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227 lines
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<document id="F6799115FFD542546CA2A4E70F8AFA72" ID-DOI="10.5281/zenodo.3708236" ID-GBIF-Dataset="6d350404-6f84-4fdc-83ef-65a70583acf8" ID-ISSN="1942-1354" ID-Zenodo-Dep="3708236" ID-ZooBank="DBEAD659-A6B3-4579-9F64-D4D2C1301D10" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584033684974" checkinUser="felipe" docAuthor="Klimaszewski, Jan, Webster, Reginald P., Davies, Anthony & Bourdon, Caroline" docDate="2018" docId="03823B711902FF8EFF7686A052C217F2" docLanguage="en" docName="InsectaMundi.0648.1-12.pdf.imf" docOrigin="Insecta Mundi 648" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Atheta (Hydrosmectomorpha) Klimaszewski, new subgenus" docType="treatment" docVersion="5" lastPageNumber="3" masterDocId="FFBB43091901FF8AFFB6825F56211416" masterDocTitle="Description of Hydrosmectomorpha Klimaszewski and Webster, a new subgenus of Atheta C. G. Thomson, with three new Canadian species (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="2" updateTime="1698727433203" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="D4628ADAF8D9862084B25975F1A6E188" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="C372AF64ABEED97873E12FAD169E73CB">
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<mods:title id="04E8A0136CDB3C7CEF42D2749EA35AD9">Description of Hydrosmectomorpha Klimaszewski and Webster, a new subgenus of Atheta C. G. Thomson, with three new Canadian species (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="AE3E38763E654A7AEFEF8B9886811097" type="personal">
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<mods:roleTerm id="8034C37DA985CCC3325C115E84FBEB6D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4FAAF468E5A6F61485E90B94760CBC1D">Klimaszewski, Jan</mods:namePart>
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</mods:name>
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<mods:name id="DC6DC862F7C2C2D7E7326483096E471D" type="personal">
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<mods:roleTerm id="EC5000F981F88211FCEFDC90617EA8D4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8D4DB9FD4CCD0051F9981AB0B6C4A3B1">Webster, Reginald P.</mods:namePart>
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</mods:name>
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<mods:name id="858EB88F7E42C96DA923DEE483A10D90" type="personal">
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<mods:roleTerm id="0E996AEA346EB0491ECB9A324C6F93C9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D1AB86672AEEA10A71CD6E653DE2E335">Davies, Anthony</mods:namePart>
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</mods:name>
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<mods:name id="A68B53A364F21634CB5ED49B03D5326A" type="personal">
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<mods:role id="DA181ED18F54DF04D6F387A5F60F679C">
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<mods:roleTerm id="557DCEE8AF87DCBF2F89BD75C70F5608">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="978B8EE1E9F49BD9F46A163D72340E5B">Bourdon, Caroline</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="4A5BACA0064F744AA8ABF3ADB4857D61">text</mods:typeOfResource>
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<mods:titleInfo id="6B27D6F47A9084E5BFF331D2F66E2E19">
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<mods:title id="3008546EAB91200921B9CEEFC85815F9">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="BCAEED3DF006BC24D44850C6094EDABE">
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<mods:date id="7DA3CC2E0B87BB61F069009B2BBF33A6">2018</mods:date>
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<mods:detail id="66B3B45783C2090133CE920BDB489317" type="pubDate">
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<mods:number id="2EFC08FDE4B4978CBD7B57A65BD3D41A">2018-08-31</mods:number>
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</mods:detail>
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<mods:detail id="59937D50B3A7CEC0FA3462BEA14197A9" type="issue">
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<mods:number id="7B68204FEAAAB9715230E6EF9AC9B329">648</mods:number>
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</mods:detail>
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<mods:extent id="DA51E3879375D7D748A5082B7EDB298D" unit="page">
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<mods:start id="54692D883E08D6EC84F4903F1D2F090F">1</mods:start>
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<mods:end id="7A2583289C6611D9AC79E7145E412F80">12</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="E8D5D50D823DAE9D0B4E0F3F20806781">journal article</mods:classification>
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<mods:identifier id="3E72890AE4635B63C992568A94C78940" type="DOI">10.5281/zenodo.3708236</mods:identifier>
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<mods:identifier id="72E5E938396C2E3AD1EFE08915114987" type="GBIF-Dataset">6d350404-6f84-4fdc-83ef-65a70583acf8</mods:identifier>
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<mods:identifier id="FE363CF6C9CEED169D742F79315F2A26" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="F642722813571065D00955E158508F1D" type="Zenodo-Dep">3708236</mods:identifier>
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<mods:identifier id="39F0E75360DA2688F00989CE11E35406" type="ZooBank">DBEAD659-A6B3-4579-9F64-D4D2C1301D10</mods:identifier>
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</mods:mods>
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<treatment id="03823B711902FF8EFF7686A052C217F2" ID-DOI="http://doi.org/10.5281/zenodo.3718347" ID-GBIF-Taxon="162485250" ID-Zenodo-Dep="3718347" LSID="urn:lsid:plazi:treatment:03823B711902FF8EFF7686A052C217F2" httpUri="http://treatment.plazi.org/id/03823B711902FF8EFF7686A052C217F2" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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<subSubSection id="C331D9EC1902FF89FF7686A052931101" box="[192,1202,1278,1303]" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="8B948A671902FF89FF7686A052931101" blockId="3.[192,1202,1278,1303]" box="[192,1202,1278,1303]" pageId="3" pageNumber="2">
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<heading id="D0DC3D0B1902FF89FF7686A052931101" bold="true" box="[192,1202,1278,1303]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
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<emphasis id="B95F56751902FF89FF7686A052931101" bold="true" box="[192,1202,1278,1303]" pageId="3" pageNumber="2">
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Subgenus
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<taxonomicName id="4C2BF1E41902FF89FEF886A1550F1101" ID-CoL="8XR36" authority="Klimaszewski" authorityName="Klimaszewski" box="[334,814,1278,1303]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Hydrosmectomorpha">
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<emphasis id="B95F56751902FF89FEF886A154481100" bold="true" box="[334,617,1278,1302]" italics="true" pageId="3" pageNumber="2">Hydrosmectomorpha</emphasis>
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Klimaszewski
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</taxonomicName>
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and
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<taxonomicName id="4C2BF1E41902FF89FCC786A055C31101" authority="Klimaszewski & Webster & Davies & Bourdon, 2018" authorityName="Klimaszewski & Webster & Davies & Bourdon" authorityYear="2018" box="[881,994,1279,1303]" pageId="3" pageNumber="2" rank="subGenus" status="subgen. nov." subGenus="Webster">Webster</taxonomicName>
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,
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<taxonomicNameLabel id="A26CEB0E1902FF89FC4686A052931101" box="[1008,1202,1279,1303]" pageId="3" pageNumber="2" rank="subGenus">new subgenus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C331D9EC1902FF89FF76877057931171" pageId="3" pageNumber="2" type="type_taxon">
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<paragraph id="8B948A671902FF89FF76877057931171" blockId="3.[192,1440,1327,1383]" pageId="3" pageNumber="2">
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<emphasis id="B95F56751902FF89FF76877054021151" bold="true" box="[192,547,1327,1351]" pageId="3" pageNumber="2">
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<typeStatus id="549034C51902FF89FF76877057251151" box="[192,260,1327,1351]" pageId="3" pageNumber="2">Type</typeStatus>
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species of subgenus.
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</emphasis>
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<taxonomicName id="4C2BF1E41902FF89FD9C8770525B1151" ID-CoL="JCRK" authorityName="Webester and Klimaszewski" authorityYear="2018" box="[554,1146,1327,1351]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="meduxnekeagensis" subGenus="Hydrosmectomorpha">
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<emphasis id="B95F56751902FF89FD9C8770545A1151" box="[554,635,1327,1351]" italics="true" pageId="3" pageNumber="2">Atheta</emphasis>
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(
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<emphasis id="B95F56751902FF89FD3D877055A91151" box="[651,904,1327,1351]" italics="true" pageId="3" pageNumber="2">Hydrosmectomorpha</emphasis>
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)
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<emphasis id="B95F56751902FF89FC2F8770525B1151" box="[921,1146,1327,1351]" italics="true" pageId="3" pageNumber="2">meduxnekeagensis</emphasis>
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</taxonomicName>
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Webster and Klimaszewski,
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<emphasis id="B95F56751902FF89FEBE8710578A1171" bold="true" box="[264,427,1359,1383]" pageId="3" pageNumber="2">
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<taxonomicNameLabel id="A26CEB0E1902FF89FEBE8710578A1171" box="[264,427,1359,1383]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C331D9EC1902FF89FF768720578A11CE" pageId="3" pageNumber="2" type="etymology">
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<paragraph id="8B948A671902FF89FF768720578A11CE" blockId="3.[192,1440,1407,1496]" pageId="3" pageNumber="2">
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<emphasis id="B95F56751902FF89FF768720577C1181" bold="true" box="[192,349,1407,1431]" pageId="3" pageNumber="2">Etymology.</emphasis>
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<taxonomicName id="4C2BF1E41902FF89FEDE872054481181" box="[360,617,1407,1431]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Hydrosmectomorpha">
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<emphasis id="B95F56751902FF89FEDE872054481181" box="[360,617,1407,1431]" italics="true" pageId="3" pageNumber="2">Hydrosmectomorpha</emphasis>
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</taxonomicName>
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, meaning body like
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<taxonomicName id="4C2BF1E41902FF89FCC7872052321181" box="[881,1043,1407,1431]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B95F56751902FF89FCC7872052321181" box="[881,1043,1407,1431]" italics="true" pageId="3" pageNumber="2">Hydrosmecta</emphasis>
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</taxonomicName>
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, an adjective derived from the generic name
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<taxonomicName id="4C2BF1E41902FF89FEDB87C0542C11A1" box="[365,525,1439,1463]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="B95F56751902FF89FEDB87C0542C11A1" box="[365,525,1439,1463]" italics="true" pageId="3" pageNumber="2">Hydrosmecta</emphasis>
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</taxonomicName>
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to which species of this subgenus are externally superficially similar. The gender is feminine.
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</paragraph>
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</subSubSection>
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<subSubSection id="C331D9EC1902FF8EFF7687B053BE15C5" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="description">
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<paragraph id="8B948A671902FF8EFF7687B053BE15C5" blockId="3.[192,1441,1519,1928]" lastBlockId="4.[192,1441,217,467]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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<emphasis id="B95F56751902FF89FF7687B0574B1211" bold="true" box="[192,362,1519,1543]" pageId="3" pageNumber="2">Description.</emphasis>
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Body narrowly elongate, subparallel, slightly flattened (
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<figureCitation id="131096E21902FF89FBA387AF527A121E" box="[1045,1115,1520,1544]" captionStart="Figures 8–15" captionStartId="10.[192,287,1819,1841]" captionTargetBox="[236,1423,250,1738]" captionTargetId="figure@10.[192,1440,242,1747]" captionTargetPageId="10" captionText="Figures 8–15. Atheta (H.) newfoundlandica (Klimaszewski and Langor). 8) Habitus in dorsal view. 9) Median lobe of aedeagus in lateral view. 10) Median lobe of aedeagus in dorsal view. 11) Male tergite VIII. 12) Male sternite VIII. 13) Female tergite VIII. 14) Female sternite VIII. 15) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708240" httpUri="https://zenodo.org/record/3708240/files/figure.png" pageId="3" pageNumber="2">Fig. 8</figureCitation>
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,
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<figureCitation id="131096E21902FF89FBD187AF52A6121E" box="[1127,1159,1520,1544]" captionStart="Figures 16–23" captionStartId="11.[192,287,1819,1841]" captionTargetBox="[212,1425,249,1783]" captionTargetId="figure@11.[192,1440,213,1792]" captionTargetPageId="11" captionText="Figures 16–23. Atheta (H.) meduxnekeagensis Webester and Klimaszewski, sp. nov. (NB specimen). 17, 18) median lobe of aedeagus in lateral view (17BC, 18 NB). 19a, b) Male tergite VIII (a - NB, b - BC). 20a, b) Male sternite VIII. 21) Female tergite VIII (a - NB, b - BC). 22) Female sternite VIII. 23) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708242" httpUri="https://zenodo.org/record/3708242/files/figure.png" pageId="3" pageNumber="2">16</figureCitation>
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,
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<figureCitation id="131096E21902FF89FB2587AF5292121E" box="[1171,1203,1520,1544]" captionStart="Figures 24–31" captionStartId="12.[192,287,1819,1841]" captionTargetBox="[207,1407,289,1718]" captionTargetId="figure@12.[192,1440,264,1741]" captionTargetPageId="12" captionText="Figures 24–31. Atheta (H.) vincenti Webester and Klimaszewski, sp. nov. 24) Habitus in dorsal view. 25) Median lobe of aedeagus in lateral view. 26) Male tergite VIII. 27) Male sternite VIII. 28) Female tergite VIII. 29) Female sternite VIII. 30, 31) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708244" httpUri="https://zenodo.org/record/3708244/files/figure.png" pageId="3" pageNumber="2">24</figureCitation>
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,
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<figureCitation id="131096E21902FF89FB0887AF52FF121E" box="[1214,1246,1520,1544]" captionStart="Figures 32–38" captionStartId="13.[192,287,1819,1841]" captionTargetBox="[222,1407,318,1714]" captionTargetId="figure@13.[192,1440,312,1723]" captionTargetPageId="13" captionText="Figures 32–38. Atheta (H.) quebecensis Webester and Klimaszewski, sp. nov. 32) Habitus in dorsal view. 33) Median lobe of aedeagus in lateral view. 34) Male tergite VIII. 35) Male sternite VIII. 36) Female tergite VIII. 37) Female sternite VIII. 38) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708246" httpUri="https://zenodo.org/record/3708246/files/figure.png" pageId="3" pageNumber="2">32</figureCitation>
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); head rounded, pubescence on disc directed obliquely towards midline, postocular carina strong and complete, eyes large, each longer than postocular area of head in dorsal view; labrum shallowly emarginate apically (
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<figureCitation id="131096E21902FF89FAFD846F53AF125E" box="[1355,1422,1584,1608]" captionStart="Figures 1–7" captionStartId="9.[192,285,1819,1841]" captionTargetBox="[234,1415,328,1668]" captionTargetId="figure@9.[192,1440,324,1677]" captionTargetPageId="9" captionText="Figures 1–7. Mouthparts. 1) Clypeus of Atheta (H.) newfoundlandica. 2, 3) Mandibles of Atheta (H.) newfoundlandica. 4, 5) Mandibles of Atheta (H.) meduxnekeagensis. 6) Mentum with labial palpi and ligula. 7) Maxilla with maxillary palpus. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3708238" httpUri="https://zenodo.org/record/3708238/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
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); antennomeres VIII-X clearly elongate (
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<figureCitation id="131096E21902FF89FD2D840F54C3127E" box="[667,738,1616,1640]" captionStart="Figures 8–15" captionStartId="10.[192,287,1819,1841]" captionTargetBox="[236,1423,250,1738]" captionTargetId="figure@10.[192,1440,242,1747]" captionTargetPageId="10" captionText="Figures 8–15. Atheta (H.) newfoundlandica (Klimaszewski and Langor). 8) Habitus in dorsal view. 9) Median lobe of aedeagus in lateral view. 10) Median lobe of aedeagus in dorsal view. 11) Male tergite VIII. 12) Male sternite VIII. 13) Female tergite VIII. 14) Female sternite VIII. 15) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708240" httpUri="https://zenodo.org/record/3708240/files/figure.png" pageId="3" pageNumber="2">Fig. 8</figureCitation>
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,
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<figureCitation id="131096E21902FF89FD58840F552F127E" box="[750,782,1616,1640]" captionStart="Figures 24–31" captionStartId="12.[192,287,1819,1841]" captionTargetBox="[207,1407,289,1718]" captionTargetId="figure@12.[192,1440,264,1741]" captionTargetPageId="12" captionText="Figures 24–31. Atheta (H.) vincenti Webester and Klimaszewski, sp. nov. 24) Habitus in dorsal view. 25) Median lobe of aedeagus in lateral view. 26) Male tergite VIII. 27) Male sternite VIII. 28) Female tergite VIII. 29) Female sternite VIII. 30, 31) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708244" httpUri="https://zenodo.org/record/3708244/files/figure.png" pageId="3" pageNumber="2">24</figureCitation>
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,
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<figureCitation id="131096E21902FF89FCAC840F551B127E" box="[794,826,1616,1640]" captionStart="Figures 32–38" captionStartId="13.[192,287,1819,1841]" captionTargetBox="[222,1407,318,1714]" captionTargetId="figure@13.[192,1440,312,1723]" captionTargetPageId="13" captionText="Figures 32–38. Atheta (H.) quebecensis Webester and Klimaszewski, sp. nov. 32) Habitus in dorsal view. 33) Median lobe of aedeagus in lateral view. 34) Male tergite VIII. 35) Male sternite VIII. 36) Female tergite VIII. 37) Female sternite VIII. 38) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708246" httpUri="https://zenodo.org/record/3708246/files/figure.png" pageId="3" pageNumber="2">32</figureCitation>
|
|||
|
), or subquadrate to slightly elongate (
|
|||
|
<figureCitation id="131096E21902FF89FABE840F537C127E" box="[1288,1373,1616,1640]" captionStart="Figures 16–23" captionStartId="11.[192,287,1819,1841]" captionTargetBox="[212,1425,249,1783]" captionTargetId="figure@11.[192,1440,213,1792]" captionTargetPageId="11" captionText="Figures 16–23. Atheta (H.) meduxnekeagensis Webester and Klimaszewski, sp. nov. (NB specimen). 17, 18) median lobe of aedeagus in lateral view (17BC, 18 NB). 19a, b) Male tergite VIII (a - NB, b - BC). 20a, b) Male sternite VIII. 21) Female tergite VIII (a - NB, b - BC). 22) Female sternite VIII. 23) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708242" httpUri="https://zenodo.org/record/3708242/files/figure.png" pageId="3" pageNumber="2">Fig. 16</figureCitation>
|
|||
|
); last maxillary palpomere short and needle-shaped (
|
|||
|
<figureCitation id="131096E21902FF89FD46842F5515129E" box="[752,820,1648,1672]" captionStart="Figures 1–7" captionStartId="9.[192,285,1819,1841]" captionTargetBox="[234,1415,328,1668]" captionTargetId="figure@9.[192,1440,324,1677]" captionTargetPageId="9" captionText="Figures 1–7. Mouthparts. 1) Clypeus of Atheta (H.) newfoundlandica. 2, 3) Mandibles of Atheta (H.) newfoundlandica. 4, 5) Mandibles of Atheta (H.) meduxnekeagensis. 6) Mentum with labial palpi and ligula. 7) Maxilla with maxillary palpus. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3708238" httpUri="https://zenodo.org/record/3708238/files/figure.png" pageId="3" pageNumber="2">Fig. 7</figureCitation>
|
|||
|
); ligula elongate, partially subdivided medially (
|
|||
|
<figureCitation id="131096E21902FF89FAC7842F56F112BE" captionStart="Figures 1–7" captionStartId="9.[192,285,1819,1841]" captionTargetBox="[234,1415,328,1668]" captionTargetId="figure@9.[192,1440,324,1677]" captionTargetPageId="9" captionText="Figures 1–7. Mouthparts. 1) Clypeus of Atheta (H.) newfoundlandica. 2, 3) Mandibles of Atheta (H.) newfoundlandica. 4, 5) Mandibles of Atheta (H.) meduxnekeagensis. 6) Mentum with labial palpi and ligula. 7) Maxilla with maxillary palpus. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3708238" httpUri="https://zenodo.org/record/3708238/files/figure.png" pageId="3" pageNumber="2">Fig. 6</figureCitation>
|
|||
|
); mandibles long and slender, internal edge of right mandible with a small or large tooth (
|
|||
|
<figureCitation id="131096E21902FF89FB4B84CF537F12BE" box="[1277,1374,1680,1704]" captionStart="Figures 1–7" captionStartId="9.[192,285,1819,1841]" captionTargetBox="[234,1415,328,1668]" captionTargetId="figure@9.[192,1440,324,1677]" captionTargetPageId="9" captionText="Figures 1–7. Mouthparts. 1) Clypeus of Atheta (H.) newfoundlandica. 2, 3) Mandibles of Atheta (H.) newfoundlandica. 4, 5) Mandibles of Atheta (H.) meduxnekeagensis. 6) Mentum with labial palpi and ligula. 7) Maxilla with maxillary palpus. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3708238" httpUri="https://zenodo.org/record/3708238/files/figure.png" pageId="3" pageNumber="2">Fig. 3, 5</figureCitation>
|
|||
|
), and left one with or without median tooth (
|
|||
|
<figureCitation id="131096E21902FF89FD3D84EF54CD12DE" box="[651,748,1712,1736]" captionStart="Figures 1–7" captionStartId="9.[192,285,1819,1841]" captionTargetBox="[234,1415,328,1668]" captionTargetId="figure@9.[192,1440,324,1677]" captionTargetPageId="9" captionText="Figures 1–7. Mouthparts. 1) Clypeus of Atheta (H.) newfoundlandica. 2, 3) Mandibles of Atheta (H.) newfoundlandica. 4, 5) Mandibles of Atheta (H.) meduxnekeagensis. 6) Mentum with labial palpi and ligula. 7) Maxilla with maxillary palpus. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3708238" httpUri="https://zenodo.org/record/3708238/files/figure.png" pageId="3" pageNumber="2">Fig. 2, 4</figureCitation>
|
|||
|
); pronotum slightly transverse, about one fifth (0.2) wider than long, broadest at apical third, pubescence on disc directed anteriad along midline, antero-laterad on each side forming arcuate lines, hind angles of disc evident to distinct, obtusely rounded, hypomera entirely visible in lateral view; elytra transverse, at suture 2/3 to 4/5 as long as wide, distinctly broader than pronotum, flat, shoulders rounded, subrectangular, posterior margin of each elytron broadly arcu- ate, latero-apical emarginations weakly defined, pubescence on disc directed obliquely posteriad from suture; abdomen subparallel; legs long.
|
|||
|
<emphasis id="B95F56751902FF89FD02852F54D0139E" box="[692,753,1904,1928]" italics="true" pageId="3" pageNumber="2">Male</emphasis>
|
|||
|
. Tergite VIII with apical margin truncate and slightly sinuate medially (
|
|||
|
<figureCitation id="131096E21905FF8EFE2C828657D114E4" box="[410,496,217,242]" captionStart="Figures 32–38" captionStartId="13.[192,287,1819,1841]" captionTargetBox="[222,1407,318,1714]" captionTargetId="figure@13.[192,1440,312,1723]" captionTargetPageId="13" captionText="Figures 32–38. Atheta (H.) quebecensis Webester and Klimaszewski, sp. nov. 32) Habitus in dorsal view. 33) Median lobe of aedeagus in lateral view. 34) Male tergite VIII. 35) Male sternite VIII. 36) Female tergite VIII. 37) Female sternite VIII. 38) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708246" httpUri="https://zenodo.org/record/3708246/files/figure.png" pageId="4" pageNumber="3">Fig. 34</figureCitation>
|
|||
|
), or sinuate and denticulate laterally and minutely denticulate at middle (
|
|||
|
<figureCitation id="131096E21905FF8EFAC6828656FF1504" captionStart="Figures 8–15" captionStartId="10.[192,287,1819,1841]" captionTargetBox="[236,1423,250,1738]" captionTargetId="figure@10.[192,1440,242,1747]" captionTargetPageId="10" captionText="Figures 8–15. Atheta (H.) newfoundlandica (Klimaszewski and Langor). 8) Habitus in dorsal view. 9) Median lobe of aedeagus in lateral view. 10) Median lobe of aedeagus in dorsal view. 11) Male tergite VIII. 12) Male sternite VIII. 13) Female tergite VIII. 14) Female sternite VIII. 15) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708240" httpUri="https://zenodo.org/record/3708240/files/figure.png" pageId="4" pageNumber="3">Fig. 11</figureCitation>
|
|||
|
), or obtusely produced laterally, without teeth and deeply emarginate medially (
|
|||
|
<figureCitation id="131096E21905FF8EFB7A82A553051504" box="[1228,1316,250,274]" captionStart="Figures 24–31" captionStartId="12.[192,287,1819,1841]" captionTargetBox="[207,1407,289,1718]" captionTargetId="figure@12.[192,1440,264,1741]" captionTargetPageId="12" captionText="Figures 24–31. Atheta (H.) vincenti Webester and Klimaszewski, sp. nov. 24) Habitus in dorsal view. 25) Median lobe of aedeagus in lateral view. 26) Male tergite VIII. 27) Male sternite VIII. 28) Female tergite VIII. 29) Female sternite VIII. 30, 31) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708244" httpUri="https://zenodo.org/record/3708244/files/figure.png" pageId="4" pageNumber="3">Fig. 26</figureCitation>
|
|||
|
); tubus of median lobe of aedeagus moderately long, in lateral view with ventral margin straight to arcuate, apex moderately curved ventrad, internal sac structures small and inconspicuous, ventral margin of crista apicalis moderate to long, projecting at an angle to bulbus, rectangular basally in some specimens also at base of median lobe (
|
|||
|
<figureCitation id="131096E21905FF8EFE0A832554221584" box="[444,515,378,402]" captionStart="Figures 8–15" captionStartId="10.[192,287,1819,1841]" captionTargetBox="[236,1423,250,1738]" captionTargetId="figure@10.[192,1440,242,1747]" captionTargetPageId="10" captionText="Figures 8–15. Atheta (H.) newfoundlandica (Klimaszewski and Langor). 8) Habitus in dorsal view. 9) Median lobe of aedeagus in lateral view. 10) Median lobe of aedeagus in dorsal view. 11) Male tergite VIII. 12) Male sternite VIII. 13) Female tergite VIII. 14) Female sternite VIII. 15) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708240" httpUri="https://zenodo.org/record/3708240/files/figure.png" pageId="4" pageNumber="3">Fig. 9</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFDB8832554781584" box="[526,601,378,402]" captionStart="Figures 16–23" captionStartId="11.[192,287,1819,1841]" captionTargetBox="[212,1425,249,1783]" captionTargetId="figure@11.[192,1440,213,1792]" captionTargetPageId="11" captionText="Figures 16–23. Atheta (H.) meduxnekeagensis Webester and Klimaszewski, sp. nov. (NB specimen). 17, 18) median lobe of aedeagus in lateral view (17BC, 18 NB). 19a, b) Male tergite VIII (a - NB, b - BC). 20a, b) Male sternite VIII. 21) Female tergite VIII (a - NB, b - BC). 22) Female sternite VIII. 23) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708242" httpUri="https://zenodo.org/record/3708242/files/figure.png" pageId="4" pageNumber="3">17, 18</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFDD2832554A51584" box="[612,644,378,402]" captionStart="Figures 24–31" captionStartId="12.[192,287,1819,1841]" captionTargetBox="[207,1407,289,1718]" captionTargetId="figure@12.[192,1440,264,1741]" captionTargetPageId="12" captionText="Figures 24–31. Atheta (H.) vincenti Webester and Klimaszewski, sp. nov. 24) Habitus in dorsal view. 25) Median lobe of aedeagus in lateral view. 26) Male tergite VIII. 27) Male sternite VIII. 28) Female tergite VIII. 29) Female sternite VIII. 30, 31) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708244" httpUri="https://zenodo.org/record/3708244/files/figure.png" pageId="4" pageNumber="3">25</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFD398325548E1584" box="[655,687,378,402]" captionStart="Figures 32–38" captionStartId="13.[192,287,1819,1841]" captionTargetBox="[222,1407,318,1714]" captionTargetId="figure@13.[192,1440,312,1723]" captionTargetPageId="13" captionText="Figures 32–38. Atheta (H.) quebecensis Webester and Klimaszewski, sp. nov. 32) Habitus in dorsal view. 33) Median lobe of aedeagus in lateral view. 34) Male tergite VIII. 35) Male sternite VIII. 36) Female tergite VIII. 37) Female sternite VIII. 38) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708246" httpUri="https://zenodo.org/record/3708246/files/figure.png" pageId="4" pageNumber="3">33</figureCitation>
|
|||
|
).
|
|||
|
<emphasis id="B95F56751905FF8EFD758325553D1584" box="[707,796,378,402]" italics="true" pageId="4" pageNumber="3">Female</emphasis>
|
|||
|
. Tergite VIII with apical margin truncate, sinuate or broadly emarginate (
|
|||
|
<figureCitation id="131096E21905FF8EFE0A83C5543115A4" box="[444,528,410,434]" captionStart="Figures 8–15" captionStartId="10.[192,287,1819,1841]" captionTargetBox="[236,1423,250,1738]" captionTargetId="figure@10.[192,1440,242,1747]" captionTargetPageId="10" captionText="Figures 8–15. Atheta (H.) newfoundlandica (Klimaszewski and Langor). 8) Habitus in dorsal view. 9) Median lobe of aedeagus in lateral view. 10) Median lobe of aedeagus in dorsal view. 11) Male tergite VIII. 12) Male sternite VIII. 13) Female tergite VIII. 14) Female sternite VIII. 15) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708240" httpUri="https://zenodo.org/record/3708240/files/figure.png" pageId="4" pageNumber="3">Fig. 13</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFDAD83C5541A15A4" box="[539,571,410,434]" captionStart="Figures 16–23" captionStartId="11.[192,287,1819,1841]" captionTargetBox="[212,1425,249,1783]" captionTargetId="figure@11.[192,1440,213,1792]" captionTargetPageId="11" captionText="Figures 16–23. Atheta (H.) meduxnekeagensis Webester and Klimaszewski, sp. nov. (NB specimen). 17, 18) median lobe of aedeagus in lateral view (17BC, 18 NB). 19a, b) Male tergite VIII (a - NB, b - BC). 20a, b) Male sternite VIII. 21) Female tergite VIII (a - NB, b - BC). 22) Female sternite VIII. 23) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708242" httpUri="https://zenodo.org/record/3708242/files/figure.png" pageId="4" pageNumber="3">21</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFDF083C5544715A4" box="[582,614,410,434]" captionStart="Figures 24–31" captionStartId="12.[192,287,1819,1841]" captionTargetBox="[207,1407,289,1718]" captionTargetId="figure@12.[192,1440,264,1741]" captionTargetPageId="12" captionText="Figures 24–31. Atheta (H.) vincenti Webester and Klimaszewski, sp. nov. 24) Habitus in dorsal view. 25) Median lobe of aedeagus in lateral view. 26) Male tergite VIII. 27) Male sternite VIII. 28) Female tergite VIII. 29) Female sternite VIII. 30, 31) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708244" httpUri="https://zenodo.org/record/3708244/files/figure.png" pageId="4" pageNumber="3">28</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFDC683C554B115A4" box="[624,656,410,434]" captionStart="Figures 32–38" captionStartId="13.[192,287,1819,1841]" captionTargetBox="[222,1407,318,1714]" captionTargetId="figure@13.[192,1440,312,1723]" captionTargetPageId="13" captionText="Figures 32–38. Atheta (H.) quebecensis Webester and Klimaszewski, sp. nov. 32) Habitus in dorsal view. 33) Median lobe of aedeagus in lateral view. 34) Male tergite VIII. 35) Male sternite VIII. 36) Female tergite VIII. 37) Female sternite VIII. 38) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708246" httpUri="https://zenodo.org/record/3708246/files/figure.png" pageId="4" pageNumber="3">36</figureCitation>
|
|||
|
); spermatheca small, capsule club-shaped, narrow or broad, stem moderately long, narrow, sinuate or straight and more or less swollen posteriorly (
|
|||
|
<figureCitation id="131096E21905FF8EFB2383E552C815C4" box="[1173,1257,442,466]" captionStart="Figures 8–15" captionStartId="10.[192,287,1819,1841]" captionTargetBox="[236,1423,250,1738]" captionTargetId="figure@10.[192,1440,242,1747]" captionTargetPageId="10" captionText="Figures 8–15. Atheta (H.) newfoundlandica (Klimaszewski and Langor). 8) Habitus in dorsal view. 9) Median lobe of aedeagus in lateral view. 10) Median lobe of aedeagus in dorsal view. 11) Male tergite VIII. 12) Male sternite VIII. 13) Female tergite VIII. 14) Female sternite VIII. 15) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708240" httpUri="https://zenodo.org/record/3708240/files/figure.png" pageId="4" pageNumber="3">Fig. 15</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFB4283E4533515C5" box="[1268,1300,443,467]" captionStart="Figures 16–23" captionStartId="11.[192,287,1819,1841]" captionTargetBox="[212,1425,249,1783]" captionTargetId="figure@11.[192,1440,213,1792]" captionTargetPageId="11" captionText="Figures 16–23. Atheta (H.) meduxnekeagensis Webester and Klimaszewski, sp. nov. (NB specimen). 17, 18) median lobe of aedeagus in lateral view (17BC, 18 NB). 19a, b) Male tergite VIII (a - NB, b - BC). 20a, b) Male sternite VIII. 21) Female tergite VIII (a - NB, b - BC). 22) Female sternite VIII. 23) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708242" httpUri="https://zenodo.org/record/3708242/files/figure.png" pageId="4" pageNumber="3">23</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFAA883E4534915C5" box="[1310,1384,443,467]" captionStart="Figures 24–31" captionStartId="12.[192,287,1819,1841]" captionTargetBox="[207,1407,289,1718]" captionTargetId="figure@12.[192,1440,264,1741]" captionTargetPageId="12" captionText="Figures 24–31. Atheta (H.) vincenti Webester and Klimaszewski, sp. nov. 24) Habitus in dorsal view. 25) Median lobe of aedeagus in lateral view. 26) Male tergite VIII. 27) Male sternite VIII. 28) Female tergite VIII. 29) Female sternite VIII. 30, 31) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708244" httpUri="https://zenodo.org/record/3708244/files/figure.png" pageId="4" pageNumber="3">30, 31</figureCitation>
|
|||
|
,
|
|||
|
<figureCitation id="131096E21905FF8EFAC483E453B315C5" box="[1394,1426,443,467]" captionStart="Figures 32–38" captionStartId="13.[192,287,1819,1841]" captionTargetBox="[222,1407,318,1714]" captionTargetId="figure@13.[192,1440,312,1723]" captionTargetPageId="13" captionText="Figures 32–38. Atheta (H.) quebecensis Webester and Klimaszewski, sp. nov. 32) Habitus in dorsal view. 33) Median lobe of aedeagus in lateral view. 34) Male tergite VIII. 35) Male sternite VIII. 36) Female tergite VIII. 37) Female sternite VIII. 38) Spermatheca. Scale line for habitus = 1 mm, remaining scale lines = 0.2 mm. Arrows indicate important diagnostic features." figureDoi="http://doi.org/10.5281/zenodo.3708246" httpUri="https://zenodo.org/record/3708246/files/figure.png" pageId="4" pageNumber="3">38</figureCitation>
|
|||
|
).
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
<subSubSection id="C331D9EC1905FF8EFF7683B552C217F2" pageId="4" pageNumber="3" type="discussion">
|
|||
|
<paragraph id="8B948A671905FF8EFF7683B552B316D5" blockId="4.[192,1440,490,997]" pageId="4" pageNumber="3">
|
|||
|
<emphasis id="B95F56751905FF8EFF7683B5577C1614" bold="true" box="[192,349,490,514]" pageId="4" pageNumber="3">Discussion.</emphasis>
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFED283B554401614" box="[356,609,490,514]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Hydrosmectomorpha">
|
|||
|
<emphasis id="B95F56751905FF8EFED283B554401614" box="[356,609,490,514]" italics="true" pageId="4" pageNumber="3">Hydrosmectomorpha</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
, is distinguishable from other subgenera of
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFB3583B552F51614" box="[1155,1236,490,514]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B95F56751905FF8EFB3583B552F51614" box="[1155,1236,490,514]" italics="true" pageId="4" pageNumber="3">Atheta</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
by the following combination of characters: body narrow, subparallel and flattened; elytra markedly broader than remain- der of body; distinctive pattern of pubescence on pronotum and elytra (described above); antennomeres VIII–X distinctly to slightly elongate; ligula weakly subdivided apically; legs long; apical margin of male tergite VIII lacking prominent teeth; spermatheca small, with stem short and simple, straight or sinuate; all known species occur among cobblestones and gravel along margins of clear, fast flowing rivers, near the outflow of brooks into rivers, and along lake margins with wave action.
|
|||
|
</paragraph>
|
|||
|
<paragraph id="8B948A671905FF8EFF5E809352C217F2" blockId="4.[192,1440,490,997]" pageId="4" pageNumber="3">
|
|||
|
The median lobe of the aedeagus in
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFD288094551F16F5" box="[670,830,715,739]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B95F56751905FF8EFD288094551F16F5" box="[670,830,715,739]" italics="true" pageId="4" pageNumber="3">Hydrosmecta</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
has a blade-like, triangular projection at the base of the tubus in lateral view to which parameres are attached, this structure is narrowly elongate along ventrally flat part of bulbus, in
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFDE1815355751732" box="[599,852,780,804]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Hydrosmectomorpha">
|
|||
|
<emphasis id="B95F56751905FF8EFDE1815355751732" box="[599,852,780,804]" italics="true" pageId="4" pageNumber="3">Hydrosmectomorpha</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
. Male tergite VIII in
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFBDB8153532C1732" box="[1133,1293,780,804]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B95F56751905FF8EFBDB8153532C1732" box="[1133,1293,780,804]" italics="true" pageId="4" pageNumber="3">Hydrosmecta</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
is truncate apically and not modified, while in
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFDD98173554D1752" box="[623,876,812,836]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Hydrosmectomorpha">
|
|||
|
<emphasis id="B95F56751905FF8EFDD98173554D1752" box="[623,876,812,836]" italics="true" pageId="4" pageNumber="3">Hydrosmectomorpha</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
it is truncate and slightly emarginate, deeply notched, or with minute teeth. The stem of spermatheca in
|
|||
|
<emphasis id="B95F56751905FF8EFC2E8113521E1772" box="[920,1087,844,868]" italics="true" pageId="4" pageNumber="3">
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFC2E8113521A1772" box="[920,1083,844,868]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">Hydrosmecta</taxonomicName>
|
|||
|
,
|
|||
|
</emphasis>
|
|||
|
is coiled posteriorly, broadly looped, or irregularly twisted, while in
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFD2F813355B71792" box="[665,918,876,900]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Hydrosmectomorpha">
|
|||
|
<emphasis id="B95F56751905FF8EFD2F813355B71792" box="[665,918,876,900]" italics="true" pageId="4" pageNumber="3">Hydrosmectomorpha</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
it is straight or slightly sinuate with small apical swelling. The external similarity of
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFD6581D355F117B2" box="[723,976,908,932]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="2" phylum="Arthropoda" rank="subGenus" subGenus="Hydrosmectomorpha">
|
|||
|
<emphasis id="B95F56751905FF8EFD6581D355F117B2" box="[723,976,908,932]" italics="true" pageId="4" pageNumber="3">Hydrosmectomorpha</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
species (e.g., flattened body, similar pronotal pubescence pattern), to those of
|
|||
|
<taxonomicName id="4C2BF1E41905FF8EFD0F81F3557817D2" box="[697,857,940,964]" class="Insecta" family="Staphylinidae" genus="Hydrosmecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
|
|||
|
<emphasis id="B95F56751905FF8EFD0F81F3557817D2" box="[697,857,940,964]" italics="true" pageId="4" pageNumber="3">Hydrosmecta</emphasis>
|
|||
|
</taxonomicName>
|
|||
|
represent convergent evolution and adaptation to live among cobblestones, gravel and sand along margins of clear, fast-flowing water.
|
|||
|
</paragraph>
|
|||
|
</subSubSection>
|
|||
|
</treatment>
|
|||
|
</document>
|