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<mods:title id="A70987D057E6D1EA5BC4831EA3609874">Notes on Iniocyphus iheringi Raffray, 1912 (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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<taxonomicName id="4C6D4D10FF97F433FF4EFBA77EDDFB85" ID-CoL="8XLZH" authority="Raffray, 1912" authorityName="Raffray" authorityYear="1912" box="[151,463,1086,1112]" class="Insecta" family="Staphylinidae" genus="Iniocyphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
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<emphasis id="B919EA81FF97F433FF4EFBA77E0CFB85" bold="true" box="[151,286,1086,1112]" italics="true" pageId="1" pageNumber="172">Iniocyphus</emphasis>
<bibRefCitation id="EFFC4B62FF97F433FEFCFBA77EDDFB85" author="Raffray, A." box="[293,463,1086,1112]" pageId="1" pageNumber="172" pagination="425 - 450" refId="ref3400" refString="Raffray, A. (1912) Especes nouvelles de Pselaphides exotiques [Coleopt.]. Annales de la Societe entomologique de France, 80 (1911), 425 - 450." type="journal article" year="1912">Raffray, 1912</bibRefCitation>
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<paragraph id="8BD23693FF97F433FF4EFBF87E32FBA7" blockId="1.[151,463,1086,1146]" box="[151,288,1121,1146]" pageId="1" pageNumber="172">
(
<figureCitation id="13562A16FF97F433FF46FBF87E05FBA7" box="[159,279,1121,1146]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–6" captionStart-2="FIGURES 7–15" captionStart-3="FIGURES 16–23" captionStartId-0="2.[151,264,1937,1960]" captionStartId-1="3.[151,264,1891,1914]" captionStartId-2="4.[151,264,1886,1909]" captionStartId-3="5.[151,264,1906,1929]" captionTargetBox-0="[157,1408,212,1906]" captionTargetBox-1="[168,1400,193,1861]" captionTargetBox-2="[159,1423,210,1848]" captionTargetBox-3="[156,1436,208,1856]" captionTargetId-0="figure@2.[150,1436,195,1913]" captionTargetId-1="figure@3.[168,1400,193,1861]" captionTargetId-2="figure@4.[151,1436,193,1865]" captionTargetId-3="figure@5.[151,1436,193,1885]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURES 1–2. Iniocyphus iheringi Raffray, 1912. lectotype. (1) Habitus habitus. (2) Labels. Scale bar for (1)." captionText-1="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." captionText-2="FIGURES 7–15. Iniocyphus iheringi Raffray, 1912. lectotype. (7) antenna. (8–11) abdomen (apical segment removed), in (8– 11) dorsal, (10) lateral and (11) ventral views. and pronotum, in (3) dorsal and (4) ventral views. (12) Anterior tibia and tarsus. (13) Middle tibia and tarsus. (14) Posterior tibia and tarsi. (15) Thorax, in ventrofrontal view. Abbreviations: blfIVbasolateral fovea of tergite IV, blfV—basolateral fovea of tergite V, blfVI—basolateral fovea of tergite VI." captionText-3="FIGURES 16–23. Iniocyphus iheringi Raffray, 1912. lectotype. (16) Aedeagus (internal sac everted), in ventral view. (17) Apical middle tarsus. (18) Left mandible. (19) Right mandible. (20) Mentum. (21) Labrum. (22) Abdominal sternite VIII. (23) Right maxillary palpus ventral views. Scale bars: vertical for (16–21, 23), and horizontal for (22)." httpUri-0="https://zenodo.org/record/2642623/files/figure.png" httpUri-1="https://zenodo.org/record/2642625/files/figure.png" httpUri-2="https://zenodo.org/record/2642627/files/figure.png" httpUri-3="https://zenodo.org/record/2642629/files/figure.png" pageId="1" pageNumber="172">Figs. 1–23</figureCitation>
)
</paragraph>
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<taxonomicName id="4C6D4D10FF97F433FF4EFB307EC1FB62" ID-CoL="8XLZH" authority="Raffray, 1912: 435" authorityName="Raffray" authorityPageNumber="435" authorityYear="1912" box="[151,467,1193,1215]" class="Insecta" family="Staphylinidae" genus="Iniocyphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="172" phylum="Arthropoda" rank="genus">
<emphasis id="B919EA81FF97F433FF4EFB307E14FB62" box="[151,262,1193,1215]" italics="true" pageId="1" pageNumber="172">Iniocyphus</emphasis>
<treatmentCitation id="0ACC1082FF97F433FED7FB307EC1FB62" author="Raffray, A." box="[270,467,1193,1215]" page="435" pageId="1" pageNumber="172" year="1912">
<bibRefCitation id="EFFC4B62FF97F433FED7FB307E8CFB62" author="Raffray, A." box="[270,414,1193,1215]" pageId="1" pageNumber="172" pagination="425 - 450" refId="ref3400" refString="Raffray, A. (1912) Especes nouvelles de Pselaphides exotiques [Coleopt.]. Annales de la Societe entomologique de France, 80 (1911), 425 - 450." type="journal article" year="1912">Raffray, 1912</bibRefCitation>
: 435
</treatmentCitation>
</taxonomicName>
; 1924: 144.
<treatmentCitation id="0ACC1082FF97F433FD8FFB307DEAFB62" author="Park, O." box="[598,760,1193,1215]" page="277" pageId="1" pageNumber="172" year="1942">
<bibRefCitation id="EFFC4B62FF97F433FD8FFB307DD1FB62" author="Park, O." box="[598,707,1193,1215]" pageId="1" pageNumber="172" pagination="1 - 403" refId="ref3284" refString="Park, O. (1942) A study in Neotropical Pselaphidae. Northwestern University studies in the biological sciences and medicine, 1, 1 - 403." type="journal article" year="1942">Park 1942</bibRefCitation>
: 277
</treatmentCitation>
;
<treatmentCitation id="0ACC1082FF97F433FCDDFB337B29FB62" author="Newton, A. F. &amp; Chandler, D. S." box="[772,1083,1193,1215]" page="50" pageId="1" pageNumber="172" year="1989">
<bibRefCitation id="EFFC4B62FF97F433FCDDFB337B01FB62" author="Newton, A. F. &amp; Chandler, D. S." box="[772,1043,1193,1215]" pageId="1" pageNumber="172" pagination="1 - 93" refId="ref3171" refString="Newton, A. F. &amp; Chandler, D. S. (1989) World catalog of the genera of Pselaphidae (Coleoptera). Fieldiana: Zoology, New Series, 53, 1 - 93." type="journal article" year="1989">Newton &amp; Chandler 1989</bibRefCitation>
: 50
</treatmentCitation>
;
<treatmentCitation id="0ACC1082FF97F433FB91FB307BE3FB62" author="Lobl, I." box="[1096,1265,1193,1215]" page="692" pageId="1" pageNumber="172" year="1994">
<bibRefCitation id="EFFC4B62FF97F433FB91FB307BAFFB62" author="Lobl, I." box="[1096,1213,1193,1215]" pageId="1" pageNumber="172" pagination="685 - 697" refId="ref3065" refString="Lobl, I. (1994) Awas giraffa gen. n., sp. n. (Coleoptera, Pselaphidae) from Malaysia and the classification of Goniacerinae. Revue suisse de zoologie, 101 (3), 685 - 697. https: // doi. org / 10.5962 / bhl. part. 79924" type="journal article" year="1994">Löbl, 1994</bibRefCitation>
: 692
</treatmentCitation>
.
<treatmentCitation id="0ACC1082FF97F433FB24FB307FE2FB03" author="Chandler, D. S." page="375" pageId="1" pageNumber="172" year="2001">
<bibRefCitation id="EFFC4B62FF97F433FB24FB307A8AFB62" author="Chandler, D. S." box="[1277,1432,1193,1215]" pageId="1" pageNumber="172" pagination="1 - 562" refId="ref2704" refString="Chandler, D. S. (2001) Biology, morphology, and systematics of the ant-like litter beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on entomology, International, 15, i-x + 1 - 562." type="journal article" year="2001">Chandler 2001</bibRefCitation>
: 375
</treatmentCitation>
.
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</paragraph>
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<emphasis id="B919EA81FF97F433FF4EFA937E20FAFE" bold="true" box="[151,306,1290,1315]" pageId="1" pageNumber="172">
<typeStatus id="54D68831FF97F433FF4EFA937FC0FAFE" box="[151,210,1290,1315]" pageId="1" pageNumber="172">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6D4D10FF97F433FEE0FA937DA5FAFE" authority="Raffray, 1912" authorityName="Raffray" authorityYear="1912" box="[313,695,1290,1315]" class="Insecta" family="Staphylinidae" genus="Iniocyphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="172" phylum="Arthropoda" rank="species" species="iheringi">
<emphasis id="B919EA81FF97F433FEE0FA937D06FAFE" box="[313,532,1290,1315]" italics="true" pageId="1" pageNumber="172">Iniocyphus iheringi</emphasis>
<bibRefCitation id="EFFC4B62FF97F433FDC3FA937DA5FAFE" author="Raffray, A." box="[538,695,1290,1315]" pageId="1" pageNumber="172" pagination="425 - 450" refId="ref3400" refString="Raffray, A. (1912) Especes nouvelles de Pselaphides exotiques [Coleopt.]. Annales de la Societe entomologique de France, 80 (1911), 425 - 450." type="journal article" year="1912">Raffray, 1912</bibRefCitation>
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, by monotypy.
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<paragraph id="8BD23693FF97F433FF1EFAB77DBBF99E" blockId="1.[151,1437,1290,2035]" pageId="1" pageNumber="172">
<emphasis id="B919EA81FF97F433FF1EFAB77E43FA9A" bold="true" box="[199,337,1326,1351]" pageId="1" pageNumber="172">Description</emphasis>
. Habitus as in
<figureCitation id="13562A16FF97F433FE2FFAB77D25FA9A" box="[502,567,1326,1351]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1937,1960]" captionTargetBox="[157,1408,212,1906]" captionTargetId="figure@2.[150,1436,195,1913]" captionTargetPageId="2" captionText="FIGURES 1–2. Iniocyphus iheringi Raffray, 1912. lectotype. (1) Habitus habitus. (2) Labels. Scale bar for (1)." httpUri="https://zenodo.org/record/2642623/files/figure.png" pageId="1" pageNumber="172">Fig. 1</figureCitation>
. Head with eyes well-developped (at least in male, female unknown), temples long, moderately convex; vertexal foveae very small, opening near upper eye margins (
<figureCitation id="13562A16FF97F433FBBEFACB7BC6FAB6" box="[1127,1236,1362,1387]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Figs. 3, 5</figureCitation>
: vf); postantennal notches narrow; ocular-mandibular carinae present; ventral surface covered with pointed microtubercles; gular area depressed at middle, with two gular foveae (
<figureCitation id="13562A16FF97F433FD52FA037DC0FA6E" box="[651,722,1434,1459]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Fig. 4</figureCitation>
: gf). Antennae (
<figureCitation id="13562A16FF97F433FC5EFA037CD8FA6E" box="[903,970,1434,1459]" captionStart="FIGURES 7–15" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[159,1423,210,1848]" captionTargetId="figure@4.[151,1436,193,1865]" captionTargetPageId="4" captionText="FIGURES 7–15. Iniocyphus iheringi Raffray, 1912. lectotype. (7) antenna. (8–11) abdomen (apical segment removed), in (8– 11) dorsal, (10) lateral and (11) ventral views. and pronotum, in (3) dorsal and (4) ventral views. (12) Anterior tibia and tarsus. (13) Middle tibia and tarsus. (14) Posterior tibia and tarsi. (15) Thorax, in ventrofrontal view. Abbreviations: blfIVbasolateral fovea of tergite IV, blfV—basolateral fovea of tergite V, blfVI—basolateral fovea of tergite VI." httpUri="https://zenodo.org/record/2642627/files/figure.png" pageId="1" pageNumber="172">Fig. 7</figureCitation>
) with eleven antennomeres, much longer than combined length of head and pronotum; club consisting of antennomeres IX–XI, with antennomeres IX wider than III–VIII, and X much longer than wide. Mouthparts as in
<figureCitation id="13562A16FF97F433FCA3FA7B7B20FA26" box="[890,1074,1506,1531]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1906,1929]" captionTargetBox="[156,1436,208,1856]" captionTargetId="figure@5.[151,1436,193,1885]" captionTargetPageId="5" captionText="FIGURES 16–23. Iniocyphus iheringi Raffray, 1912. lectotype. (16) Aedeagus (internal sac everted), in ventral view. (17) Apical middle tarsus. (18) Left mandible. (19) Right mandible. (20) Mentum. (21) Labrum. (22) Abdominal sternite VIII. (23) Right maxillary palpus ventral views. Scale bars: vertical for (16–21, 23), and horizontal for (22)." httpUri="https://zenodo.org/record/2642629/files/figure.png" pageId="1" pageNumber="172">Figs. 18–21, 23</figureCitation>
; maxillary palpi (
<figureCitation id="13562A16FF97F433FADAFA7B7A4AFA26" box="[1283,1368,1506,1531]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1906,1929]" captionTargetBox="[156,1436,208,1856]" captionTargetId="figure@5.[151,1436,193,1885]" captionTargetPageId="5" captionText="FIGURES 16–23. Iniocyphus iheringi Raffray, 1912. lectotype. (16) Aedeagus (internal sac everted), in ventral view. (17) Apical middle tarsus. (18) Left mandible. (19) Right mandible. (20) Mentum. (21) Labrum. (22) Abdominal sternite VIII. (23) Right maxillary palpus ventral views. Scale bars: vertical for (16–21, 23), and horizontal for (22)." httpUri="https://zenodo.org/record/2642629/files/figure.png" pageId="1" pageNumber="172">Fig. 23</figureCitation>
) with article 3 distinctly quandrangular, with mesal margin long, almost as long as external margin; no trichome at the outside of the article 4 (see
<bibRefCitation id="EFFC4B62FF97F433FE15F9B27D60F99E" author="Nomura, Sh." box="[460,626,1579,1603]" pageId="1" pageNumber="172" pagination="1 - 462" refId="ref3251" refString="Nomura, Sh. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1 - 462." type="journal article" year="1991">Nomura, 1991</bibRefCitation>
: 40).
</paragraph>
<paragraph id="8BD23693FF97F433FF1EF9D67CBAF82E" blockId="1.[151,1437,1290,2035]" pageId="1" pageNumber="172">
Pronotum fairly hexagonal, sides slightly sinuate posteriorly; median antebasal fovea present (
<figureCitation id="13562A16FF97F433FAD1F9D77A43F9BA" box="[1288,1361,1614,1639]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Fig. 3</figureCitation>
: maf); antebasal sulcus present (
<figureCitation id="13562A16FF97F433FE6FF9EB7EEEF956" box="[438,508,1650,1675]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Fig. 3</figureCitation>
: as), joining the lateral foveae (
<figureCitation id="13562A16FF97F433FCBBF9EB7CBAF956" box="[866,936,1650,1675]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Fig. 3</figureCitation>
: laf); areas posterior to antebasal sulcus and near most protruding portion of lateral edges covered with pointed microtubercles. Prosternum lacking prosternal foveae; paranotal carinae present, entire (
<figureCitation id="13562A16FF97F433FDB3F9237DB8F90E" box="[618,682,1722,1747]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Fig 4</figureCitation>
: pc); center covered with pointed microtubercles. Elytra very long, longer than their maximal width combined, and two times longer than abdomen in dorsal view, without foveae, sulci or carinae; with small concavity at posterolateral angles (as one of locking devices
<emphasis id="B919EA81FF97F433FB5BF89D7BD2F8C6" box="[1154,1216,1796,1819]" italics="true" pageId="1" pageNumber="172">sensu</emphasis>
<bibRefCitation id="EFFC4B62FF97F433FB1EF89A7A7FF8C6" author="Nomura, Sh." box="[1223,1389,1795,1819]" pageId="1" pageNumber="172" pagination="1 - 462" refId="ref3251" refString="Nomura, Sh. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1 - 462." type="journal article" year="1991">Nomura, 1991</bibRefCitation>
: 16, Figs. 28–29). Wings well-developped. Mesoventrite with pairs of median mesoventral foveae (
<figureCitation id="13562A16FF97F433FB36F8BF7A2BF8E2" box="[1263,1337,1830,1855]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Fig. 6</figureCitation>
: mmvf), lateral mesoventral foveae (
<figureCitation id="13562A16FF97F433FE15F8D37D03F8BE" box="[460,529,1866,1891]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Fig. 6</figureCitation>
: lmvf), longitudinal anapleural carinae (
<emphasis id="B919EA81FF97F433FC0BF8D57B02F8BE" box="[978,1040,1868,1891]" italics="true" pageId="1" pageNumber="172">sensu</emphasis>
<bibRefCitation id="EFFC4B62FF97F433FBCEF8D27BAEF8BE" author="Nomura, Sh." box="[1047,1212,1867,1891]" pageId="1" pageNumber="172" pagination="1 - 462" refId="ref3251" refString="Nomura, Sh. (1991) Systematic study on the genus Batrisoplisus and its allied genera from Japan (Coleoptera, Pselaphidae). Esakia, 30, 1 - 462." type="journal article" year="1991">Nomura, 1991</bibRefCitation>
: 12, fig. 18a;
<figureCitation id="13562A16FF97F433FA8AF8D37A85F8BE" box="[1363,1431,1866,1891]" captionStart="FIGURES 3–6" captionStartId="3.[151,264,1891,1914]" captionTargetBox="[168,1400,193,1861]" captionTargetId="figure@3.[168,1400,193,1861]" captionTargetPageId="3" captionText="FIGURES 3–6. Iniocyphus iheringi Raffray, 1912. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea." httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="1" pageNumber="172">Fig. 6</figureCitation>
: lapc). Metaventrite apparently with pair of small lateral mesocoxal foveae (not clearly visible without preparation); posterior edge between metacoxae in dorsal view evenly concave. Legs (
<figureCitation id="13562A16FF97F433FBDBF80B7B9DF876" box="[1026,1167,1938,1963]" captionStart="FIGURES 7–15" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[159,1423,210,1848]" captionTargetId="figure@4.[151,1436,193,1865]" captionTargetPageId="4" captionText="FIGURES 7–15. Iniocyphus iheringi Raffray, 1912. lectotype. (7) antenna. (8–11) abdomen (apical segment removed), in (8– 11) dorsal, (10) lateral and (11) ventral views. and pronotum, in (3) dorsal and (4) ventral views. (12) Anterior tibia and tarsus. (13) Middle tibia and tarsus. (14) Posterior tibia and tarsi. (15) Thorax, in ventrofrontal view. Abbreviations: blfIVbasolateral fovea of tergite IV, blfV—basolateral fovea of tergite V, blfVI—basolateral fovea of tergite VI." httpUri="https://zenodo.org/record/2642627/files/figure.png" pageId="1" pageNumber="172">Figs. 12–13</figureCitation>
) rather long; pro- and mesocoxae conical, contiguous, oblique (
<figureCitation id="13562A16FF97F433FDADF82F7DDEF812" box="[628,716,1974,1999]" captionStart="FIGURES 7–15" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[159,1423,210,1848]" captionTargetId="figure@4.[151,1436,193,1865]" captionTargetPageId="4" captionText="FIGURES 7–15. Iniocyphus iheringi Raffray, 1912. lectotype. (7) antenna. (8–11) abdomen (apical segment removed), in (8– 11) dorsal, (10) lateral and (11) ventral views. and pronotum, in (3) dorsal and (4) ventral views. (12) Anterior tibia and tarsus. (13) Middle tibia and tarsus. (14) Posterior tibia and tarsi. (15) Thorax, in ventrofrontal view. Abbreviations: blfIVbasolateral fovea of tergite IV, blfV—basolateral fovea of tergite V, blfVI—basolateral fovea of tergite VI." httpUri="https://zenodo.org/record/2642627/files/figure.png" pageId="1" pageNumber="172">Fig. 15</figureCitation>
); metacoxae moderately distants (
<figureCitation id="13562A16FF97F433FB80F82F7BA3F812" box="[1113,1201,1974,1999]" captionStart="FIGURES 7–15" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[159,1423,210,1848]" captionTargetId="figure@4.[151,1436,193,1865]" captionTargetPageId="4" captionText="FIGURES 7–15. Iniocyphus iheringi Raffray, 1912. lectotype. (7) antenna. (8–11) abdomen (apical segment removed), in (8– 11) dorsal, (10) lateral and (11) ventral views. and pronotum, in (3) dorsal and (4) ventral views. (12) Anterior tibia and tarsus. (13) Middle tibia and tarsus. (14) Posterior tibia and tarsi. (15) Thorax, in ventrofrontal view. Abbreviations: blfIVbasolateral fovea of tergite IV, blfV—basolateral fovea of tergite V, blfVI—basolateral fovea of tergite VI." httpUri="https://zenodo.org/record/2642627/files/figure.png" pageId="1" pageNumber="172">Fig. 15</figureCitation>
); tarsomeres 1 very short, tarsomeres 2 slightly shorter than 3, latter with 1 claw (
<figureCitation id="13562A16FF97F433FC9EF8437C89F82E" box="[839,923,2010,2035]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1906,1929]" captionTargetBox="[156,1436,208,1856]" captionTargetId="figure@5.[151,1436,193,1885]" captionTargetPageId="5" captionText="FIGURES 16–23. Iniocyphus iheringi Raffray, 1912. lectotype. (16) Aedeagus (internal sac everted), in ventral view. (17) Apical middle tarsus. (18) Left mandible. (19) Right mandible. (20) Mentum. (21) Labrum. (22) Abdominal sternite VIII. (23) Right maxillary palpus ventral views. Scale bars: vertical for (16–21, 23), and horizontal for (22)." httpUri="https://zenodo.org/record/2642629/files/figure.png" pageId="1" pageNumber="172">Fig. 17</figureCitation>
).
</paragraph>
<caption id="DF12661BFF94F430FF4EF8087BE8F875" box="[151,1274,1937,1960]" httpUri="https://zenodo.org/record/2642623/files/figure.png" pageId="2" pageNumber="173" startId="2.[151,264,1937,1960]" targetBox="[157,1408,212,1906]" targetPageId="2">
<paragraph id="8BD23693FF94F430FF4EF8087BE8F875" blockId="2.[151,1274,1937,1960]" box="[151,1274,1937,1960]" pageId="2" pageNumber="173">
<emphasis id="B919EA81FF94F430FF4EF8087E28F87A" bold="true" box="[151,314,1937,1960]" pageId="2" pageNumber="173">FIGURES 1–2.</emphasis>
<taxonomicName id="4C6D4D10FF94F430FE99F80B7D88F875" authority="Raffray, 1912" authorityName="Raffray" authorityYear="1912" box="[320,666,1938,1960]" class="Insecta" family="Staphylinidae" genus="Iniocyphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="173" phylum="Arthropoda" rank="species" species="iheringi">
<emphasis id="B919EA81FF94F430FE99F80B7D14F875" box="[320,518,1938,1960]" italics="true" pageId="2" pageNumber="173">Iniocyphus iheringi</emphasis>
Raffray, 1912
</taxonomicName>
. lectotype. (1) Habitus habitus. (2) Labels. Scale bar for (1).
</paragraph>
</caption>
<caption id="DF12661BFF95F431FF4EF8FA7D93F829" httpUri="https://zenodo.org/record/2642625/files/figure.png" pageId="3" pageNumber="174" startId="3.[151,264,1891,1914]" targetBox="[168,1400,193,1861]" targetPageId="3">
<paragraph id="8BD23693FF95F431FF4EF8FA7D93F829" blockId="3.[151,1436,1891,2036]" pageId="3" pageNumber="174">
<emphasis id="B919EA81FF95F431FF4EF8FA7E29F8A4" bold="true" box="[151,315,1891,1914]" pageId="3" pageNumber="174">FIGURES 3–6.</emphasis>
<taxonomicName id="4C6D4D10FF95F431FE9DF8FD7DB0F8A7" authority="Raffray, 1912" authorityName="Raffray" authorityYear="1912" box="[324,674,1892,1914]" class="Insecta" family="Staphylinidae" genus="Iniocyphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="174" phylum="Arthropoda" rank="species" species="iheringi">
<emphasis id="B919EA81FF95F431FE9DF8FD7D19F8A7" box="[324,523,1892,1914]" italics="true" pageId="3" pageNumber="174">Iniocyphus iheringi</emphasis>
Raffray, 1912
</taxonomicName>
. lectotype. (3–4) Head (mouthparts removed) and pronotum, in (3) dorsal and (4) ventral views. (5–6) Head (mouthparts removed) and thorax, in (5) lateral and (6) ventral views. Scale bar 200 µm for (3–4), and scale bar 1 mm for (5–6). Abbreviations: as—antebasal sulcus, gf—gular fovea, laf—lateral antebasal fovea, lapclongitudinal anapleural carina, lmvf—lateral mesoventral fovea, maf—median antebasal fovea, mmvf—median mesoventral fovea, pc—paranotal carina. vf—vertexal fovea.
</paragraph>
</caption>
<caption id="DF12661BFF92F436FF4EF8C77BDFF80D" httpUri="https://zenodo.org/record/2642627/files/figure.png" pageId="4" pageNumber="175" startId="4.[151,264,1886,1909]" targetBox="[159,1423,210,1848]" targetPageId="4">
<paragraph id="8BD23693FF92F436FF4EF8C77BDFF80D" blockId="4.[151,1437,1886,2000]" pageId="4" pageNumber="175">
<emphasis id="B919EA81FF92F436FF4EF8C77E55F8A9" bold="true" box="[151,327,1886,1909]" pageId="4" pageNumber="175">FIGURES 7–15.</emphasis>
<taxonomicName id="4C6D4D10FF92F436FE97F8C77DBBF8A9" authority="Raffray, 1912" authorityName="Raffray" authorityYear="1912" box="[334,681,1886,1908]" class="Insecta" family="Staphylinidae" genus="Iniocyphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="175" phylum="Arthropoda" rank="species" species="iheringi">
<emphasis id="B919EA81FF92F436FE97F8C77D06F8A9" box="[334,532,1886,1908]" italics="true" pageId="4" pageNumber="175">Iniocyphus iheringi</emphasis>
Raffray, 1912
</taxonomicName>
. lectotype. (7) antenna. (8–11) abdomen (apical segment removed), in (8– 11) dorsal, (10) lateral and (11) ventral views. and pronotum, in (3) dorsal and (4) ventral views. (12) Anterior tibia and tarsus. (13) Middle tibia and tarsus. (14) Posterior tibia and tarsi. (15) Thorax, in ventrofrontal view. Abbreviations: blfIVbasolateral fovea of tergite IV, blfV—basolateral fovea of tergite V, blfVI—basolateral fovea of tergite VI.
</paragraph>
</caption>
<caption id="DF12661BFF93F437FF4EF8EB7B61F81B" httpUri="https://zenodo.org/record/2642629/files/figure.png" pageId="5" pageNumber="176" startId="5.[151,264,1906,1929]" targetBox="[156,1436,208,1856]" targetPageId="5">
<paragraph id="8BD23693FF93F437FF4EF8EB7B61F81B" blockId="5.[151,1436,1906,1990]" pageId="5" pageNumber="176">
<emphasis id="B919EA81FF93F437FF4EF8EB7E44F855" bold="true" box="[151,342,1906,1929]" pageId="5" pageNumber="176">FIGURES 16–23.</emphasis>
<taxonomicName id="4C6D4D10FF93F437FE86F8EA7DD3F854" authority="Raffray, 1912" authorityName="Raffray" authorityYear="1912" box="[351,705,1907,1929]" class="Insecta" family="Staphylinidae" genus="Iniocyphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="176" phylum="Arthropoda" rank="species" species="iheringi">
<emphasis id="B919EA81FF93F437FE86F8EA7D3AF854" box="[351,552,1907,1929]" italics="true" pageId="5" pageNumber="176">Iniocyphus iheringi</emphasis>
Raffray, 1912
</taxonomicName>
. lectotype. (16) Aedeagus (internal sac everted), in ventral view. (17) Apical middle tarsus. (18) Left mandible. (19) Right mandible. (20) Mentum. (21) Labrum. (22) Abdominal sternite VIII. (23) Right maxillary palpus ventral views. Scale bars: vertical for (16–21, 23), and horizontal for (22).
</paragraph>
</caption>
<paragraph id="8BD23693FF90F434FF1EFF0E7E71FEC0" blockId="6.[151,1436,151,680]" pageId="6" pageNumber="177">
Abdomen with five visibles tergites (IV–VIII) and six visibles sternites (III–VIII); tergites IV–VI each with pair of small basolateral foveae (
<figureCitation id="13562A16FF90F434FDC9FF257D4AFF09" box="[528,600,188,213]" captionStart="FIGURES 7–15" captionStartId="4.[151,264,1886,1909]" captionTargetBox="[159,1423,210,1848]" captionTargetId="figure@4.[151,1436,193,1865]" captionTargetPageId="4" captionText="FIGURES 7–15. Iniocyphus iheringi Raffray, 1912. lectotype. (7) antenna. (8–11) abdomen (apical segment removed), in (8– 11) dorsal, (10) lateral and (11) ventral views. and pronotum, in (3) dorsal and (4) ventral views. (12) Anterior tibia and tarsus. (13) Middle tibia and tarsus. (14) Posterior tibia and tarsi. (15) Thorax, in ventrofrontal view. Abbreviations: blfIVbasolateral fovea of tergite IV, blfV—basolateral fovea of tergite V, blfVI—basolateral fovea of tergite VI." httpUri="https://zenodo.org/record/2642627/files/figure.png" pageId="6" pageNumber="177">Fig. 9</figureCitation>
: blfIV, blfV and blfVI); sternite III with apical margin visible on entire length, including posteriorly metacoxae; sternite IV with pair of basolateral foveae; sternite VIII (
<figureCitation id="13562A16FF90F434FADDFF467A4AFF25" box="[1284,1368,223,248]" captionStart="FIGURES 16–23" captionStartId="5.[151,264,1906,1929]" captionTargetBox="[156,1436,208,1856]" captionTargetId="figure@5.[151,1436,193,1885]" captionTargetPageId="5" captionText="FIGURES 16–23. Iniocyphus iheringi Raffray, 1912. lectotype. (16) Aedeagus (internal sac everted), in ventral view. (17) Apical middle tarsus. (18) Left mandible. (19) Right mandible. (20) Mentum. (21) Labrum. (22) Abdominal sternite VIII. (23) Right maxillary palpus ventral views. Scale bars: vertical for (16–21, 23), and horizontal for (22)." httpUri="https://zenodo.org/record/2642629/files/figure.png" pageId="6" pageNumber="177">Fig. 22</figureCitation>
) with mediobasal gland.
</paragraph>
<paragraph id="8BD23693FF90F434FF1EFEB17CC7FEB8" blockId="6.[151,1436,151,680]" pageId="6" pageNumber="177">Upper part of head and mesotrochanters apparently sexually modified. Aedeagus membranous, with 2 parameres fused at base and affixed slightly asymetrically to median lobe.</paragraph>
</subSubSection>
<subSubSection id="C3776518FF90F434FF1EFEF67A24FD75" pageId="6" pageNumber="177" type="discussion">
<paragraph id="8BD23693FF90F434FF1EFEF67A24FD75" blockId="6.[151,1436,151,680]" pageId="6" pageNumber="177">
<emphasis id="B919EA81FF90F434FF1EFEF67E5DFE55" bold="true" box="[199,335,367,392]" pageId="6" pageNumber="177">Comments.</emphasis>
With respect to the various genera currently assigned to Iniocyphini, Proterini, and Brachyglutini, the genus
<taxonomicName id="4C6D4D10FF90F434FED3FE0D7E97FE70" authorityName="Raffray" authorityYear="1912" box="[266,389,404,429]" class="Insecta" family="Staphylinidae" genus="Iniocyphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B919EA81FF90F434FED3FE0D7E97FE70" box="[266,389,404,429]" italics="true" pageId="6" pageNumber="177">Iniocyphus</emphasis>
</taxonomicName>
is characterized by the following unique features: 1) Presence of pointed microtubercles on pronotum, prosternum and ventral surface of head; 2) Elytra unusually long, longer than their maximal width combined; 3) Mesocoxae conical and oblique. Within these three tribes the presence of markedly elongate antennomeres X is quite rare and occurs only here and there, such as for example in
<taxonomicName id="4C6D4D10FF90F434FB82FE667A8EFDC5" authority="Chandler, 2001" authorityName="Chandler" authorityYear="2001" box="[1115,1436,511,536]" class="Insecta" family="Staphylinidae" genus="Simplicorfa" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B919EA81FF90F434FB82FE667BCCFDC5" box="[1115,1246,511,536]" italics="true" pageId="6" pageNumber="177">Simplicorfa</emphasis>
<bibRefCitation id="EFFC4B62FF90F434FB3FFE667A8EFDC5" author="Chandler, D. S." box="[1254,1436,511,536]" pageId="6" pageNumber="177" pagination="1 - 562" refId="ref2704" refString="Chandler, D. S. (2001) Biology, morphology, and systematics of the ant-like litter beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on entomology, International, 15, i-x + 1 - 562." type="journal article" year="2001">Chandler, 2001</bibRefCitation>
</taxonomicName>
(Inyociphini: Natypleurina), and males of
<taxonomicName id="4C6D4D10FF90F434FD5CFDBC7BD9FDE0" authority="Cuccodoro &amp; Kurbatov, 2006" authorityName="Cuccodoro &amp; Kurbatov" authorityYear="2006" box="[645,1227,548,573]" class="Insecta" family="Staphylinidae" genus="Exoterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="177" phylum="Arthropoda" rank="genus">
<emphasis id="B919EA81FF90F434FD5CFDBC7DFBFDE1" box="[645,745,549,572]" italics="true" pageId="6" pageNumber="177">Exoterus</emphasis>
<bibRefCitation id="EFFC4B62FF90F434FD2AFDBD7B5BFDE0" author="Cuccodoro, G. &amp; Kurbatov, S. A." box="[755,1097,548,573]" pageId="6" pageNumber="177" pagination="251 - 273" refId="ref2748" refString="Cuccodoro, G. &amp; Kurbatov, S. A. (2006) Revision of Proterus Raffray, 1897, with description of a new affiliated genus from Thailand (Coleoptera, Staphylinidae, Pselaphinae). Mitteilungen der schweizerischen entomologischen Gesellschaft, 79, 251 - 273." type="journal article" year="2006">Cuccodoro &amp; Kurbatov, 2006</bibRefCitation>
(Proterini)
</taxonomicName>
. Also notable and potentially of high significance in further reassessments of phylogenetic position of the genus in the system are the quandrangular shape of the third segment of the maxillary palpi, the lack of trichome at the outside of the fourth segment of the maxillary palpi, and the apical margin of abdominal sternite III visible on its entire length.
</paragraph>
</subSubSection>
</treatment>
</document>
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