22.[151,250,1208,1230]" captionTargetBox="[182,1404,196,1182]" captionTargetId="figure@22.[178,1409,193,1187]" captionTargetPageId="22" captionText="FIGURE 6. Map of Limpopo and Mpumalanga provinces, South Africa and adjacent regions showing the distributions of previously described species of Afroedura endemic to this region and known localities of the nine new species described in this paper. For previously recognized forms an approximate outline of the range based on Bates et al. (2014) is shown; genetically sampled localities are marked by circles in Figure 4. For new species all known localities are plotted as dots (see relevant species accounts), with genetically sampled site marked by circles with a a red center, type locality marked by a star, and all other localities indicated by open circles colored to correspond to the key in the figure. The very restricted distributions of A. maripi sp. nov., A. rondavelica sp. nov., and A. rupestris sp. nov. are indicated by arrows and are labeled directly on the map." httpUri="https://zenodo.org/record/250501/files/figure.png" pageId="31" pageNumber="482">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="FC5536C3B113A133EBFDFE9DE226FE28" blockId="31.[151,1436,151,680]" pageId="31" pageNumber="482">
<emphasis id="CE9EEAD1B113A133EBFDFE9DE1E0FEE0" bold="true" box="[199,394,332,357]" pageId="31" pageNumber="482">Natural history.</emphasis>
This gecko retreats into narrow crevices between rocks and under exfoliating flakes of granite. It is found in Granite Lowveld (SVl 3) and Northern Zululand Sourveld (SVl 22) (
<bibRefCitation id="987B4B32B113A133EF99FEBEE0BBFE28" author="Mucina" pageId="31" pageNumber="482" refString="Mucina, L. &amp; Rutherford, M. C. (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina &amp; Rutherford 2006</bibRefCitation>
) at elevations of 
<quantity id="3B129B26B113A133EAABFE45E27AFE29" box="[401,528,404,429]" metricMagnitude="2" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="31" pageNumber="482" unit="m" value="800.0" valueMax="850.0" valueMin="750.0">750–850 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="B4F06548B113A133EBFDFE66E08DFD2D" pageId="31" pageNumber="482" type="discussion">
<paragraph id="FC5536C3B113A133EBFDFE66E08DFD2D" blockId="31.[151,1436,151,680]" pageId="31" pageNumber="482">
<emphasis id="CE9EEAD1B113A133EBFDFE66E151FE55" bold="true" box="[199,315,439,464]" pageId="31" pageNumber="482">Remarks.</emphasis>
<taxonomicName id="3BEA4D40B113A133EA7CFE66E24AFE55" box="[326,544,439,464]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="pongola" status="sp. nov.">
<emphasis id="CE9EEAD1B113A133EA7CFE66E24AFE55" box="[326,544,439,464]" italics="true" pageId="31" pageNumber="482">Afroedura pongola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5AD57AAB113A133E911FE66E2E3FE55" box="[555,649,439,464]" pageId="31" pageNumber="482" rank="species">
<emphasis id="CE9EEAD1B113A133E911FE66E2E3FE55" bold="true" box="[555,649,439,464]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
was included by 
<bibRefCitation id="987B4B32B113A133E865FE66E47EFE55" author="Jacobsen" box="[863,1044,439,464]" pageId="31" pageNumber="482" refString="Jacobsen, N. H. G. (1990) A Herpetological Survey of the Transvaal. Unpublished Ph. D. Dissertation, University of Natal, Durban, South Africa, 1621 pp." type="book" year="1990">Jacobsen (1990</bibRefCitation>
, 
<bibRefCitation id="987B4B32B113A133EF1EFE69E401FE55" author="Jacobsen" box="[1060,1131,440,464]" pageId="31" pageNumber="482" refString="Jacobsen, N. H. G. (1992 a) Flat geckos (genus Afroedura) in the Transvaal. Journal of the Herpetological Association of Africa, 40, 22 - 25. http: // dx. doi. org / 10.1080 / 04416651.1992.9650313" type="journal article" year="1992" yearSuffix="a">1992a</bibRefCitation>
, 
<bibRefCitation id="987B4B32B113A133EF40FE69E4DEFE55" author="Jacobsen" box="[1146,1204,440,464]" pageId="31" pageNumber="482" refString="Jacobsen, N. H. G. (1997) Family Gekkonidae, Genus Afroedura. In: Van Wyk, J. H. (Ed.), Proceedings of the FitzSimons Commemorative Symposium, South African Lizards: 50 years of Progress and Third H. A. A. Symposium on African Herpetology, Transvaal Museum, Pretoria, South Africa, 11 - 15 October 1993. Herpetological Association of Africa, Stellenbosch, pp. 34 - 39." type="journal article" year="1997">1997</bibRefCitation>
) in his 
<taxonomicName id="3BEA4D40B113A133EE2FFE69E5F6FE55" box="[1301,1436,439,464]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="pondolia">
<emphasis id="CE9EEAD1B113A133EE2FFE69E5F6FE55" box="[1301,1436,439,464]" italics="true" pageId="31" pageNumber="482">A. pondolia</emphasis>
</taxonomicName>
complex, but our molecular results demonstrate that whereas it forms a clade with other members of this proposed complex, 
<taxonomicName id="3BEA4D40B113A133EA3FFDD1E1E2FD9D" box="[261,392,511,536]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="pondolia">
<emphasis id="CE9EEAD1B113A133EA3FFDD1E1E2FD9D" box="[261,392,511,536]" italics="true" pageId="31" pageNumber="482">A. pondolia</emphasis>
</taxonomicName>
is only distantly related. 
<taxonomicName id="3BEA4D40B113A133E998FE2EE31DFD9D" box="[674,887,511,536]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="pongola" status="sp. nov.">
<emphasis id="CE9EEAD1B113A133E998FE2EE31DFD9D" box="[674,887,511,536]" italics="true" pageId="31" pageNumber="482">Afroedura pongola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5AD57AAB113A133E847FE2EE3BDFD9D" box="[893,983,511,536]" pageId="31" pageNumber="482" rank="species">
<emphasis id="CE9EEAD1B113A133E847FE2EE3BDFD9D" bold="true" box="[893,983,511,536]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
may be distinguished from the similarly sized 
<taxonomicName id="3BEA4D40B113A133EBEDFDF5E120FDB8" box="[215,330,548,573]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="marleyi">
<emphasis id="CE9EEAD1B113A133EBEDFDF5E120FDB8" box="[215,330,548,573]" italics="true" pageId="31" pageNumber="482">A. marleyi</emphasis>
</taxonomicName>
on the basis of its greater number of midbody scale rows (p &lt;0.001, t20 d.f.). and from this species and the much larger 
<taxonomicName id="3BEA4D40B113A133EABAFD99E180FDE5" box="[384,490,584,608]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="maripi" status="sp. nov.">
<emphasis id="CE9EEAD1B113A133EABAFD99E180FDE5" box="[384,490,584,608]" italics="true" pageId="31" pageNumber="482">A. maripi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5AD57AAB113A133EAC8FD96E226FDE5" box="[498,588,583,608]" pageId="31" pageNumber="482" rank="species">
<emphasis id="CE9EEAD1B113A133EAC8FD96E226FDE5" bold="true" box="[498,588,583,608]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
and 
<taxonomicName id="3BEA4D40B113A133E9BEFD99E343FDE5" box="[644,809,583,608]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="rondavelica" status="sp. nov.">
<emphasis id="CE9EEAD1B113A133E9BEFD99E343FDE5" box="[644,809,583,608]" italics="true" pageId="31" pageNumber="482">A. rondavelica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5AD57AAB113A133E80BFD96E3E1FDE5" box="[817,907,583,608]" pageId="31" pageNumber="482" rank="species">
<emphasis id="CE9EEAD1B113A133E80BFD96E3E1FDE5" bold="true" box="[817,907,583,608]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
(maximum SVL 
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and 
<quantity id="3B129B26B113A133EFE3FD96E543FDDA" box="[1241,1321,583,608]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="31" pageNumber="482" unit="mm" value="55.0">55 mm</quantity>
<emphasis id="CE9EEAD1B113A133EE0BFD98E512FDE5" box="[1329,1400,585,608]" italics="true" pageId="31" pageNumber="482">versus</emphasis>
<quantity id="3B129B26B113A133EEBAFD99E0A9FD01" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="31" pageNumber="482" unit="mm" value="40.0">40 mm</quantity>
for 
<taxonomicName id="3BEA4D40B113A133EBC2FDBDE11DFD00" box="[248,375,620,645]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="482" phylum="Chordata" rank="species" species="pongola" status="sp. nov.">
<emphasis id="CE9EEAD1B113A133EBC2FDBDE163FD06" box="[248,265,620,643]" italics="true" pageId="31" pageNumber="482">A</emphasis>
. 
<emphasis id="CE9EEAD1B113A133EA21FDBDE11DFD00" box="[283,375,620,645]" italics="true" pageId="31" pageNumber="482">pongola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5AD57AAB113A133EABBFDBDE1B4FD00" box="[385,478,620,645]" pageId="31" pageNumber="482" rank="species">
<emphasis id="CE9EEAD1B113A133EABBFDBDE1B4FD00" bold="true" box="[385,478,620,645]" pageId="31" pageNumber="482">sp. nov.</emphasis>
</taxonomicNameLabel>
) by its greater number of precloacal pores in males (mean of 22.4 
<emphasis id="CE9EEAD1B113A133EFCDFDBCE554FD01" box="[1271,1342,621,644]" italics="true" pageId="31" pageNumber="482">versus</emphasis>
13.4 or fewer).
</paragraph>
</subSubSection>
</treatment>
</document>