148 lines
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148 lines
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.803.28903" ID-GBIF-Dataset="6c6c0b59-636f-4502-8b48-642ba231eb16" ID-PMC="PMC6390078" ID-Pensoft-Pub="1313-2970-803-7" ID-PubMed="30814899" ID-ZBK="936CFD88F297440EA9BE4C258AE9BD09" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-803-7" ModsDocOrigin="ZooKeys 803" ModsDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1545315537716" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2018" docId="05F7A20804B9F09FD9808916361DBA77" docLanguage="en" docName="ZooKeys 803: 7-70" docOrigin="ZooKeys 803" docSource="http://dx.doi.org/10.3897/zookeys.803.28903" docTitle="Exocelina cyclops Shaverdo & Balke, sp. n." docType="treatment" docUuid="236B93F4-6599-499D-8325-5FB446714AF2" docUuidSource="ZooBank" docVersion="7" lastPageNumber="16" masterDocId="5133FFF6125A1065FF83FF96FFF3FFD4" masterDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="70" masterPageNumber="7" pageNumber="15" updateTime="1668166558239" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>803</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>7</mods:start>
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<mods:end>70</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.803.28903</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.803.28903</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-803-7</mods:identifier>
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<mods:identifier type="ZBK">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
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<mods:identifier type="ZooBank">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154125614" LSID="urn:lsid:zoobank.org:act:236B93F4-6599-499D-8325-5FB446714AF2" httpUri="http://treatment.plazi.org/id/05F7A20804B9F09FD9808916361DBA77" lastPageId="9" lastPageNumber="16" pageId="8" pageNumber="15">
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<subSubSection pageId="8" pageNumber="15" type="nomenclature">
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<paragraph pageId="8" pageNumber="15">
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5.
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<taxonomicName LSID="http://zoobank.org/236B93F4-6599-499D-8325-5FB446714AF2" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina cyclops" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="species" species="cyclops">Exocelina cyclops Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="8" pageNumber="15">sp. n.</taxonomicNameLabel>
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Figs 12, 36
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="reference_group">
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<paragraph pageId="8" pageNumber="15">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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undescribed sp. MB3330:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/ncomms5001" year="2014">Toussaint et al. 2014</bibRefCitation>
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: supplementary figs 1-4, table 2;
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary figs S1-S2, table S3.
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</paragraph>
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<paragraph pageId="8" pageNumber="15">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina cyclops" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="species" species="cyclops">Exocelina cyclops</taxonomicName>
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New Guinea MB3330:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary information S5-S6.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="type locality">
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<paragraph pageId="8" pageNumber="15">Type locality.</paragraph>
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<paragraph pageId="8" pageNumber="15">
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Papua: Jayapura Regency, Cyclops Mts,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-2.5338333">02°32.03'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="140.50684">140°30.41'E</geoCoordinate>
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, 710 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="type material">
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<paragraph pageId="8" pageNumber="15">Type material.</paragraph>
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<paragraph pageId="8" pageNumber="15">
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Holotype: male "Indonesia: Papua, Cyclops Mts., 710 m,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-2.53385">02°32.031'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="140.50687">140°30.412'E</geoCoordinate>
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, local collector ca. 1997" (ZSM). Paratypes: 1 male with the same label as the holotype (NHMW). 1 male, 2 females "Indonesia: Papua, Cyclops Mts., 1120 m,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-2.5252666">02°31.516'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="140.50726">140°30.436'E</geoCoordinate>
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, local collector ca. 1997" (MZB, ZSM). 2 males, 1 female "Indonesia: Papua, Cyclops Mts., 615 m,
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<geoCoordinate direction="south" orientation="latitude" precision="1" value="-2.53385">02°32.031'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="140.50687">140°30.412'E</geoCoordinate>
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, local collector ca. 1997" (NHMW, ZSM). 2 males "Indonesia: Papua, Cyclops Mts., Doyo, 365 m, local collector ca. 1997" (ZSM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="description">
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<paragraph pageId="8" pageNumber="15">Description.</paragraph>
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<paragraph pageId="8" pageNumber="15">Body size and form: Beetle small: TL-H 3.0-3.25 mm, TL 3.25-3.55 mm, MW 1.65-1.8 mm (holotype: TL-H 3.1 mm, TL 3.4 mm, MW 1.7 mm), with oblong-oval habitus.</paragraph>
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<paragraph pageId="8" pageNumber="15">Coloration: Reddish. Dorsal surface almost uniformly yellowish red to reddish brown, with paler anterior part of head and pronotum laterally; head appendages and legs yellowish red (Fig. 12). All type specimens are teneral, therefore, coloration may be darker.</paragraph>
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<paragraph pageId="8" pageNumber="15">
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Surface sculpture: Submatt dorsally. As in
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<taxonomicName lsidName="E. pseudopusilla" pageId="8" pageNumber="15" rank="species" species="pseudopusilla">E. pseudopusilla</taxonomicName>
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sp. n. but microreticulation more weakly impressed, dorsal surface shinier.
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</paragraph>
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<paragraph pageId="8" pageNumber="15">Structures: Pronotum with lateral bead. Its lateral sides with distinct longitudinal impressions. Base of prosternum and neck of prosternal process with distinct ridge, rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 slightly truncate.</paragraph>
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<paragraph pageId="8" pageNumber="15">
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Male: Antennae simple (Fig. 12). Protarsomere 4 with medium-sized, thick, curved anterolateral hook-like seta. Protarsomere 5 long and narrow, with anterior band of 30 and posterior row of 7 relatively long setae (Fig. 36D). Median lobe in lateral view slightly curved, with slightly thickened, straight apex; in ventral view, subparallel, very slightly narrowed distally, with broad, truncate apex. Paramere slightly concave on dorsal side, with long, dense subdistal setae, median and proximal ones finer and sparser (Fig. 36
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Abdominal ventrite 6 with 4-6 lateral striae on each side.
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</paragraph>
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<paragraph pageId="8" pageNumber="15">Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="affinities">
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<paragraph pageId="8" pageNumber="15">Affinities.</paragraph>
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<paragraph pageId="8" pageNumber="15">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina cyclops" order="Coleoptera" pageId="8" pageNumber="15" phylum="Arthropoda" rank="species" species="cyclops">Exocelina cyclops</taxonomicName>
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sp. n. is similar to
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<taxonomicName lsidName="E. sumokedi" pageId="8" pageNumber="15" rank="species" species="sumokedi">E. sumokedi</taxonomicName>
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sp. n.,
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<taxonomicName lsidName="E. adelbertensis" pageId="8" pageNumber="15" rank="species" species="adelbertensis">E. adelbertensis</taxonomicName>
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sp. n., and
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<taxonomicName lsidName="E. bewani" pageId="8" pageNumber="15" rank="species" species="bewani">E. bewani</taxonomicName>
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sp. n., but has coarser dorsal punctation. In this character, the species is more similar to
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<taxonomicName lsidName="E. pusilla" pageId="8" pageNumber="15" rank="species" species="pusilla">E. pusilla</taxonomicName>
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sp. n. and
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<taxonomicName lsidName="E. pseudopusilla" pageId="8" pageNumber="15" rank="species" species="pseudopusilla">E. pseudopusilla</taxonomicName>
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sp. n. but it differs from them in much broader and differently shaped median lobe (not distinctly narrowed distally, with apex thicker in lateral and ventral views) and from the latter, also in smaller size and slightly shinier dorsal surface.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="15" type="distribution">
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<paragraph pageId="8" pageNumber="15">Distribution.</paragraph>
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<paragraph pageId="8" pageNumber="15">Papua: Jayapura Regency. The species is known only from the Cyclops Mountains (Fig. 50).</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="16" type="etymology">
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<paragraph pageId="9" pageNumber="16">
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<pageBreakToken pageId="9" pageNumber="16" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="9" pageNumber="16">The species is named after the Cyclops Mountains. The name is a noun in the nominative singular standing in apposition.</paragraph>
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</subSubSection>
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</treatment>
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</document>
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