<documentid="44BB15B12D46E297BCF1B2BA7AC4CAEC"ID-DOI="10.11646/zootaxa.5492.4.8"ID-ISSN="1175-5326"ID-Zenodo-Dep="13269124"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="guilherme"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="felipe"IM.treatments_approvedBy="felipe"checkinTime="1723108120852"checkinUser="plazi"docAuthor="Asima, A., Sankaran, Pradeep M. & Prasad, Gopal"docDate="2024"docId="03914A50FFA5FFD146E8F9D980C630A9"docLanguage="en"docName="zootaxa.5492.4.8.pdf"docOrigin="Zootaxa 5492 (4)"docSource="http://dx.doi.org/10.11646/zootaxa.5492.4.8"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Asceua tertia Asima, Sankaran & Prasad, 2024, sp. nov."docType="treatment"docVersion="3"lastPageNumber="599"masterDocId="FFA83228FFA5FFD2467FFF968205344E"masterDocTitle="A new species of Asceua Thorell, 1887 (Araneae, Zodariidae) from the Western Ghats of India"masterLastPageNumber="599"masterPageNumber="596"pageNumber="596"updateTime="1723551340250"updateUser="guilherme"zenodo-license-document="CLOSED">
<mods:titleid="D9026D06924863042DE16CAE6DC3CA86">A new species of Asceua Thorell, 1887 (Araneae, Zodariidae) from the Western Ghats of India</mods:title>
<mods:affiliationid="1C59D1516CAF4254ECDF9BE4E6DC449A">Department of Zoology, University of Kerala, Kariavattom, Thiruvananthapuram, Kerala 695 581, India.</mods:affiliation>
<mods:affiliationid="4C9E9474C47A5220718660DB90F6E526">Division of Arachnology, Department of Zoology, Sacred Heart College, Thevara, Cochin, Kerala 682 013, India.</mods:affiliation>
<mods:affiliationid="7ADE815039DACA2900FB8B121B4CE335">Department of Zoology, University of Kerala, Kariavattom, Thiruvananthapuram, Kerala 695 581, India.</mods:affiliation>
<taxonomicNameid="4C3880C5FFA5FFD246E8F9D983323227"authorityName="Asima & Sankaran & Prasad"authorityYear="2024"box="[151,311,1615,1641]"class="Arachnida"family="Zodariidae"genus="Asceua"kingdom="Animalia"order="Araneae"pageId="0"pageNumber="596"phylum="Arthropoda"rank="species"species="tertia"status="sp. nov.">
<collectionCodeid="ED296383FFA5FFD244D8F97780BC32B7"box="[679,697,1761,1785]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="0"pageNumber="596">R</collectionCode>
<collectionCodeid="ED296383FFA5FFD2477EF89383113353"box="[257,276,1797,1821]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="0"pageNumber="596">R</collectionCode>
<collectingDateid="EFC2246EFFA5FFD2442AF8BF80FE330F"box="[597,763,1833,1857]"pageId="0"pageNumber="596"value="2021-03-12">12 March 2021</collectingDate>
<materialsCitationid="3B50F11BFFA5FFD2478FF8DB868233C7"collectingDate="2013-12-04"collectionCode="WGRC, R"collectorName="M. S. Pradeep"country="India"county="Ernakulam"elevation="40"latitude="10.183"location="Kurisumudi"longLatPrecision="549"longitude="76.3"municipality="Malayatoor"pageId="0"pageNumber="596"specimenCount="1"specimenCount-female="1"stateProvince="Kerala">
<collectionCodeid="ED296383FFA5FFD24509F8DB818D332B"box="[886,904,1869,1893]"country="Chile"name="Departamento de Geologia, Universidad de Chile"pageId="0"pageNumber="596">R</collectionCode>
<collectingDateid="EFC2246EFFA5FFD24787F8E780C033C7"box="[504,709,1905,1929]"pageId="0"pageNumber="596"value="2013-12-04">04 December 2013</collectingDate>
<taxonomicNameid="4C3880C5FFA5FFD24742F80383C133E3"authorityName="Asima & Sankaran & Prasad"authorityYear="2024"box="[317,452,1941,1965]"class="Arachnida"family="Zodariidae"genus="Asceua"kingdom="Animalia"order="Araneae"pageId="0"pageNumber="596"phylum="Arthropoda"rank="species"species="tertia"status="sp. nov.">
as both share laterally compressed cymbium, cymbium with prominent posteroretrolateral cymbial process, horizontally oriented, hyaline conductor, branched median apophysis, W-shaped posterior epigynal margin, epigyne with anterior hood, and copulatory ducts with wide proximal and narrow distal parts, but can be separated by the following combination of features: males by tibia with prominent PTA (
<figureCitationid="1303E7C3FFA4FFD344A4FE9581833552"box="[731,902,259,284]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="1"pageNumber="597">Fig. 2A–C, E–G</figureCitation>
,
<bibRefCitationid="EFA986B7FFA4FFD345EDFE95864E3552"author="Zhang, B. S. & Zhang, F. & Jia, X. M."box="[914,1099,259,284]"pageId="1"pageNumber="597"pagination="62 - 68"refId="ref2880"refString="Zhang, B. S., Zhang, F. & Jia, X. M. (2012) Two new species of the ant spider genus Asceua Thorell, 1887 (Araneae: Zodariidae) from China. Zootaxa, 3307 (1), 62 - 68. https: // doi. org / 10.11646 / zootaxa. 3307.1.4"type="journal article"year="2012">
<bibRefCitationid="EFA986B7FFA4FFD34289FE9582C8350E"author="Zhang, B. S. & Zhang, F."pageId="1"pageNumber="597"pagination="285 - 300"refId="ref2832"refString="Zhang, B. S. & Zhang, F. (2018) Two new species of the spider family Zodariidae from Laos. Zootaxa, 4459 (2), 285 - 300. https: // doi. org / 10.11646 / zootaxa. 4459.2.4"type="journal article"year="2018">Zhang & Zhang 2018</bibRefCitation>
: fig. 8A–B, D–E). Males of
<taxonomicNameid="4C3880C5FFA4FFD3478CFEBE80423571"authorityName="Asima & Sankaran & Prasad"authorityYear="2024"box="[499,583,296,319]"class="Arachnida"family="Zodariidae"genus="Asceua"kingdom="Animalia"order="Araneae"pageId="1"pageNumber="597"phylum="Arthropoda"rank="species"species="tertia"status="sp. nov.">
. absent in both species); females can be separated from the females of both species by epigyne with anterior hood and median epigynal depression with median bulging, which are absent in the latter two species (cf.
<figureCitationid="1303E7C3FFA4FFD34355FE2182C135BD"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="1"pageNumber="597">Fig. 2A–B, D–F</figureCitation>
and
<bibRefCitationid="EFA986B7FFA4FFD3468AFE4D838A35BA"author="Sankaran, P. M."box="[245,399,475,500]"pageId="1"pageNumber="597"pagination="381 - 405"refId="ref2726"refString="Sankaran, P. M. (2023) Taxonomic notes on the ant-eating spider genera Asceua Thorell, 1887 and Cydrela Thorell, 1873 from India, with comment on Indian species of Euryeidon Dankittipakul & Jocque, 2004 (Araneae: Zodariidae). Zootaxa, 5296 (3), 381 - 405. https: // doi. org / 10.11646 / zootaxa. 5296.3.4"type="journal article"year="2023">Sankaran 2023</bibRefCitation>
<taxonomicNameid="4C3880C5FFA4FFD34762F8FD83A033CC"authorityName="Asima & Sankaran & Prasad"authorityYear="2024"box="[285,421,1899,1922]"class="Arachnida"family="Zodariidae"genus="Asceua"kingdom="Animalia"order="Araneae"pageId="1"pageNumber="597"phylum="Arthropoda"rank="species"species="tertia"status="sp. nov.">
, holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal. I–J left palp: I prolateral; J ventral. K–L genitalia of paratypes: K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm.
<taxonomicNameid="4C3880C5FFA7FFD04767FAA583993105"authorityName="Asima & Sankaran & Prasad"authorityYear="2024"box="[280,412,1331,1355]"class="Arachnida"family="Zodariidae"genus="Asceua"kingdom="Animalia"order="Araneae"pageId="2"pageNumber="598"phylum="Arthropoda"rank="species"species="tertia"status="sp. nov.">
A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm.
<figureCitationid="1303E7C3FFA7FFD0440AF9E180FA32C1"box="[629,767,1655,1679]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="2"pageNumber="598">Fig. 1C, E, G</figureCitation>
). Body length 2.54. Carapace 1.24 long, 0.10 wide. Opisthosoma 1.30 long, 0.10 wide. Prosoma overall yellow-brown. Leg and palp segments creamy white to yellow-brown, with broad brown patches and narrow longitudinal stripes on femora, patellae and tibiae of all legs; dorsum of opisthosoma with three pairs of obliquely oriented narrow, creamy white stripes and a white dot near the rear end (
<figureCitationid="1303E7C3FFA7FFD04200F97586D632B5"box="[1151,1235,1763,1787]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="2"pageNumber="598">Fig. 1C</figureCitation>
), sides posteriorly with a pair of short, narrow, creamy white stripes and numerous tiny creamy white spots (
<figureCitationid="1303E7C3FFA7FFD04229F89186A23351"box="[1110,1191,1799,1823]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="2"pageNumber="598">Fig. 1E</figureCitation>
), venter creamy white, with broad brown patch posteriorly and with a pair of pale black patches anteriorly; dorsal scutum red-brown (
<figureCitationid="1303E7C3FFA7FFD0430EF8BD82BC3329"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="2"pageNumber="598">Fig. 1C</figureCitation>
). Fovea short, longitudinal (
<figureCitationid="1303E7C3FFA7FFD04788F8D9804A3329"box="[503,591,1871,1895]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="2"pageNumber="598">Fig. 1C</figureCitation>
). Chilum inverted triangular, unipartite. Cheliceral promargin with two tiny teeth, retromargin without tooth visible. Sternum shield-shaped, with coxal and intercoxal extensions, former one well developed. Opisthosoma elongate-oval (
<figureCitationid="1303E7C3FFA7FFD04433F80180A433E1"box="[588,673,1943,1967]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="2"pageNumber="598">Fig. 1C</figureCitation>
); dorsum anteromedially with narrow membranous scutum (
<figureCitationid="1303E7C3FFA7FFD04346F801878A33E1"box="[1337,1423,1943,1967]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="2"pageNumber="598">Fig. 1C</figureCitation>
). Metatarsi II–III with distal preening brush. Eye diameters and interdistances: ALE 0.13, AME 0.12, PLE 0.11, PME 0.14; AME–ALE 0.04, AME–AME 0.03, AME–PME 0.12, ALE–PLE 0.03, PME–PLE 0.08, PME–PME 0.07. Clypeus height at AMEs 0.48, at ALEs 0.51. Chelicerae 0.52 long. Sternum 0.69 long, 0.75 wide. Measurements of palp and legs: palp 2.10 [0.68, 0.22, 0.20, 1.00], I 4.94 [1.24, 0.35, 1.21, 1.47, 0.67], II 4.22 [1.12, 0.38, 0.92, 1.24, 0.56], III 4.16 [1.15, 0.40, 0.86, 1.28, 0.47], IV 5.08 [1.30, 0.38, 1.10, 1.68, 0.62]. Leg formula: 4123. Spination of palp: femur pld 1 rld 1, patella spineless, tibia spineless, tarsus/ cymbium spineless; legs: femora I–IV do 2; patellae I–IV spineless; tibiae I–III spineless, IV rlv 1; metatarsus I spineless, II–IV plv 1 rlv 1; tarsi I–IV spineless.
<figureCitationid="1303E7C3FFA6FFD145C1FEB18626350E"box="[958,1059,295,320]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1I–J</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD14251FEBE8671350E"box="[1070,1140,296,320]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2A–D</figureCitation>
). Tibia with flat retrolateral, short, conical ventral, and broad prolateral apophyses (
<figureCitationid="1303E7C3FFA6FFD14496FEDD813D352A"box="[745,824,331,356]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1I</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD1453AFEDA818C352A"box="[837,905,332,356]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2A–C</figureCitation>
); RTA unbranched, with smoothly rounded distal part, directed at 12-o’ clock ventrally (
<figureCitationid="1303E7C3FFA6FFD1443EFEF9809635C6"box="[577,659,367,392]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1J</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD144DDFEE680E135C6"box="[674,740,368,392]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2B–C</figureCitation>
); PTA distally bifurcated, with narrow, claw-like prolateral and broad, conical retrolateral branches (
<figureCitationid="1303E7C3FFA6FFD1445CFE05807735E2"box="[547,626,403,428]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1I</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD14400FE0280C735E2"box="[639,706,404,428]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2A, C</figureCitation>
; arrow 1). Cymbium modified, with prominent cymbial retrolateral fold, with highly modified posteroretrolateral extremity forming cymbial process, dorsally with cymbial scopula (
<figureCitationid="1303E7C3FFA6FFD14312FE2182C235BA"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1I–J</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD146ABFE4A831235BA"box="[212,279,476,500]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2A–C</figureCitation>
). Tegulum broad (
<figureCitationid="1303E7C3FFA6FFD147A2FE4D802835BA"box="[477,557,475,500]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1J</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD14444FE4A805835BA"box="[571,605,476,500]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2B</figureCitation>
). Median apophysis flat, with broad prolateral, and long, slender, retrolateral branches, lying adjacent to conductor (
<figureCitationid="1303E7C3FFA6FFD1444EFE6980923656"box="[561,663,511,536]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1I–J</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD144DEFD9680E13656"box="[673,740,512,536]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2A–B</figureCitation>
<figureCitationid="1303E7C3FFA6FFD147A4FDB5802C3672"box="[475,553,547,572]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1J</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD1444BFDB280723672"box="[564,631,548,572]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2B–C</figureCitation>
). Embolus long, filiform, basolateral to tegulum, with short embolar base, at first running anteriorly along the prolateral side of cymbium, then turning retrolaterally, without forming distal loop, with blunt, flat tip having tiny thorn-like process distodorsally and membranous extension distoventrally (
<figureCitationid="1303E7C3FFA6FFD142A0FDFD874236CA"box="[1247,1351,619,644]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1I–J</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD1432BFDFA879D36CA"box="[1364,1432,620,644]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2A–D</figureCitation>
<figureCitationid="1303E7C3FFA6FFD1447DFD2580893682"box="[514,652,691,716]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Fig. 1D, F, H</figureCitation>
). Body length 3.20. Carapace 1.40 long, 1.00 wide. Opisthosoma 1.80 long, 1.50 wide. Habitus and details like male except for the following: colour of prosoma slightly dark brown. Opisthosoma widely oval, without dorsal scutum (
<figureCitationid="1303E7C3FFA6FFD14461FD6D8076375A"box="[542,627,763,788]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Fig. 1D</figureCitation>
); dorsum with four pairs of creamy-white stripes (
<figureCitationid="1303E7C3FFA6FFD142F1FD6D86E6375A"box="[1166,1251,763,788]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Fig. 1D</figureCitation>
); sides anteriorly with small and posteriorly with broad creamy white patches (
<figureCitationid="1303E7C3FFA6FFD14560FC89816B3776"box="[799,878,799,824]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Fig. 1F</figureCitation>
); venter uniformly creamy white. Eye diameters and interdistances: ALE 0.10, AME 0.12, PLE 0.09, PME 0.12; AME–ALE 0.03, AME–AME 0.04, AME–PME 0.14, ALE– PLE 0.02, PME–PLE 0.14, PME–PME 0.12. Clypeus height at AMEs 0.43, at ALEs 0.41. Chelicerae 0.50 long. Sternum 0.90 long, 0.66 wide. Measurements of palp and legs: palp 1.57 [0.68, 0.16, 0.33, 0.40], I 4.43 [1.00, 0.37, 1.10, 1.19, 0.77], II 3.73 [1.00, 0.30, 0.92, 1.01, 0.50], III 4.13 [1.00, 0.38, 0.90, 1.19, 0.66], IV 4.89 [1.01, 0.40, 1.19, 1.60, 0.60]. Leg formula: 4132. Spination of palp: femur do 1, patella spineless, tibia pl 1, tarsus pl 1 plv 2 rlv 1; legs: tibia IV spineless.
<figureCitationid="1303E7C3FFA6FFD14775FC618365305E"box="[266,352,1015,1040]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1L</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD14713FC6E83AB305E"box="[364,430,1016,1040]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2E–G</figureCitation>
). Epigyne with W-shaped posterior margin, with median broad, membranous depression, with median bulging visible only in lateral view, with anterior hood (
<figureCitationid="1303E7C3FFA6FFD14524FB8D81C9307A"box="[859,972,1051,1076]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">Fig. 2E–F</figureCitation>
; arrow 6). Copulatory openings indistinct. Copulatory ducts long, with multiple windings, with wide proximal and narrow distal parts (
<figureCitationid="1303E7C3FFA6FFD14210FBA986C13016"box="[1135,1220,1087,1112]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1L</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD142AFFBD686F13016"box="[1232,1268,1088,1112]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2G</figureCitation>
). Spermathecae small, oval, lying adjacent to each other, situated posteriorly (
<figureCitationid="1303E7C3FFA6FFD14548FBF581883032"box="[823,909,1123,1148]"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1L</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD145E4FBF281BA3032"box="[923,959,1124,1148]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2G</figureCitation>
). Fertilization ducts narrow, diverging (
<figureCitationid="1303E7C3FFA6FFD14312FBF582B330EE"captionStart="FIGURE 1"captionStartId="1.[151,250,1899,1923]"captionTargetBox="[164,1423,524,1875]"captionTargetId="figure-239@1.[164,1423,524,1875]"captionTargetPageId="1"captionText="FIGURE 1. Asceua tertia sp. nov., holotype male (ZSI/WGRC/I.R.INV.26826; A, C, E, G, I–J) and paratype females (ZSI/ WGRC/1.R.INV.26828 & 26831; B, D, F, H, K–L). A–B live images. C–D habitus, dorsal. E–F same, retrolateral. G–H same, frontal.I–Jleft palp:I prolateral; J ventral.K–L genitalia of paratypes:K epigyne intact of female 2 (ZSI/WGRC/1.R.INV.26831); L vulva of female 1(ZSI/WGRC/1.R.INV.26828). Scale bars: C–J, 0.5 mm; K–L, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269126"httpUri="https://zenodo.org/record/13269126/files/figure.png"pageId="3"pageNumber="599">Figs 1L</figureCitation>
,
<figureCitationid="1303E7C3FFA6FFD146BDFB1E82E230EE"box="[194,231,1160,1184]"captionStart="FIGURE 2"captionStartId="2.[151,248,1331,1355]"captionTargetBox="[160,1427,191,1297]"captionTargetId="figure-17@2.[151,1436,181,1307]"captionTargetPageId="2"captionText="FIGURE 2. Asceua tertia sp. nov. A–D left palp (ZSI/WGRC/I.R.INV.26826): A prolateral; B ventral; C retrolateral; D enlarged view of the tip of embolus, retrolateral. E–G female genitalia (ZSI/WGRC/1.R.INV.26828): E epigyne, ventral; F same, retrolateral; G vulva, dorsal. Abbreviations: C, conductor; CD, copulatory duct; CS, cymbial scopula; CP, cymbial process; E, embolus; EB, embolar base; ED, epigynal depression; EH, epigynal hood; FD, fertilisation duct; MA, median apophysis; PTA, prolateral tibial apophysis; RTA, retrolateral tibial apophysis; S, spermatheca; SD, sperm duct; T, tegulum; VTA, ventral tibial apophysis. Arrows indicate retrolateral branch of PTA (1), branches of MA (2 & 3), distodorsal thorn-like process of embolus (4), ventral membranous extension of embolus (5), and median bulging of epigyne (6). Scale bars: A–G, 0.2 mm."figureDoi="http://doi.org/10.5281/zenodo.13269128"httpUri="https://zenodo.org/record/13269128/files/figure.png"pageId="3"pageNumber="599">2G</figureCitation>
