treatments-xml/data/8C/D7/BE/8CD7BE05252F53729494CDA75FA4DA80.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1004.57526" ID-GBIF-Dataset="4e633bc1-16fc-4e83-b8a0-cb9289897616" ID-PMC="PMC7758311" ID-Pensoft-Pub="1313-2970-1004-27" ID-Pensoft-UUID="BEECC18115E95A798CA5250990A350E3" ID-PubMed="33384565" ID-ZooBank="320559CF19B5423CB7FB72555290241A" ModsDocID="1313-2970-1004-27" checkinTime="1608157838683" checkinUser="pensoft" docAuthor="Maddison, Wayne P., Beattie, Imara, Marathe, Kiran, Ng, Paul Y. C., Kanesharatnam, Nilani, Benjamin, Suresh P. &amp; Kunte, Krushnamegh" docDate="2020" docId="8CD7BE05252F53729494CDA75FA4DA80" docLanguage="en" docName="ZooKeys 1004: 27-97" docOrigin="ZooKeys 1004" docPubDate="2020-12-16" docSource="http://dx.doi.org/10.3897/zookeys.1004.57526" docTitle="Maripanthus draconis Maddison, sp. nov." docType="treatment" docVersion="4" id="BEECC18115E95A798CA5250990A350E3" lastPageNumber="27" masterDocId="BEECC18115E95A798CA5250990A350E3" masterDocTitle="A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)" masterLastPageNumber="97" masterPageNumber="27" pageNumber="27" updateTime="1668146624181" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>A phylogenetic and taxonomic review of baviine jumping spiders (Araneae, Salticidae, Baviini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Maddison, Wayne P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4953-4575</mods:nameIdentifier>
<mods:affiliation>Departments of Zoology and Botany and Beaty Biodiversity Museum, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
<mods:nameIdentifier type="email">wmaddisn@mail.ubc.ca</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Beattie, Imara</mods:namePart>
<mods:affiliation>Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marathe, Kiran</mods:namePart>
<mods:affiliation>Department of Zoology, University of British Columbia, 6270 University Boulevard, Vancouver, British Columbia, V 6 T 1 Z 4, Canada &amp; National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Ng, Paul Y. C.</mods:namePart>
<mods:affiliation>205 River Valley Road, # 16 - 53, Singapore 238274, Republic of Singapore</mods:affiliation>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kanesharatnam, Nilani</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6070-8989</mods:nameIdentifier>
<mods:affiliation>National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka &amp; Department of Zoology, Faculty of Science, Eastern University, Vantharumoolai, Sri Lanka</mods:affiliation>
</mods:name>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Benjamin, Suresh P.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4666-0330</mods:nameIdentifier>
<mods:affiliation>National Institute of Fundamental Studies, Hantana Road, Kandy, Sri Lanka</mods:affiliation>
</mods:name>
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<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kunte, Krushnamegh</mods:namePart>
<mods:affiliation>National Centre for Biological Sciences, Tata Institute of Fundamental Research, GKVK Campus, Bellary Road, Bengaluru 560065, India</mods:affiliation>
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<mods:date>2020</mods:date>
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<mods:number>2020-12-16</mods:number>
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<mods:number>1004</mods:number>
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<treatment ID-GBIF-Taxon="176088103" LSID="urn:lsid:zoobank.org:act:B511C74F-1343-48B8-ACDD-B01B20549324" httpUri="http://treatment.plazi.org/id/8CD7BE05252F53729494CDA75FA4DA80" lastPageNumber="27" pageId="0" pageNumber="27">
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<paragraph pageId="0" pageNumber="27">
<taxonomicName LSID="http://zoobank.org/B511C74F-1343-48B8-ACDD-B01B20549324" authority="Maddison, sp. nov." authorityName="Maddison, sp. nov." family="Pittosporaceae" genus="Maripanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maripanthus draconis" order="Apiales" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="draconis">Maripanthus draconis Maddison, sp. nov.</taxonomicName>
<figureCitation captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">Figs 27</figureCitation>
<figureCitation captionStart="Figures 4–35" captionStartId="F2" captionText="Figures 4 - 35. Variation in traits among baviines 4, 5 oblique views of female prosoma 4 Bavia nessagyna (specimen IDWM. 20004; note convex front face of chelicerae, and typical thorax) 5 Indopadilla redunca (holotype IDWM. 20011; note concave front face of chelicerae, and thoracic bulges) 6 - 12 left male endite (9 also with chelicera) 6 Bavia nessagyna (holotype IDWM. 20005) 7 Indopadilla redunca (specimen SWK 12 - M 0009) 8 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 9 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 10 Padillothorax badut (specimen SWK 12 - 4688) 11 Maripanthus jubatus (specimen AS 19.4373) 12 Piranthus planolancis (specimen AS 19.5970) 13 - 20 carapace (all females except 15) 13 Bavia nessagyna (specimen IDWM. 20004) 14 Indopadilla redunca (holotype IDWM. 20011) 15 Stagetillus opaciceps male (specimen JK. 13.02.26.0017) 16 Stagetillus cf. opaciceps female (specimen MRB 079). 17 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 18 Padillothorax mulu (from Mulu Nat. Pk.) 19 Maripanthus reinholdae (specimen SWK 12 - 1934) 20 Piranthus planolancis (specimen AS 19.5940) 21 - 28 Metatarsus of first leg, retrolateral view. All female except 24 male. All are of left legs except 23, 25, 28 which are of right leg, digitally flipped 21 Bavia nessagyna (specimen IDWM. 20004) 22 Indopadilla kahariana (specimen SWK 12 - 1876) 23 Stagetillus cf. opaciceps (specimen MRB 079) 24 Padillothorax semiostrinus (specimen SWK 12 - EP 0105) 25 Padillothorax mulu (specimen SWK 12 - EP 0105) 26 Maripanthus reinholdae (specimen SWK 12 - 1934) 27 Maripanthus draconis (from Gunung Belemut, Johor) 28 Piranthus planolancis (specimen AS 19.5940) 29 - 35 prolateral surface of male first leg femur. 29 Bavia cf. capistrata (specimen from Singapore) 30 Indopadilla kahariana (specimen from Lambir Hills Nat. Pk.) 31 Stagetillus opaciceps (specimen JK. 08.08.19.0001) 32 Padillothorax semiostrinus (specimen JK. 20.06.20.001) 33 Padillothorax badut (specimen SWK 12 - 4688) 34 Maripanthus draconis (specimen from Johor, Gunung Lambak) 35 Piranthus bakau (holotype). Scale bars: 0.1 mm (6 - 12, 21 - 35), 1.0 mm (4, 5, 13 - 20)." figureDoi="10.3897/zookeys.1004.57526.figures4-35" httpUri="https://binary.pensoft.net/fig/487533" pageId="0" pageNumber="27">, 34</figureCitation>
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">, 188-201</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="type material">
<paragraph pageId="0" pageNumber="27">Type material.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Holotype</emphasis>
male (specimen AS19.2232), two 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">paratype</emphasis>
females (specimens AS19.2250 and d547), all in LKCNHM, from Singapore: Bukit Timah Nature Reserve, stream at Jungle Falls Path. 
<geoCoordinate degrees="1.3562" direction="north" orientation="latitude" precision="5" value="1.3562">1.3562°N</geoCoordinate>
, 
<geoCoordinate degrees="103.7748" direction="east" orientation="longitude" precision="5" value="103.7748">103.7748°E</geoCoordinate>
to 
<geoCoordinate degrees="1.3572" direction="north" orientation="latitude" precision="5" value="1.3572">1.3572°N</geoCoordinate>
, 
<geoCoordinate degrees="103.7734" direction="east" orientation="longitude" precision="5" value="103.7734">103.7734°E</geoCoordinate>
110-150 m elev. 12 June 2019 Maddison, Morehouse, &amp; Marathe WPM#19-051. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Paratype</emphasis>
male (specimen WSG018) from Singapore: Nee Soon Swamp Forest. Beating vegetation. 
<geoCoordinate degrees="1.39" direction="north" orientation="latitude" precision="555" value="1.39">1.39°N</geoCoordinate>
, 
<geoCoordinate degrees="103.81" direction="east" orientation="longitude" precision="555" value="103.81">103.81°E</geoCoordinate>
12 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-015.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="etymology">
<paragraph pageId="0" pageNumber="27">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="27">
Greek, 
<emphasis italics="true" pageId="0" pageNumber="27">
<normalizedToken originalValue="δράκων">δράκων</normalizedToken>
</emphasis>
, referring to the fiery colours of the male. 
<emphasis italics="true" pageId="0" pageNumber="27">Other names</emphasis>
: In 
<bibRefCitation author="Maddison, WP" journalOrPublisher="Journal of Arachnology" pageId="0" pageNumber="27" publicationUrl="http://salticidae.org/salticidImages/" refId="B23" refString="Maddison, WP, 2015b. Salticid images. http://salticidae.org/salticidImages/" title="Salticid images." url="http://salticidae.org/salticidImages/" year="2015 b">Maddison (2015b)</bibRefCitation>
and 
<normalizedToken originalValue="WPM’s">WPM's</normalizedToken>
field or lab notebooks the informal code for this species was 
<normalizedToken originalValue="“CFMAR”">&quot;CFMAR&quot;</normalizedToken>
. A specimen that is likely a closely related but distinct species is shown in 
<bibRefCitation author="Koh, JKH" journalOrPublisher="Sabah Forestry Department, city" pageId="0" pageNumber="27" refId="B18" refString="Koh, JKH, Bay, N, 2019. Borneo Spiders: A photographic field guide. Sabah Forestry Department, city" title="Borneo Spiders: A photographic field guide." year="2019">Koh and Bay 2019</bibRefCitation>
as &quot; 
<taxonomicName authorityName="Simon" authorityYear="1877" class="Arachnida" family="Pittosporaceae" genus="Bavia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bavia" order="Araneae" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Bavia</emphasis>
</taxonomicName>
&quot; sp. B Black-collared long-bellied jumping spider.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="diagnosis">
<paragraph pageId="0" pageNumber="27">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="27">
Most similar to 
<taxonomicName lsidName="M. smedleyi" pageId="0" pageNumber="27" rank="species" species="smedleyi">
<emphasis italics="true" pageId="0" pageNumber="27">M. smedleyi</emphasis>
</taxonomicName>
, of which only the female is known; 
<taxonomicName lsidName="M. draconis" pageId="0" pageNumber="27" rank="species" species="draconis">
<emphasis italics="true" pageId="0" pageNumber="27">M. draconis</emphasis>
</taxonomicName>
differs in having longer epigynal atria (greater than half of epigynal length) and less distinct atrial cliff (
<normalizedToken originalValue="“ac”">&quot;ac&quot;</normalizedToken>
, Fig. 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">191</figureCitation>
). Similar to 
<taxonomicName lsidName="M. jubatus" pageId="0" pageNumber="27" rank="species" species="jubatus">
<emphasis italics="true" pageId="0" pageNumber="27">M. jubatus</emphasis>
</taxonomicName>
in being large bodied, but 
<taxonomicName lsidName="M. draconis" pageId="0" pageNumber="27" rank="species" species="draconis">
<emphasis italics="true" pageId="0" pageNumber="27">M. draconis</emphasis>
</taxonomicName>
differs in having a honey-coloured translucent thorax, which in the female is speckled with black, and which in the male contrasts strongly against the black ocular area. Male front is impressively red-orange, with red-orange first femora, palps, and rings around eyes, and a golden face. Embolus slightly shorter than the other three species for which males are known. Ventral bump on palp femur.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="description">
<paragraph pageId="0" pageNumber="27">Description.</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Male</emphasis>
(based on holotype, specimen AS19.2232). Carapace length 4.0; abdomen length 5.0. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Carapace</emphasis>
(Figs 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">196</figureCitation>
, 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">197</figureCitation>
) black in ocular area and in a thin line along ventral and posterior margin; thorax strongly contrasting honey coloured, glabrous except for a few vertical black setae. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Clypeus</emphasis>
covered with yellow-cream setae. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Chelicerae</emphasis>
vertical, a soft orange-red, covered with yellow-cream setae. Three promarginal and five retromarginal teeth. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Palp</emphasis>
femur pale, darkening to red-brown distally. Bulb elongate (Figs 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">188</figureCitation>
, 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">189</figureCitation>
); embolus arising on the prolateral basal corner and proceeding distally. Femur with small but distinct distal ventral bump. Endite with small corner projection. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Legs</emphasis>
with pale femora and darker markings more distally. First femur striking orange-red, patella to tarsus dark brown to black. First patella through metatarsus with sparse ventral fringe of setae that are black at base, white at tip. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Abdomen</emphasis>
with brown longitudinal medial band dorsally, flanked by scattered red scales, with some scattered white scales basally.
</paragraph>
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Female</emphasis>
(paratype, specimen AS19.2250). Carapace length 4.2; abdomen length 6.1. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Carapace</emphasis>
(Fig. 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">199</figureCitation>
) integument black in ocular area, except for a pale spot on each side just medial to the PME (in other females, these pale spots form a transverse band that intersects with a longitudinal medial pale band, forming a pale cross in the ocular area). Thorax honey-coloured with distinct black speckles. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Clypeus</emphasis>
with scattered white setae. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Chelicerae</emphasis>
brown with darker patches, with scattered white setae. Three promarginal and five retromarginal teeth. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Legs</emphasis>
pale honey-coloured with various darker patches. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Abdomen</emphasis>
with central longitudinal pale chevroned band flanked by red scales. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="27">Epigyne</emphasis>
with large atria leading to broad ducts (Figs 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">191</figureCitation>
, 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">192</figureCitation>
). Atria shallow laterally, declining rapidly at an &quot;atrial cliff&quot; (
<normalizedToken originalValue="“ac”">&quot;ac&quot;</normalizedToken>
, Fig. 
<figureCitation captionStart="Figures 188–201" captionStartId="F21" captionText="Figures 188 - 201. Maripanthus draconis sp. nov. 188 male left palp, prolateral view (holotype AS 19.2232) 189 same, ventral view 190 same, retrolateral view 191 epigyne, ventral (specimen AS 19.2250) 192 vulva, dorsal 193 - 197 holotype 198 - 201 female AS 19.2250. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27">191</figureCitation>
) medial to which the surface is distinctly deeper. The ECP is apparently medial and small.
</paragraph>
<caption doi="10.3897/zookeys.1004.57526.figures188-201" httpUri="https://binary.pensoft.net/fig/487552" pageId="0" pageNumber="27" start="Figures 188–201" startId="F21">
<paragraph pageId="0" pageNumber="27">
<emphasis bold="true" pageId="0" pageNumber="27">Figures 188-201.</emphasis>
<taxonomicName family="Pittosporaceae" genus="Maripanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maripanthus draconis" order="Apiales" pageId="0" pageNumber="27" phylum="Arthropoda" rank="species" species="draconis">
<emphasis italics="true" pageId="0" pageNumber="27">Maripanthus draconis</emphasis>
</taxonomicName>
sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="27">188</emphasis>
male left palp, prolateral view (holotype AS19.2232) 
<emphasis bold="true" pageId="0" pageNumber="27">189</emphasis>
same, ventral view 
<emphasis bold="true" pageId="0" pageNumber="27">190</emphasis>
same, retrolateral view 
<emphasis bold="true" pageId="0" pageNumber="27">191</emphasis>
epigyne, ventral (specimen AS19.2250) 
<emphasis bold="true" pageId="0" pageNumber="27">192</emphasis>
vulva, dorsal 
<emphasis bold="true" pageId="0" pageNumber="27">193-197</emphasis>
holotype 
<emphasis bold="true" pageId="0" pageNumber="27">198-201</emphasis>
female AS19.2250. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="natural history">
<paragraph pageId="0" pageNumber="27">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="27">In Singapore, beating vegetation in forest understory.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="27" type="materials_examined">
<paragraph pageId="0" pageNumber="27">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="27">
In UBCZ. Singapore: Bukit Timah Nature Reserve, stream at Jungle Falls Path. 
<geoCoordinate degrees="1.3562" direction="north" orientation="latitude" precision="5" value="1.3562">1.3562°N</geoCoordinate>
, 
<geoCoordinate degrees="103.7748" direction="east" orientation="longitude" precision="5" value="103.7748">103.7748°E</geoCoordinate>
to 
<geoCoordinate degrees="1.3572" direction="north" orientation="latitude" precision="5" value="1.3572">1.3572°N</geoCoordinate>
, 
<geoCoordinate degrees="103.7734" direction="east" orientation="longitude" precision="5" value="103.7734">103.7734°E</geoCoordinate>
, 110-150 m elev. 12 June 2019 Maddison, Morehouse, &amp; Marathe WPM#19-051 (one female); Bukit Timah Nature Reserve. 
<geoCoordinate degrees="1.355" direction="north" orientation="latitude" precision="55" value="1.355">1.355°N</geoCoordinate>
, 
<geoCoordinate degrees="103.78" direction="east" orientation="longitude" precision="555" value="103.78">103.78°E</geoCoordinate>
, 29 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-041 (1 female 1 juvenile). Malaysia: Johor: Gunung Belumut Recreational Forest. 
<geoCoordinate degrees="2.066" direction="north" orientation="latitude" precision="55" value="2.066">2.066°N</geoCoordinate>
, 
<geoCoordinate degrees="103.527" direction="east" orientation="longitude" precision="55" value="103.527">103.527°E</geoCoordinate>
, 60-100 m el. 24 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-038 (1 male, 1 female, 6 juveniles); near Kluang, Gunung Lambak. 
<geoCoordinate degrees="2.025" direction="north" orientation="latitude" precision="55" value="2.025">2.025°N</geoCoordinate>
, 
<geoCoordinate degrees="103.344" direction="east" orientation="longitude" precision="55" value="103.344">103.344°E</geoCoordinate>
, 50-100 m el. 25 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-039 (1 male). Pahang: Tanah Rata. Jungle Trail 9 from Robinson Falls. 
<geoCoordinate degrees="4.46" direction="north" orientation="latitude" precision="555" value="4.46">4.46°N</geoCoordinate>
, 
<geoCoordinate degrees="101.40" direction="east" orientation="longitude" precision="555" value="101.4">101.40°E</geoCoordinate>
, 1200-1500 m el. 21-22 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-035 (1 female). Selangor: Ulu Gombak Field Station, 
<geoCoordinate degrees="3.325" direction="north" orientation="latitude" precision="55" value="3.325">3.325°N</geoCoordinate>
, 
<geoCoordinate degrees="101.753" direction="east" orientation="longitude" precision="55" value="101.753">101.753°E</geoCoordinate>
, 250 m el. 16-19 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-026 (1 female 1 juvenile); canyon near Ulu Gombak, 
<geoCoordinate degrees="3.325" direction="north" orientation="latitude" precision="55" value="3.325">3.325°N</geoCoordinate>
, 
<geoCoordinate degrees="101.765" direction="east" orientation="longitude" precision="55" value="101.765">101.765°E</geoCoordinate>
, 275 m el. 17 May 2005. W. Maddison, D. Li, I. Agnarsson, J. X. Zhang. WPM#05-027 (1 female 1 juvenile). Sarawak: Fairy Caves, near Kuching, 1.381-2°N, 110.117-9°E, 20 m el. 10 March 2012 Maddison/Piascik/Ang/Lee WPM#12-011 (1 female). The female from Sarawak is listed with some hesitation. It may be conspecific with a male 
<taxonomicName authorityName="Maddison" authorityYear="2020" family="Pittosporaceae" genus="Maripanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Maripanthus" order="Apiales" pageId="0" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="27">Maripanthus</emphasis>
</taxonomicName>
from Brunei, which appears to be a closely related but distinct species, with embolus initially directed distinctly more to the dorsal, and slight different carapace markings (specimen JK 08.08.23.0004, in LKCNHM, from Brunei: Ulu Temburong National Park, Ashton Trail 
<geoCoordinate degrees="4.5428" direction="north" orientation="latitude" precision="5" value="4.5428">4.5428°N</geoCoordinate>
, 
<geoCoordinate degrees="115.1528" direction="east" orientation="longitude" precision="5" value="115.1528">115.1528°E</geoCoordinate>
J K H Koh 23 August 2008).
</paragraph>
</subSubSection>
</treatment>
</document>