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205 lines
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<document id="A4664849554247B151F6AB84109D9455" ID-CLB-Dataset="20208" ID-DOI="10.11646/zootaxa.5115.4.1" ID-GBIF-Dataset="b99d2dc5-e0a6-4f73-b2c6-b21b6350c69f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6361552" ID-ZooBank="320C2E00-8035-431A-8AD7-D483D81923F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647419732391" checkinUser="plazi" docAuthor="Ullah, Mishkat, Naeem, Muhammad, Mahmood, Khalid & Garner, Beulah" docDate="2022" docId="291CD3215309DC62FF6DF93B82D3FE43" docLanguage="en" docName="zootaxa.5115.4.1.pdf" docOrigin="Zootaxa 5115 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Stenochlaenius) dostojevskii" docType="treatment" docVersion="7" lastPageNumber="482" masterDocId="D525AB595317DC7DFFFAFFFA836DFF91" masterDocTitle="Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan" masterLastPageNumber="486" masterPageNumber="451" pageNumber="481" updateTime="1699318106451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="D96844E0EBA8C2A188AA8EFD2C278608">Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan</mods:title>
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<mods:namePart id="9199E9675A05DD3FFA56DB47E4E9EF4A">Ullah, Mishkat</mods:namePart>
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<mods:affiliation id="90F4975704228CF3C8D248BA2F7FDE1A">Insects Division, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom.</mods:affiliation>
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</mods:role>
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<mods:namePart id="A8669DE620D3F3652604C349A0AD7871">Naeem, Muhammad</mods:namePart>
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<mods:nameIdentifier id="2623209402FDE4977D5C457BF9D82999" type="ORCID">0000-0002-2025-2024</mods:nameIdentifier>
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<mods:affiliation id="AB11146AD30B72698B4D30A89BF373FC">muhammad. naeem @ uaar. edu. pk; https: // orcid. org / 0000 - 0002 - 2025 - 2024</mods:affiliation>
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<mods:nameIdentifier id="ED94C8DF0352FB8118ACC4E1B39E6CDF" type="email">muhammad.naeem@uaar.edu.pk</mods:nameIdentifier>
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</mods:name>
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</mods:role>
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<mods:namePart id="7450479726C99DBA7F9E505C7552BCD7">Mahmood, Khalid</mods:namePart>
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<mods:nameIdentifier id="242C6D4370037D962BF78D15FBDF31E9" type="ORCID">0000-0001-7162-3430</mods:nameIdentifier>
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<mods:affiliation id="A9E214D8E7E14A14E3702D4478F31654">dkmahmood @ msn. com; https: // orcid. org / 0000 - 0001 - 7162 - 3430</mods:affiliation>
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<mods:nameIdentifier id="D6A4E94A37677ED80905A4D37A7515B6" type="email">dkmahmood@msn.com</mods:nameIdentifier>
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</mods:role>
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<mods:namePart id="390AB02AE245C212BB9ADC29D11D4D1F">Garner, Beulah</mods:namePart>
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<mods:nameIdentifier id="C369E7AB3236E03BC6D3D7A820E2A0F5" type="ORCID">0000-0002-5229-2450</mods:nameIdentifier>
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<mods:affiliation id="4C693FA16A96376ED66A370A88803F6E">Insects Division, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom. & b. garner @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 5229 - 2450</mods:affiliation>
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<mods:nameIdentifier id="AB3076D9388478DE4CEFE028936629CE" type="email">b.garner@nhm.ac.uk</mods:nameIdentifier>
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<mods:typeOfResource id="9ACC6E74AA7773BA13C0EF189B165088">text</mods:typeOfResource>
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<mods:title id="2B2623BE0BD7024606C682B72DE52E2B">Zootaxa</mods:title>
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<mods:part id="D0BD3A7922E881CED3A66C6C9537A756">
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<mods:date id="D525D82DBEF5C1943360CDA6A644B9C5">2022</mods:date>
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<mods:number id="EBEB84DB4E902D84188FCCCCC6F063F6">2022-03-16</mods:number>
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<mods:number id="500E75EC051FC2A72A726874C577D2BD">5115</mods:number>
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<mods:number id="7F0C2F66120DCF10094257876166A9F8">4</mods:number>
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<mods:identifier id="CCEEDD37F7C8F0369DB2BA60C26FD3B7" type="DOI">10.11646/zootaxa.5115.4.1</mods:identifier>
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</mods:mods>
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<treatment id="291CD3215309DC62FF6DF93B82D3FE43" ID-DOI="http://doi.org/10.5281/zenodo.6361524" ID-GBIF-Taxon="194051276" ID-Zenodo-Dep="6361524" LSID="urn:lsid:plazi:treatment:291CD3215309DC62FF6DF93B82D3FE43" httpUri="http://treatment.plazi.org/id/291CD3215309DC62FF6DF93B82D3FE43" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481">
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<subSubSection id="E9AF31BC5309DC63FF6DF93B80FEF94A" box="[151,915,1729,1756]" pageId="30" pageNumber="481" type="nomenclature">
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<paragraph id="A10A62375309DC63FF6DF93B80FEF94A" blockId="30.[151,915,1729,1792]" box="[151,915,1729,1756]" pageId="30" pageNumber="481">
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<heading id="FA42D55B5309DC63FF6DF93B80FEF94A" bold="true" box="[151,915,1729,1756]" fontSize="11" level="1" pageId="30" pageNumber="481" reason="1">
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<taxonomicName id="66B519B45309DC63FF6DF93B80FEF94A" ID-CoL="69VNM" authority="(Tschitscherine, 1896)" baseAuthorityName="Tschitscherine" baseAuthorityYear="1896" box="[151,915,1729,1756]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="dostojevskii" subGenus="Stenochlaenius">
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<emphasis id="93C1BE255309DC63FF6DF93B80FEF94A" bold="true" box="[151,915,1729,1756]" pageId="30" pageNumber="481">
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<emphasis id="93C1BE255309DC63FF6DF93B827EF94D" bold="true" box="[151,275,1729,1756]" italics="true" pageId="30" pageNumber="481">Chlaenius</emphasis>
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(
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<emphasis id="93C1BE255309DC63FEDEF93B82B3F94D" bold="true" box="[292,478,1729,1756]" italics="true" pageId="30" pageNumber="481">Stenochlaenius</emphasis>
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)
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<emphasis id="93C1BE255309DC63FE15F93B8116F94D" bold="true" box="[495,635,1729,1756]" italics="true" pageId="30" pageNumber="481">dostojevskii</emphasis>
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(
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<bibRefCitation id="C5241FC65309DC63FD70F93B80E6F94A" author="Tschitscherine, T." box="[650,907,1729,1755]" pageId="30" pageNumber="481" pagination="494 - 503" refId="ref27997" refString="Tschitscherine, T. (1896) Description de quelques carabiques nouveaux ou peu connus. Horae Societatis Entomologicae Rossicae, 29, 494 - 503." type="journal article" year="1896">Tschitscherine, 1896</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9AF31BC5309DC63FF6DF91C8268F891" box="[151,261,1766,1792]" pageId="30" pageNumber="481" type="description">
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<paragraph id="A10A62375309DC63FF6DF91C8268F891" blockId="30.[151,915,1729,1792]" box="[151,261,1766,1792]" pageId="30" pageNumber="481">
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(
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<figureCitation id="398E7EB25309DC63FF5AF91C8390F891" box="[160,253,1766,1792]" captionStart="FIGURE 10" captionStartId="31.[151,250,1495,1519]" captionTargetBox="[151,1436,489,1472]" captionTargetId="figure-122@31.[151,1436,489,1472]" captionTargetPageId="31" captionText="FIGURE 10. Dorsal habitus (a) Chlaenius (Stenochlaenius) dostojevskii Tschitscherine, 1896 imaged from NHMUK collection; (b) Harpaglossus opacus (Chaudoir, 1857) imaged from NHMUK collection." figureDoi="http://doi.org/10.5281/zenodo.6361581" httpUri="https://zenodo.org/record/6361581/files/figure.png" pageId="30" pageNumber="481">Fig. 10a</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="E9AF31BC5309DC63FF6DF8D081ADF8D3" box="[151,704,1834,1858]" pageId="30" pageNumber="481" type="reference_group">
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<paragraph id="A10A62375309DC63FF6DF8D081ADF8D3" blockId="30.[151,704,1834,1858]" box="[151,704,1834,1858]" pageId="30" pageNumber="481">
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<treatmentCitationGroup id="81A545195309DC63FF6DF8D081ADF8D3" box="[151,704,1834,1858]" pageId="30" pageNumber="481">
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<treatmentCitation id="201444265309DC63FF6DF8D081D1F8D3" author="Tschitscherine, T." box="[151,700,1834,1858]" page="497" pageId="30" pageNumber="481" year="1896">
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<taxonomicName id="66B519B45309DC63FF6DF8D081D1F8D3" ID-CoL="69VNM" authority="Tschitscherine, 1896: 497" authorityName="Tschitscherine" authorityPageNumber="497" authorityYear="1896" box="[151,700,1834,1858]" class="Insecta" family="Carabidae" genus="Stenochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="dostojevskii">
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<emphasis id="93C1BE255309DC63FF6DF8D082C2F8D3" box="[151,431,1834,1858]" italics="true" pageId="30" pageNumber="481">Stenochlaenius dostojevskii</emphasis>
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<bibRefCitation id="C5241FC65309DC63FE4EF8D081D1F8D3" author="Tschitscherine, T." box="[436,700,1834,1858]" pageId="30" pageNumber="481" pagination="494 - 503" refId="ref27997" refString="Tschitscherine, T. (1896) Description de quelques carabiques nouveaux ou peu connus. Horae Societatis Entomologicae Rossicae, 29, 494 - 503." type="journal article" year="1896">Tschitscherine, 1896: 497</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9AF31BC5309DC63FF6DF88A825DF842" pageId="30" pageNumber="481" type="materials_examined">
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<paragraph id="A10A62375309DC63FF6DF88A825DF842" blockId="30.[151,1436,1904,2039]" pageId="30" pageNumber="481">
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<emphasis id="93C1BE255309DC63FF6DF88A8212F81B" bold="true" box="[151,383,1904,1930]" pageId="30" pageNumber="481">Material examined.</emphasis>
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<materialsCitation id="11DD686A5309DC63FE7FF88B825DF842" ID-GBIF-Occurrence="3709719315" collectingDate="1929-07-22" collectingDateMax="1929-07-25" collectingDateMin="1929-07-22" collectionCode="NHMUK" collectorName="B. N. Chopra" country="Pakistan" county="Chitral" location="Karakal" municipality="Bumburat Valley" pageId="30" pageNumber="481" specimenCount="1" specimenCount-male="1" stateProvince="Khyber Pakhtunkhwa">
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<collectingCountry id="D9A222A75309DC63FE7FF88B8166F81A" box="[389,523,1905,1931]" name="Pakistan" pageId="30" pageNumber="481">PAKISTAN</collectingCountry>
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•
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<specimenCount id="B7B3A9BE5309DC63FDDAF88B8126F81A" box="[544,587,1905,1931]" pageId="30" pageNumber="481" type="male">1 ♂</specimenCount>
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;
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<collectingRegion id="6371ACD55309DC63FDACF88A8027F81B" box="[598,842,1904,1930]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="30" pageNumber="481">Khyber Pakhtunkhwa</collectingRegion>
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,
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<collectingCounty id="486B1ABB5309DC63FCAEF88A80C8F81B" box="[852,933,1904,1930]" pageId="30" pageNumber="481">Chitral</collectingCounty>
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,
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<collectingMunicipality id="416EF84D5309DC63FC55F88A8700F81B" box="[943,1133,1904,1930]" pageId="30" pageNumber="481">Bumburat Valley</collectingMunicipality>
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,
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<location id="A46A34EC5309DC63FB8CF88A87B8F81B" LSID="urn:lsid:plazi:treatment:291CD3215309DC62FF6DF93B82D3FE43:A46A34EC5309DC63FB8CF88A87B8F81B" box="[1142,1237,1904,1930]" country="Pakistan" county="Chitral" municipality="Bumburat Valley" name="Karakal" pageId="30" pageNumber="481" stateProvince="Khyber Pakhtunkhwa">Karakal</location>
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;,
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<date id="D50B44F75309DC63FB19F88A86FAF81A" box="[1251,1431,1904,1931]" pageId="30" pageNumber="481" value="1929-07-22" valueMax="1929-07-25" valueMin="1929-07-22">
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<collectingDate id="C54FBD1F5309DC63FB19F88A86FAF81A" box="[1251,1431,1904,1931]" pageId="30" pageNumber="481" value="1929-07-22" valueMax="1929-07-25" valueMin="1929-07-22">22–25 Jul. 1929</collectingDate>
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</date>
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;
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<collectorName id="0C4007E15309DC63FF6DF86F8248F83F" box="[151,293,1940,1967]" pageId="30" pageNumber="481">B.N. Chopra</collectorName>
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(
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<location id="A46A34EC5309DC63FECEF86E810BF83E" LSID="urn:lsid:plazi:treatment:291CD3215309DC62FF6DF93B82D3FE43:A46A34EC5309DC63FECEF86E810BF83E" box="[308,614,1940,1967]" country="Pakistan" county="Chitral" municipality="Bumburat Valley" name="Zoological Survey of India" pageId="30" pageNumber="481" stateProvince="Khyber Pakhtunkhwa">Zoological Survey of India</location>
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) leg.; ex coll.
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<collectorName id="0C4007E15309DC63FCFDF86E8772F83F" box="[775,1055,1940,1967]" pageId="30" pageNumber="481">Indian Museum Calcutta</collectorName>
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;
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<collectionCode id="C7A4FAF25309DC63FBD1F86F87F8F83E" box="[1067,1173,1941,1967]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="30" pageNumber="481" type="Museum">NHMUK</collectionCode>
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(
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<collectorName id="0C4007E15309DC63FB5EF86F83AEF843" pageId="30" pageNumber="481">H.E. Andrews Collection</collectorName>
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1945-97)
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9AF31BC5309DC62FF3DF82783B7FF44" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481" type="description">
|
|||
|
|
<paragraph id="A10A62375309DC62FF3DF82783B7FF44" blockId="30.[151,1436,1904,2039]" lastBlockId="31.[151,1437,151,466]" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481">
|
|||
|
|
<emphasis id="93C1BE255309DC63FF3DF8278213F866" bold="true" box="[199,382,2013,2039]" pageId="30" pageNumber="481">Measurements.</emphasis>
|
|||
|
|
Male (n= 2): TL= 12.00 mm, HW=
|
|||
|
|
<quantity id="664DCFD25309DC63FCDCF82780E3F866" box="[806,910,2013,2039]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="30" pageNumber="481" unit="mm" value="2.2">2.20 mm</quantity>
|
|||
|
|
, PL=
|
|||
|
|
<quantity id="664DCFD25309DC63FC2FF8268750F866" box="[981,1085,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="481" unit="mm" value="2.5">2.50 mm</quantity>
|
|||
|
|
, PW=
|
|||
|
|
<quantity id="664DCFD25309DC63FB77F8268798F866" box="[1165,1269,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="481" unit="mm" value="2.5">2.50 mm</quantity>
|
|||
|
|
, EL=
|
|||
|
|
<quantity id="664DCFD25309DC63FAC4F82686F5F866" box="[1342,1432,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="30" pageNumber="481" unit="mm" value="7.5">7.5 mm</quantity>
|
|||
|
|
, EW=
|
|||
|
|
<quantity id="664DCFD25308DC62FF26FF6D8228FF20" box="[220,325,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="31" pageNumber="482" unit="mm" value="2.3">2.30 mm</quantity>
|
|||
|
|
. Ratios PL/PW= 1.0, HW/PW= 0.88 and EL/EW= 3.26. No females were studied / available for study.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="E9AF31BC5308DC62FF3DFF248149FED0" pageId="31" pageNumber="482" type="diagnosis">
|
|||
|
|
<paragraph id="A10A62375308DC62FF3DFF248149FED0" blockId="31.[151,1437,151,466]" pageId="31" pageNumber="482">
|
|||
|
|
<emphasis id="93C1BE255308DC62FF3DFF24822FFF69" bold="true" box="[199,322,222,248]" pageId="31" pageNumber="482">Diagnosis.</emphasis>
|
|||
|
|
This species is almost like
|
|||
|
|
<taxonomicName id="66B519B45308DC62FD7FFF25800CFF68" authorityName="J. Schmidt" authorityYear="2007" baseAuthorityName="Grundmann" baseAuthorityYear="1955" box="[645,865,223,249]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="kashmiricus" subGenus="Stenochlaenius">
|
|||
|
|
<emphasis id="93C1BE255308DC62FD7FFF2581F2FF68" box="[645,671,223,249]" italics="true" pageId="31" pageNumber="482">C.</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="93C1BE255308DC62FD48FF2581AAFF68" box="[690,711,223,249]" italics="true" pageId="31" pageNumber="482">S.</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="93C1BE255308DC62FD20FF25800CFF68" box="[730,865,223,249]" italics="true" pageId="31" pageNumber="482">kashmiricus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
with the following differences, pronotum anterolateral angles protruding; pronotum lateral border less convex and continuous with less oblique deflection towards basal border. Basal margin straight.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="E9AF31BC5308DC62FF3DFEB0809AFE18" pageId="31" pageNumber="482" type="discussion">
|
|||
|
|
<paragraph id="A10A62375308DC62FF3DFEB0809AFE18" blockId="31.[151,1437,151,466]" pageId="31" pageNumber="482">
|
|||
|
|
<emphasis id="93C1BE255308DC62FF3DFEB08222FEF5" bold="true" box="[199,335,330,356]" pageId="31" pageNumber="482">Comments.</emphasis>
|
|||
|
|
<taxonomicName id="66B519B45308DC62FEACFEB1815AFEF4" authorityName="Tschitscherine" authorityYear="1896" box="[342,567,331,357]" class="Insecta" family="Carabidae" genus="Stenochlaenius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="dostojevskii" subGenus="Stenochlaenius">
|
|||
|
|
<emphasis id="93C1BE255308DC62FEACFEB18213FEF4" box="[342,382,331,357]" italics="true" pageId="31" pageNumber="482">Ch.</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="93C1BE255308DC62FE74FEB182CEFEF4" box="[398,419,331,357]" italics="true" pageId="31" pageNumber="482">S.</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="93C1BE255308DC62FE49FEB1815AFEF4" box="[435,567,331,357]" italics="true" pageId="31" pageNumber="482">dostojevskii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
was collected from Chitral valley during 1929. It was not encountered during this current study. This is the first time this species is reported from
|
|||
|
|
<collectingCountry id="D9A222A75308DC62FC6BFE958099FE18" box="[913,1012,367,393]" name="Pakistan" pageId="31" pageNumber="482">Pakistan</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="E9AF31BC5308DC62FF3DFE6882D3FE43" pageId="31" pageNumber="482" type="distribution">
|
|||
|
|
<paragraph id="A10A62375308DC62FF3DFE6882D3FE43" blockId="31.[151,1437,151,466]" pageId="31" pageNumber="482">
|
|||
|
|
<emphasis id="93C1BE255308DC62FF3DFE68820CFE3D" bold="true" box="[199,353,402,428]" pageId="31" pageNumber="482">Distribution.</emphasis>
|
|||
|
|
<collectingCountry id="D9A222A75308DC62FE9CFE69829FFE3C" box="[358,498,403,429]" name="Afghanistan" pageId="31" pageNumber="482">Afghanistan</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="D9A222A75308DC62FE04FE698143FE3C" box="[510,558,403,429]" name="Iran" pageId="31" pageNumber="482">Iran</collectingCountry>
|
|||
|
|
, Kashmir,
|
|||
|
|
<collectingCountry id="D9A222A75308DC62FD5DFE698040FE3C" box="[679,813,403,429]" name="Kazakhstan" pageId="31" pageNumber="482">Kazakhstan</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="D9A222A75308DC62FCC2FE6980C4FE3C" box="[824,937,403,429]" name="Tajikistan" pageId="31" pageNumber="482">Tajikistan</collectingCountry>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="D9A222A75308DC62FC4FFE69875EFE3C" box="[949,1075,403,429]" name="Uzbekistan" pageId="31" pageNumber="482">Uzbekistan</collectingCountry>
|
|||
|
|
(Anichtchenko 2017) and
|
|||
|
|
<collectingCountry id="D9A222A75308DC62FA9BFE6983A9FE40" name="Pakistan" pageId="31" pageNumber="482">Pakistan</collectingCountry>
|
|||
|
|
(first country record).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|