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<document id="A1095D92B0BCA07F43DC9F412A0ED6E9" ID-DOI="10.5281/zenodo.172764" ID-GBIF-Dataset="57f77567-9f74-4e9f-a79a-a19a41e960c8" ID-ISSN="1175­5326" ID-Zenodo-Dep="172764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459761781846" checkinUser="plazi" docAuthor="Blahnik, Roger J. &amp; Holzenthal, Ralph W." docDate="2006" docId="03FC8793FFCE6F4A846AFA5BC43219CD" docLanguage="en" docName="zt01233p052.pdf" docOrigin="Zootaxa 1233" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Culoptila nahuatl Flint" docType="treatment" docVersion="10" lastPageNumber="34" masterDocId="FFC5FFEBFFD16F6B8562FFAFC6051D3B" masterDocTitle="Revision of the genus Culoptila (Trichoptera: Glossosomatidae)" masterLastPageNumber="52" masterPageNumber="1" pageNumber="32" updateTime="1698215326134" updateUser="plazi">
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<mods:title id="4CA3E86194CA17A16A49ED2D46D4A261">Revision of the genus Culoptila (Trichoptera: Glossosomatidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="5C939DD416780CDBEA9EB876AE7C6A9D">Blahnik, Roger J.</mods:namePart>
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<mods:namePart id="5A0D6C5C05844F388B30D8F11DE33E20">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="D5D526EFA0A8DB967DD6D7F51745140C">text</mods:typeOfResource>
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<mods:title id="869E6EC954D0315CAB815497D6EE5540">Zootaxa</mods:title>
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<mods:part id="4B04DC49C74036445ED59F565AFA87EC">
<mods:date id="FC6F7258263E1BCB35F29E81EECEB985">2006</mods:date>
<mods:detail id="798C86604BBC288FE48FE9FB2370426B" type="volume">
<mods:number id="2A7B076FC2F5320EEBA74A71EE072E32">1233</mods:number>
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<mods:classification id="8E1A82C13E715E20E588EC4586EAEB15">journal article</mods:classification>
<mods:identifier id="525070900F5C27D997EFE15F407D9FC4" type="DOI">10.5281/zenodo.172764</mods:identifier>
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<mods:identifier id="64008D65F46A68B974160D9014B55C8C" type="ISSN">1175­5326</mods:identifier>
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<treatment id="03FC8793FFCE6F4A846AFA5BC43219CD" ID-DOI="http://doi.org/10.5281/zenodo.6256657" ID-GBIF-Taxon="119345074" ID-Zenodo-Dep="6256657" LSID="urn:lsid:plazi:treatment:03FC8793FFCE6F4A846AFA5BC43219CD" httpUri="http://treatment.plazi.org/id/03FC8793FFCE6F4A846AFA5BC43219CD" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
<subSubSection id="C34F650EFFCE6F74846AFA5BC7971B0A" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BEA3685FFCE6F74846AFA5BC4271B35" blockId="31.[264,546,1524,1585]" box="[264,546,1524,1550]" pageId="31" pageNumber="32">
<heading id="D0A281E9FFCE6F74846AFA5BC4271B35" bold="true" box="[264,546,1524,1550]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<taxonomicName id="4C554D06FFCE6F74846AFA5BC4271B35" ID-CoL="32DPX" authority="Flint" authorityName="Flint" box="[264,546,1524,1550]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nahuatl">
<emphasis id="B921EA97FFCE6F74846AFA5BC4271B35" bold="true" box="[264,546,1524,1550]" pageId="31" pageNumber="32">
<emphasis id="B921EA97FFCE6F74846AFA5BC7DB1B35" bold="true" box="[264,478,1524,1550]" italics="true" pageId="31" pageNumber="32">Culoptila nahuatl</emphasis>
Flint
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BEA3685FFCE6F74846AF9B8C7971B0A" blockId="31.[264,546,1524,1585]" box="[264,402,1559,1585]" pageId="31" pageNumber="32">
<figureCitation id="136E2A00FFCE6F74846AF9B8C75F1B0A" box="[264,346,1559,1585]" captionStart="FIGURES 27 – 28" captionStartId="32.[264,383,1827,1851]" captionTargetBox="[316,1282,486,1803]" captionTargetId="figure@32.[316,1282,486,1803]" captionTargetPageId="32" captionText="FIGURES 27 – 28. Male Genitalia. 27. Culoptila moselyi Denning: 27 A—lateral; 27 B—ventral. 28. Culoptila nahuatl Flint: 28 A—lateral; 28 B—ventral; 28 C—ventral process of sternum VI, lateral." httpUri="https://zenodo.org/record/172776/files/figure.png" pageId="31" pageNumber="32">Fig. 28</figureCitation>
A–C
</paragraph>
</subSubSection>
<subSubSection id="C34F650EFFCE6F4B846AF9C8C79C1C95" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32" type="reference_group">
<paragraph id="8BEA3685FFCE6F74846AF9C8C46D1B46" blockId="31.[264,616,1638,1662]" box="[264,616,1638,1662]" pageId="31" pageNumber="32">
<treatmentCitationGroup id="AB4511ABFFCE6F74846AF9C8C46D1B46" box="[264,616,1638,1662]" pageId="31" pageNumber="32">
<taxonomicName id="4C554D06FFCE6F74846AF9C8C4611B46" ID-CoL="32DPX" authority="Flint 1974a: 8" authorityName="Flint" authorityPageNumber="8" authorityYear="1974" box="[264,612,1638,1662]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nahuatl">
<emphasis id="B921EA97FFCE6F74846AF9C8C7C01B45" box="[264,453,1639,1662]" italics="true" pageId="31" pageNumber="32">Culoptila nahuatl</emphasis>
<treatmentCitation id="0AF41094FFCE6F7484AEF9C9C4611B46" author="Flint" box="[460,612,1638,1661]" page="8" pageId="31" pageNumber="32" year="1974">
<bibRefCitation id="EFC44B74FFCE6F7484AEF9C9C44C1B46" author="Flint" box="[460,585,1638,1661]" pageId="31" pageNumber="51" refString="Flint, O. S., Jr. (1974 a) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974" yearSuffix="a">Flint 1974a</bibRefCitation>
: 8
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEA3685FFCE6F4B846AF900C79C1C95" blockId="31.[264,1323,1711,1897]" lastBlockId="32.[264,1323,284,430]" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
This species is similar to
<taxonomicName id="4C554D06FFCE6F74874CF900C4E11BF3" ID-CoL="32DPL" box="[558,740,1711,1736]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="aluca">
<emphasis id="B921EA97FFCE6F74874CF900C4E11BF3" box="[558,740,1711,1736]" italics="true" pageId="31" pageNumber="32">Culoptila aluca</emphasis>
</taxonomicName>
,
<taxonomicName id="4C554D06FFCE6F748791F900C5651BF3" ID-CoL="32DPN" box="[755,864,1711,1736]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="azulae">
<emphasis id="B921EA97FFCE6F748791F900C5651BF3" box="[755,864,1711,1736]" italics="true" pageId="31" pageNumber="32">C. azulae</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C554D06FFCE6F7486C2F900C2351BF3" box="[928,1072,1711,1736]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="B921EA97FFCE6F7486C2F900C2351BF3" box="[928,1072,1711,1736]" italics="true" pageId="31" pageNumber="32">C. bidentata</emphasis>
</taxonomicName>
, agreeing in having a relatively short, subtruncate tergum X and an inflated phallobase with 2 prominent, curved spines. In
<taxonomicName id="4C554D06FFCE6F7484E1F950C7FA1A23" ID-CoL="32DPX" box="[387,511,1791,1816]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nahuatl">
<emphasis id="B921EA97FFCE6F7484E1F950C7FA1A23" box="[387,511,1791,1816]" italics="true" pageId="31" pageNumber="32">C. nahuatl</emphasis>
</taxonomicName>
, however, the curved phallic spine, while enlarged basally, does not have the basal enlargement strongly sclerotized.
<taxonomicName id="4C554D06FFCE6F748637F888C22D1A7B" ID-CoL="32DPX" box="[853,1064,1831,1856]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="nahuatl">
<emphasis id="B921EA97FFCE6F748637F888C22D1A7B" box="[853,1064,1831,1856]" italics="true" pageId="31" pageNumber="32">Culoptila nahuatl</emphasis>
</taxonomicName>
differs most from
<taxonomicName id="4C554D06FFCE6F748072F888C74C1A53" ID-CoL="32DPL" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="aluca">
<emphasis id="B921EA97FFCE6F748072F888C74C1A53" italics="true" pageId="31" pageNumber="32">C. aluca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D06FFCE6F7484E3F8E0C7EA1A53" ID-CoL="32DPN" box="[385,495,1871,1896]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="azulae">
<emphasis id="B921EA97FFCE6F7484E3F8E0C7EA1A53" box="[385,495,1871,1896]" italics="true" pageId="31" pageNumber="32">C. azulae</emphasis>
</taxonomicName>
in having the posterodorsal apex of the phallobase distinctly upturned, and by the very narrow ventral margin of tergum IX. It differs from
<taxonomicName id="4C554D06FFF16F4B8107FEB3C2C81C0E" ID-CoL="32DPL" box="[1125,1229,284,309]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="aluca">
<emphasis id="B921EA97FFF16F4B8107FEB3C2C81C0E" box="[1125,1229,284,309]" italics="true" pageId="32" pageNumber="33">C. aluca</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D06FFF16F4B8072FEB3C7721C66" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bidentata">
<emphasis id="B921EA97FFF16F4B8072FEB3C7721C66" italics="true" pageId="32" pageNumber="33">C. bidentata</emphasis>
</taxonomicName>
by the absence of a strong spine (and apparent fork) of the ventrolateral processes of tergum X, and from
<taxonomicName id="4C554D06FFF16F4B87D8FEC3C5351CBE" ID-CoL="32DPN" box="[698,816,364,389]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="azulae">
<emphasis id="B921EA97FFF16F4B87D8FEC3C5351CBE" box="[698,816,364,389]" italics="true" pageId="32" pageNumber="33">C. azulae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C554D06FFF16F4B861AFEC3C5FC1CBE" box="[888,1017,364,389]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bidenta">
<emphasis id="B921EA97FFF16F4B861AFEC3C5FC1CBE" box="[888,1017,364,389]" italics="true" pageId="32" pageNumber="33">C. bidenta</emphasis>
</taxonomicName>
ta by its shorter inferior appendages.
</paragraph>
</subSubSection>
<caption id="DF2A660DFFF16F4B846AF88CC3251A5B" httpUri="https://zenodo.org/record/172776/files/figure.png" pageId="32" pageNumber="33" targetBox="[316,1282,486,1803]" targetPageId="32">
<paragraph id="8BEA3685FFF16F4B846AF88CC3251A5B" blockId="32.[264,1323,1827,1888]" pageId="32" pageNumber="33">
<emphasis id="B921EA97FFF16F4B846AF88CC7C81A00" bold="true" box="[264,461,1827,1851]" pageId="32" pageNumber="33">FIGURES 27–28.</emphasis>
Male Genitalia. 27.
<taxonomicName id="4C554D06FFF16F4B87C4F88BC5CC1A00" authority="Denning" authorityName="Denning" box="[678,969,1828,1851]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="moselyi">
<emphasis id="B921EA97FFF16F4B87C4F88BC5671A00" box="[678,866,1828,1851]" italics="true" pageId="32" pageNumber="33">Culoptila moselyi</emphasis>
Denning
</taxonomicName>
: 27A—lateral; 27B—ventral. 28.
<taxonomicName id="4C554D06FFF16F4B846AF8E6C4061A5B" authority="Flint" authorityName="Flint" box="[264,515,1865,1888]" class="Insecta" family="Glossosomatidae" genus="Culoptila" kingdom="Animalia" order="Trichoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="nahuatl">
<emphasis id="B921EA97FFF16F4B846AF8E6C7C01A5B" box="[264,453,1865,1888]" italics="true" pageId="32" pageNumber="33">Culoptila nahuatl</emphasis>
Flint
</taxonomicName>
: 28A—lateral; 28B—ventral; 28C—ventral process of sternum VI, lateral.
</paragraph>
</caption>
<subSubSection id="C34F650EFFF06F4A845AFEB3C4BB1EB5" pageId="33" pageNumber="34" type="description">
<paragraph id="8BEA3685FFF06F4A845AFEB3C50A1CED" blockId="33.[264,1324,284,910]" pageId="33" pageNumber="34">
Adult. Length of forewing: male
<quantity id="4CAD9B60FFF06F4A87A0FEB3C5251C0D" box="[706,800,284,310]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="33" pageNumber="34" unit="mm" value="2.5">2.5 mm</quantity>
. Color light brown; wing without markings, anastomosis only indistinctly evident. Mesothorax of male narrow and greatly acute anteromesally; mesoscutal wart short, diamond­shaped, sharply angulate anteromesally. Mesothoracic tegulae of male rounded, greatly enlarged; tegular gland large, concertinalike, with small number of pleats (ca. 8–10).
</paragraph>
<paragraph id="8BEA3685FFF06F4A845AFE4BC4BB1EB5" blockId="33.[264,1324,284,910]" pageId="33" pageNumber="34">Male genitalia. Sternum VI process short, subtruncate. Tergum IX ventral margin rounded, tapered from anterolateral margin, distinctly produced posteroventrally. Inferior appendages short, subacute apically. Tergum X short, length slightly less than width, apex in ventral view truncate to subtruncate; ventrolateral processes with apices incurved, mesal margin acute, or nearly so, apices slightly posteriorly bent, apical bend near apex and relatively weak, mesal margin of apex bluntly rounded, not acute. Phallobase large, broadly rounded in lateral view, apicodorsal projection short, with very distinct upward inflection at base, apex subacute. Phallic apparatus with 2 phallic spines, both elongate, narrow, weakly sinuous; longer spine about 2/3 length of phallobase, basal part distinctly recurved, with weakly sclerotized basal enlargement; shorter spine subequal, but lacking recurved base and basal enlargement.</paragraph>
</subSubSection>
<subSubSection id="C34F650EFFF06F4A846AFC6BC2051945" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="8BEA3685FFF06F4A846AFC6BC7E31EE6" blockId="33.[264,1325,964,1150]" box="[264,486,964,989]" pageId="33" pageNumber="34">
<emphasis id="B921EA97FFF06F4A846AFC6BC7E31EE6" box="[264,486,964,989]" italics="true" pageId="33" pageNumber="34">Material examined</emphasis>
</paragraph>
<paragraph id="8BEA3685FFF06F4A845AFC43C2051945" blockId="33.[264,1325,964,1150]" pageId="33" pageNumber="34">
<emphasis id="B921EA97FFF06F4A845AFC43C443193D" bold="true" box="[312,582,1004,1030]" pageId="33" pageNumber="34">
<collectingCountry id="F3427615FFF06F4A845AFC43C7BC193D" box="[312,441,1004,1030]" name="Mexico" pageId="33" pageNumber="34">MEXICO</collectingCountry>
: Veracruz:
</emphasis>
Fortín de las
<collectingCountry id="F3427615FFF06F4A8799FC43C542193D" box="[763,839,1004,1030]" name="Indonesia" pageId="33" pageNumber="34">Flores</collectingCountry>
,
<date id="FFEB1045FFF06F4A863AFC43C5C9193D" box="[856,972,1004,1030]" pageId="33" pageNumber="34" value="1964-05-17">17.v.1964</date>
(Blanton et al.) —
<specimenCount id="9D53FD0CFFF06F4A81ABFC43C329193D" box="[1225,1324,1004,1030]" pageId="33" pageNumber="34" type="male">32 male</specimenCount>
<typeStatus id="54EE8827FFF06F4A846AFBBBC77E1915" box="[264,379,1044,1070]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
,
<specimenCount id="9D53FD0CFFF06F4A84EAFBBBC7EB1915" box="[392,494,1044,1070]" pageId="33" pageNumber="34" type="female">9 female</specimenCount>
<typeStatus id="54EE8827FFF06F4A8494FBBBC4631915" box="[502,614,1044,1070]" pageId="33" pageNumber="34" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED44AE40FFF06F4A8714FBBBC4D21915" box="[630,727,1044,1070]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34">NMNH</collectionCode>
); same, except
<date id="FFEB1045FFF06F4A86E9FBBBC2141915" box="[907,1041,1044,1070]" pageId="33" pageNumber="34" value="1966-07-24">24.vii.1966</date>
(Flint &amp; Ortiz) — male
<typeStatus id="54EE8827FFF06F4A846AFB93C76A196D" box="[264,367,1084,1110]" pageId="33" pageNumber="34" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED44AE40FFF06F4A841FFB93C7D1196D" LSID="urn:lsid:biocol.org:col:34871" box="[381,468,1084,1110]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34">USNM</collectionCode>
<typeStatus id="54EE8827FFF06F4A84B9FB93C410196D" box="[475,533,1084,1110]" pageId="33" pageNumber="34">Type</typeStatus>
No. 72731) (
<collectionCode id="ED44AE40FFF06F4A87D0FB93C517196D" box="[690,786,1084,1110]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="33" pageNumber="34">NMNH</collectionCode>
); Barranca de Metlac, ca. Fortín de las
<collectingCountry id="F3427615FFF06F4A81BEFB93C32D196D" box="[1244,1320,1084,1110]" name="Indonesia" pageId="33" pageNumber="34">Flores</collectingCountry>
,
<date id="FFEB1045FFF06F4A846AFBCBC7821945" box="[264,391,1124,1150]" pageId="33" pageNumber="34" value="1976-03-30">30.iii.1976</date>
(J. Bueno­Soria) —
<specimenCount id="9D53FD0CFFF06F4A871FFBCBC4F21945" box="[637,759,1124,1150]" pageId="33" pageNumber="34" type="male">129 males</specimenCount>
,
<specimenCount id="9D53FD0CFFF06F4A8661FBCBC5861945" box="[771,899,1124,1150]" pageId="33" pageNumber="34" type="female">33 females</specimenCount>
(
<collectionCode id="ED44AE40FFF06F4A86F1FBCBC5F11945" box="[915,1012,1124,1150]" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="33" pageNumber="34">UNAM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F650EFFF06F4A846AFB1BC43219CD" box="[264,567,1204,1270]" pageId="33" pageNumber="34" type="distribution">
<paragraph id="8BEA3685FFF06F4A846AFB1BC79219F6" blockId="33.[264,567,1204,1270]" box="[264,407,1204,1229]" pageId="33" pageNumber="34">
<emphasis id="B921EA97FFF06F4A846AFB1BC79219F6" box="[264,407,1204,1229]" italics="true" pageId="33" pageNumber="34">Distribution</emphasis>
</paragraph>
<paragraph id="8BEA3685FFF06F4A845AFB73C43219CD" blockId="33.[264,567,1204,1270]" box="[312,567,1244,1270]" pageId="33" pageNumber="34">
<collectingCountry id="F3427615FFF06F4A845AFB73C7A919CD" box="[312,428,1244,1270]" name="Mexico" pageId="33" pageNumber="34">MEXICO</collectingCountry>
(Veracruz).
</paragraph>
</subSubSection>
</treatment>
</document>
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