203 lines
26 KiB
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203 lines
26 KiB
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<document id="B768309136CF111D30A773FE2AD893FF" ID-ISSN="0024-4082" ID-ZooBank="4ED8513E-2C4F-4F17-81A1-4D4E1F2EABF5" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632744481225" checkinUser="plazi" docAuthor="Abello, María Alejandra, Martin, Gabriel M. & Cardoso, Yamila" docDate="2021" docId="006E64249F16FFDAA551FCD6361AFE70" docLanguage="en" docName="zlaa165.pdf" docOrigin="Zoological Journal of the Linnean Society 193" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Caenolestoides miocaenicus Abello & Martin & Cardoso 2021, SP. NOV." docType="treatment" docUuid="42675F71-06BC-49C9-83C0-62689384476C" docUuidSource="ZooBank" docVersion="2" masterDocId="FC571C5C9F0CFFC6A62BFFDA3424FFE3" masterDocTitle="Review of the extinct ‘ shrew-opossums’ (Marsupialia: Caenolestidae), with descriptions of two new genera and three new species from the Early Miocene of southern South America" updateTime="1699415251967" updateUser="plazi">
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<mods:mods id="18B3CDBE55BDDC4E73506599C7C41B47" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="04F3545F6D2CEEC76286B6E5288B4DF7">
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<mods:title id="37FC0D565DA832C8ACD7C5E3E8AFABAF">Review of the extinct ‘ shrew-opossums’ (Marsupialia: Caenolestidae), with descriptions of two new genera and three new species from the Early Miocene of southern South America</mods:title>
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</mods:titleInfo>
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<mods:name id="2A30A11C5BD58F4BC3E79869F02772C1" type="personal">
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<mods:roleTerm id="8EB065E81404B3D79217651CD57CE9A5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DA2E49028370873F7764A17204A2535C">Abello, María Alejandra</mods:namePart>
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</mods:name>
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<mods:name id="8A7D33FF4DD3D49347E95F097B6E0771" type="personal">
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<mods:role id="4E05EDA0BEB189D0729D1C1C321137F4">
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<mods:roleTerm id="489650C93A8E68013004D1BEECFF28A3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EF32515F2EFA330FE469A429EFAB1E4D">Martin, Gabriel M.</mods:namePart>
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</mods:name>
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<mods:name id="851004B8D54F054F69177C6C54E861C0" type="personal">
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<mods:role id="C7EDF07A1B0FB62D68ACE1DB7AD89241">
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<mods:roleTerm id="57E1A3FBCAD706A79CF2FF18FCA73AE4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ED3F4985A152318366DABB960872FD49">Cardoso, Yamila</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7D466E0D51AF9065EB7D37E07C1EAB84">text</mods:typeOfResource>
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<mods:relatedItem id="0499B7419D6B422B9B44C81418045A40" type="host">
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<mods:titleInfo id="CCD1A1098B91224DCA9ABD5542ADCB64">
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<mods:title id="A897A4C27BAAEC0ACDCAEC264D0890C9">Zoological Journal of the Linnean Society</mods:title>
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</mods:titleInfo>
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<mods:part id="D01D5DE6490917B0DBE8E2EAE2D5EDC1">
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<mods:date id="BDE458E9F331DB4DD6EF310B572CCD7F">2021</mods:date>
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<mods:detail id="AA243A6F486ABB83F969F474C95F1BB1" type="volume">
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<mods:number id="0943D632DFE7446154532060B2996BC4">193</mods:number>
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</mods:detail>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="C0369E2848F8D56FFA0D95C0490222B2">journal volume</mods:classification>
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<mods:identifier id="97B78BDBB528E756AD8D329D493D4651" type="ISSN">0024-4082</mods:identifier>
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<mods:identifier id="D59C46F28B5ED4CFEF2173412394AC76" type="ZooBank">4ED8513E-2C4F-4F17-81A1-4D4E1F2EABF5</mods:identifier>
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</mods:mods>
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<treatment id="006E64249F16FFDAA551FCD6361AFE70" LSID="urn:lsid:zoobank.org:act:42675F71-06BC-49C9-83C0-62689384476C" httpUri="http://treatment.plazi.org/id/006E64249F16FFDAA551FCD6361AFE70" lastPageId="28" pageId="26">
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<subSubSection id="C0DD86B99F16FFDCA551FCD63147FCC7" box="[890,1379,780,804]" pageId="26" type="nomenclature">
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<paragraph id="8878D5329F16FFDCA551FCD63147FCC7" blockId="26.[890,1379,780,804]" box="[890,1379,780,804]" pageId="26">
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<heading id="D330625E9F16FFDCA551FCD63147FCC7" box="[890,1379,780,804]" centered="true" fontSize="9" level="2" pageId="26" reason="2">
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<taxonomicName id="4FC7AEB19F16FFDCA551FCD630DDFCC1" authority="Abello & Martin & Cardoso, 2021" authorityName="Abello & Martin & Cardoso" authorityYear="2021" box="[890,1273,780,804]" class="Mammalia" family="Caenolestidae" genus="Caenolestoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="26" phylum="Chordata" rank="species" species="miocaenicus" status="sp. nov.">CAENOLESTOIDES MIOCAENICUS</taxonomicName>
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<emphasis id="BAB309209F16FFDCA32BFCD53147FCC7" bold="true" box="[1280,1379,780,804]" pageId="26">
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<taxonomicNameLabel id="A180B45B9F16FFDCA32BFCD53147FCC7" box="[1280,1379,780,804]" pageId="26" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C0DD86B99F16FFDAA20CFCE8361AFE70" lastPageId="28" pageId="26" type="description">
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<paragraph id="8878D5329F16FFDCA20CFCE83092FCA8" blockId="26.[1063,1206,818,843]" box="[1063,1206,818,843]" pageId="26">
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(
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<figureCitation id="10FCC9B79F16FFDCA204FCE8308BFCA8" box="[1071,1199,818,843]" captionStart="Figure 10" captionStartId="24.[163,244,1789,1811]" captionTargetBox="[228,1378,198,1746]" captionTargetId="figure-27@24.[225,1380,195,1749]" captionTargetPageId="24" captionText="Figure 10. Gaimanlestes pascuali and Caenolestoides miocaenicus. A–E, G. pascuali: A–C, type MPEF-PV 4849, left mandibular fragment with m1-3 and alveolus for the m4 in labial (A), lingual (B) and occlusal (C) views; D, E, MPEF-PV 5735, right mandibular fragment with p3-m1 and alveoli of m2-4 in lingual (D) and occlusal (E) views. F–J, C. miocaenicus:" pageId="26">FIG. 10F–J</figureCitation>
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)
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</paragraph>
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<paragraph id="8878D5329F16FFDCA510FC8031B3FC71" blockId="26.[827,1442,858,914]" pageId="26">
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<emphasis id="BAB309209F16FFDCA510FC80308CFC91" box="[827,1192,858,882]" italics="true" pageId="26">Zoobank registration: lsid</emphasis>
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:
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<uri id="FC56D9309F16FFDCA285FC8031B3FC71" pageId="26">
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urn:lsid:zoobank. org:act:
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<uuid id="FC61EFE79F16FFDCA5A0FCA031B3FC71" box="[907,1431,890,914]" pageId="26">42675F71-06BC-49C9-83C0-62689384476C</uuid>
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</uri>
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</paragraph>
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<paragraph id="8878D5329F16FFDCA510FC743118FBFC" blockId="26.[827,1442,942,1056]" pageId="26">
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<emphasis id="BAB309209F16FFDCA510FC74379FFC20" box="[827,955,942,963]" italics="true" pageId="26">Etymology:</emphasis>
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The epithet is composed of the geological era Miocene (in turn from Ancient Greek μείων, less and καινός, new) and the Latin suffix
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<emphasis id="BAB309209F16FFDCA2C0FC363102FBE2" box="[1259,1318,1004,1025]" italics="true" pageId="26">–icus</emphasis>
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, belonging to, in reference to its chronologic provenance.
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</paragraph>
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<paragraph id="8878D5329F16FFDCA510FB923036FB59" blockId="26.[827,1442,1096,1210]" pageId="26">
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<materialsCitation id="38AFDF6F9F16FFDCA510FB923036FB59" pageId="26" specimenCode="MPEF-PV 553" specimenCount="1" typeStatus="holotype">
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<emphasis id="BAB309209F16FFDCA510FB923783FBBE" box="[827,935,1096,1117]" italics="true" pageId="26">
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<typeStatus id="577C6B909F16FFDCA510FB923785FBBE" box="[827,929,1096,1117]" pageId="26" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<specimenCode id="D8617D499F16FFDCA598FB933078FBBD" box="[947,1116,1096,1118]" collectionCode="MPEF-PV" country="Argentina" httpUri="http://grbio.org/cool/pnqa-7dnx" name="Muso Paleontologico Egidio Fergulio" pageId="26">MPEF-PV 553</specimenCode>
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, right mandibular fragment with alveoli of the i2, c and p1, fragmentary p2, roots of p3, posterior root and talonid of m1 and complete m2-4 (
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<figureCitation id="10FCC9B79F16FFDCA5A8FB7E3027FB5A" box="[899,1027,1188,1210]" captionStart="Figure 10" captionStartId="24.[163,244,1789,1811]" captionTargetBox="[228,1378,198,1746]" captionTargetId="figure-27@24.[225,1380,195,1749]" captionTargetPageId="24" captionText="Figure 10. Gaimanlestes pascuali and Caenolestoides miocaenicus. A–E, G. pascuali: A–C, type MPEF-PV 4849, left mandibular fragment with m1-3 and alveolus for the m4 in labial (A), lingual (B) and occlusal (C) views; D, E, MPEF-PV 5735, right mandibular fragment with p3-m1 and alveoli of m2-4 in lingual (D) and occlusal (E) views. F–J, C. miocaenicus:" pageId="26">Fig. 10F–H</figureCitation>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8878D5329F16FFDCA510FB39377CFAD5" blockId="26.[827,1444,1251,1334]" pageId="26">
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<emphasis id="BAB309209F16FFDCA510FB39302EFB1B" box="[827,1034,1251,1272]" italics="true" pageId="26">Referred material:</emphasis>
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MLP 85-VII-3–9, right mandibular fragment with m2 and alveoli of the p3-m1 (
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<figureCitation id="10FCC9B79F16FFDCA368FAD8376DFAD5" captionStart="Figure 10" captionStartId="24.[163,244,1789,1811]" captionTargetBox="[228,1378,198,1746]" captionTargetId="figure-27@24.[225,1380,195,1749]" captionTargetPageId="24" captionText="Figure 10. Gaimanlestes pascuali and Caenolestoides miocaenicus. A–E, G. pascuali: A–C, type MPEF-PV 4849, left mandibular fragment with m1-3 and alveolus for the m4 in labial (A), lingual (B) and occlusal (C) views; D, E, MPEF-PV 5735, right mandibular fragment with p3-m1 and alveoli of m2-4 in lingual (D) and occlusal (E) views. F–J, C. miocaenicus:" pageId="26">Fig. 10I, J</figureCitation>
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).
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</paragraph>
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<paragraph id="8878D5329F16FFDCA510FA8537B9FA70" blockId="26.[827,1443,1375,1427]" pageId="26">
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<emphasis id="BAB309209F16FFDCA510FA85301DFA97" box="[827,1081,1375,1396]" italics="true" pageId="26">Temporal distribution:</emphasis>
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Early Miocene, Colhuehuapian SALMA.
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</paragraph>
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<paragraph id="8878D5329F16FFDCA510FA66305BF9F3" blockId="26.[827,1442,1468,1552]" pageId="26">
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<emphasis id="BAB309209F16FFDCA510FA66313DFA32" box="[827,1305,1468,1489]" italics="true" pageId="26">Stratigraphic and geographic distribution:</emphasis>
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Sarmiento Formation, Colhue-Huapi Member. Gran Barranca,
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<collectingRegion id="4A031BD09F16FFDCA510FA2037DEF9F3" box="[827,1018,1530,1552]" country="Argentina" name="Chubut" pageId="26">Chubut province</collectingRegion>
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,
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<collectingCountry id="F0D095A29F16FFDCA22FFA20305FF9F3" box="[1028,1147,1530,1552]" name="Argentina" pageId="26">Argentina</collectingCountry>
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.
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</paragraph>
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<paragraph id="8878D5329F16FFDCA510F9E33787F98F" blockId="26.[827,1441,1592,1645]" pageId="26">
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<emphasis id="BAB309209F16FFDCA510F9E337D1F9AD" box="[827,1013,1593,1614]" italics="true" pageId="26">Measurements:</emphasis>
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Supporting Information, File S6,
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<tableCitation id="C545E0899F16FFDCA510F98D37BFF98F" box="[827,923,1623,1645]" captionStart="Table 1" captionStartId="1.[145,209,834,855]" captionText="Table 1. Temporal and geographical range of caenolestid species" pageId="26">Table 1a</tableCitation>
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.
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</paragraph>
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<paragraph id="8878D5329F16FFDDA510F94B35CAFF1A" blockId="26.[827,1443,1680,1856]" lastBlockId="27.[145,761,197,250]" lastPageId="27" pageId="26">
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<emphasis id="BAB309209F16FFDCA510F94B312BF945" box="[827,1295,1680,1702]" italics="true" pageId="26">
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Diagnosis:
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<taxonomicName id="4FC7AEB19F16FFDCA5E0F94A312BF945" authorityName="Abello & Martin & Cardoso" authorityYear="2021" box="[971,1295,1680,1702]" class="Mammalia" family="Caenolestidae" genus="Caenolestoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="26" phylum="Chordata" rank="species" species="miocaenicus">Caenolestoides miocaenicus</taxonomicName>
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</emphasis>
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differs from the remaining caenolestids by a (1) long anterobasal cingulum in m2-3, (2) transversally aligned meta- and protoconid in m2-3, (3) close para- and metaconids in m2-3, (4) presence of a shelf lingual to the entocristid of m1-3, (5) entoconid of m2-3 posteriorly situated with respect to the hypoconid, (6) deep protocristid notch in m2-3 and (7) birradiculate m4.
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</paragraph>
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<paragraph id="8878D5329F17FFDDA6BAFEF937BEFB91" blockId="27.[145,761,290,1907]" lastBlockId="27.[809,1425,197,1476]" pageId="27">
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<emphasis id="BAB309209F17FFDDA6BAFEF9353AFEDB" box="[145,286,291,312]" italics="true" pageId="27">Description:</emphasis>
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Mandibular body of the
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<typeStatus id="577C6B909F17FFDDA461FEF93659FEDB" box="[586,637,291,312]" pageId="27">type</typeStatus>
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specimen, MPEF-PV 553, is somewhat higher below the molars than in the antemolar region. Labially, it has three mental foramina, with the posterior one smaller and opening below the m1-2 boundary, the intermediate foramen opens below p3, and the anterior largest one, opens below p2. Despite the mandibular body being broken in front of the anterior mental foramen, a well developed, anterior and forwardly directed groove is present. Anterior to the partially preserved p2, there are two alveoli; the anterior one is half the size of the posterior one and both are forwardly inclined, suggesting that they housed somewhat procumbent teeth. The p2 is biradiculate, has the anterior root smaller than the posterior one, is long and narrow, and probably was twice the size of the anterior tooth. The main cusp is missing, but comparing its base with the preserved talonid, it could be inferred that the talonid was as long as the main cusp. The talonid narrows backwards, ending in a small cuspule. Given the dimensions of the roots of p3, this premolar was clearly larger than p2. The m1 is only represented by its posterior root and the lingual half of the talonid, would have been equal or somewhat larger than m2. On the preserved part, the entoconid base shows this cusp might be similar in size to that of m2. The morphology of the talonid in front of, and lingual to, the entoconid suggest that a small shelf was developed lingually to this cusp; however, being incompletely preserved, the shelf cannot be compared in detail with that present in other caenolestids, such as
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<taxonomicName id="4FC7AEB19F17FFDDA43AFB6036E5FB33" box="[529,705,1210,1232]" class="Mammalia" family="Caenolestidae" genus="Pliolestes" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="undetermined">
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<emphasis id="BAB309209F17FFDDA43AFB6036A1FB2C" box="[529,645,1210,1231]" italics="true" pageId="27">Pliolestes</emphasis>
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spp.
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</taxonomicName>
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and
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<taxonomicName id="4FC7AEB19F17FFDDA6BAFB03350AFB0D" box="[145,302,1241,1262]" class="Mammalia" family="Caenolestidae" genus="Caenolestes" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="fuliginosus">
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<emphasis id="BAB309209F17FFDDA6BAFB03350AFB0D" box="[145,302,1241,1262]" italics="true" pageId="27">C. fuliginosus</emphasis>
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</taxonomicName>
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. The m2 is an elongated molar, with an anteroposteriorly shortened trigonid. The anterobasal cingulum is long and well developed, extending posteriorly to the hypoconid base. The protoconid is the largest trigonid cusp and is transversely aligned with the metaconid. The paraconid is the smallest trigonid cusp, located next to the metaconid and separated from this cusp by a narrow valley formed by the posterior wall of the paraconid, and the anterior wall of the metaconid. As in all other caenolestids, the paracristid is divided into a postparacristid and a preprotocristid. The postparacristid is transversely oriented and the preprotocristid is somewhat parallel to the anteroposterior axis of the molar. In contrast to extant cenolestids, the protocristid has a deep notch. The talonid is elongated with respect to the trigonid, and the most notable characteristic of the talonid is the position of the entoconid, which is posteriorly displaced relative to the hypoconid. The entoconid is relatively small and labiolingually compressed. The m2-3 entocristid is represented by a proximal portion that is oriented towards the talonid basin, as in all other caenolestids. The distal portion, which in other caenolestids is extended forward to the posterior wall of the trigonid, is absent or, more probably, worn. Therefore, entocristids do not show the typical curved morphology observed in the caenolestids. The talonid basin of m2 is anteroposteriorly wide and is labially limited by a high cristid obliqua. Towards the lingual border of the basin, a well-developed shelf extends between the metaconid and the entoconid. The hypoconid is labially salient and the postcristid is oblique, and although worn, it can be inferred that the postcristid should have been high. The hypoconulid is anteroposteriorlly compressed and is placed somewhat posterior to the entoconid. The morphology of m3 is close to that of m2, specially the talonid structure. The major differences among both molars are in the trigonid cusp arrangement. In m3, the paraconid is more reduced in relation to the metaconid and is more labially situated with respect to this cusp than in m2. The para- and metaconid are also more closely situated between each other than in m2. The m4 is the smallest molar of the series, it is biradiculate and reduced with respect to m3. As in the anterior molars, the trigonid is anteroposteriorly shortened. The meta- and protoconid are coalescent and the paraconid, although reduced, is well differentiated and placed somewhat labial to the paraconid. The two portions of the paracristid have the same orientation than in the anterior molars. The talonid is quadrangular in occlusal view, somewhat wider than the trigonid, and lacks any recognizable structure.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8878D5329F17FFDDA56AFBA1378CFA27" blockId="27.[809,1425,197,1476]" pageId="27">
|
|||
|
|
<materialsCitation id="38AFDF6F9F17FFDDA56AFBA13780FA27" collectionCode="MLP" pageId="27" specimenCode="MLP 85, MPEF-PV 553" specimenCount="1">
|
|||
|
|
When compared with the
|
|||
|
|
<typeStatus id="577C6B909F17FFDDA2A5FBA630E7FB72" box="[1166,1219,1148,1169]" pageId="27">type</typeStatus>
|
|||
|
|
specimen,
|
|||
|
|
<specimenCode id="D8617D499F17FFDDA37DFBA63761FB53" collectionCode="MLP" country="Argentina" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="27">MLP 85</specimenCode>
|
|||
|
|
-VII-3–9 has a shallower mandibular ramus but similar alveolar dimensions of p3-m1. In the preserved portion of this specimen there is no mental foramina. The only preserved molar, m2, is morphologically similar to that of
|
|||
|
|
<specimenCode id="D8617D499F17FFDDA5DBFACF30B2FAC9" box="[1008,1174,1300,1322]" collectionCode="MPEF-PV" country="Argentina" httpUri="http://grbio.org/cool/pnqa-7dnx" name="Muso Paleontologico Egidio Fergulio" pageId="27">MPEF-PV 553</specimenCode>
|
|||
|
|
, but somewhat larger. It is also less worn so that some molar traits are more clearly visible: the hypoconid, cristid obliqua and postcristid are high, and the cristid oblicua has a notch just behind the contact with the posterior wall of the protoconid
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8878D5329F17FFDAA502FA37361AFE70" blockId="27.[809,1426,1517,1907]" lastBlockId="28.[163,779,197,403]" lastPageId="28" pageId="27">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA502FA3737B3F9E1" box="[809,919,1517,1538]" italics="true" pageId="27">Remarks:</emphasis>
|
|||
|
|
Of all extinct caenolestids,
|
|||
|
|
<taxonomicName id="4FC7AEB19F17FFDDA2CEFA37379CF9C2" authorityName="Abello & Martin & Cardoso" authorityYear="2021" class="Mammalia" family="Caenolestidae" genus="Caenolestoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="miocaenicus">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA2CEFA37379CF9C2" italics="true" pageId="27">Caenolestoides miocaenicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is the most closely related to the living species, with which it shares three main characters of m2-3: long anterobasal cinguli, entoconid posterior to the hyponconid, and lingual platforms between meta and entoconid.
|
|||
|
|
<taxonomicName id="4FC7AEB19F17FFDDA5F1F95C313FF97F" authorityName="Abello & Martin & Cardoso" authorityYear="2021" box="[986,1307,1670,1692]" class="Mammalia" family="Caenolestidae" genus="Caenolestoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="miocaenicus">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA5F1F95C313FF97F" box="[986,1307,1670,1692]" italics="true" pageId="27">Caenolestoides miocaenicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is similar to
|
|||
|
|
<taxonomicName id="4FC7AEB19F17FFDDA56DF97F3067F958" authority="(Tomes)" baseAuthorityName="Tomes" baseAuthorityYear="1863" box="[838,1091,1701,1723]" class="Mammalia" family="Caenolestidae" genus="Caenolestes" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="fuliginosus">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA56DF97F37C6F959" box="[838,994,1701,1722]" italics="true" pageId="27">C. fuliginosus</emphasis>
|
|||
|
|
(Tomes)
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4FC7AEB19F17FFDDA256F97F3109F958" authority="Thomas" authorityName="Thomas" authorityYear="1917" box="[1149,1325,1701,1723]" class="Mammalia" family="Caenolestidae" genus="Lestoros" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="inca">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA256F97F30EFF959" box="[1149,1227,1701,1722]" italics="true" pageId="27">L. inca</emphasis>
|
|||
|
|
Thomas
|
|||
|
|
</taxonomicName>
|
|||
|
|
, sharing with these extant species the proximity between para- and metaconid, and a larger, compared size between m4 and m3, clearly different from the reduced m
|
|||
|
|
<quantity id="4F3F78D79F17FFDDA3A9F8DB3766F8D6" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="27" unit="in" value="4.0">4 in</quantity>
|
|||
|
|
<taxonomicName id="4FC7AEB19F17FFDDA564F8FA3034F8D7" authorityName="Osgood" authorityYear="1924" box="[847,1040,1823,1845]" class="Mammalia" family="Caenolestidae" genus="Rhyncholestes" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="raphanurus">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA564F8FA3034F8D7" box="[847,1040,1823,1845]" italics="true" pageId="27">R. raphanurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4FC7AEB19F17FFDDA20DF8C531B5F8D6" authorityName="Abello & Martin & Cardoso" authorityYear="2021" box="[1062,1425,1823,1845]" class="Mammalia" family="Caenolestidae" genus="Caenolestoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="miocaenicus">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA20DF8C531B5F8D6" box="[1062,1425,1823,1845]" italics="true" pageId="27">Caenolestoides miocaenicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differs from extant species
|
|||
|
|
<taxonomicName id="4FC7AEB19F17FFDDA25EF8E530E3F8B7" authorityName="Thomas" authorityYear="1917" box="[1141,1223,1855,1876]" class="Mammalia" family="Caenolestidae" genus="Lestoros" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="inca">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA25EF8E530E3F8B7" box="[1141,1223,1855,1876]" italics="true" pageId="27">L. inca</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4FC7AEB19F17FFDDA32CF8E4377CF891" class="Mammalia" family="Caenolestidae" genus="Caenolestes" kingdom="Animalia" order="Paucituberculata" pageId="27" phylum="Chordata" rank="species" species="undetermined">
|
|||
|
|
<emphasis id="BAB309209F17FFDDA32CF8E431B5F8B0" box="[1287,1425,1854,1875]" italics="true" pageId="27">Caenolestes</emphasis>
|
|||
|
|
spp.
|
|||
|
|
</taxonomicName>
|
|||
|
|
in some plesiomorphic characters such as a p2 shorter than p3 and deep protocristid notch in m2-3. Compared with
|
|||
|
|
<taxonomicName id="4FC7AEB19F10FFDAA74AFF3E3626FF1A" baseAuthorityName="Tomes" baseAuthorityYear="1863" box="[353,514,228,249]" class="Mammalia" family="Caenolestidae" genus="Caenolestes" kingdom="Animalia" order="Paucituberculata" pageId="28" phylum="Chordata" rank="species" species="fuliginosus">
|
|||
|
|
<emphasis id="BAB309209F10FFDAA74AFF3E3626FF1A" box="[353,514,228,249]" italics="true" pageId="28">C. fuliginosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4FC7AEB19F10FFDAA415FF3F36B4FF1A" authorityName="Thomas" authorityYear="1917" box="[574,656,228,250]" class="Mammalia" family="Caenolestidae" genus="Lestoros" kingdom="Animalia" order="Paucituberculata" pageId="28" phylum="Chordata" rank="species" species="inca">
|
|||
|
|
<emphasis id="BAB309209F10FFDAA415FF3F36B4FF1A" box="[574,656,228,250]" italics="true" pageId="28">L. inca</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the para- and metaconid in m2-3 are less twinned, the metaconid is slightly larger than the paraconid and the trigonids are proportionately wider than in the living species.
|
|||
|
|
<taxonomicName id="4FC7AEB19F10FFDAA688FE8435FEFE97" authorityName="Abello & Martin & Cardoso" authorityYear="2021" box="[163,474,350,372]" class="Mammalia" family="Caenolestidae" genus="Caenolestoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="28" phylum="Chordata" rank="species" species="miocaenicus">
|
|||
|
|
<emphasis id="BAB309209F10FFDAA688FE8435FEFE97" box="[163,474,350,372]" italics="true" pageId="28">Caenolestoides miocaenicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4FC7AEB19F10FFDAA7CEFE853646FE90" authorityName="Abello & Martin & Cardoso" authorityYear="2021" box="[485,610,350,372]" class="Mammalia" family="Caenolestidae" genus="Gaimanlestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="28" phylum="Chordata" rank="species" species="pascuali">
|
|||
|
|
<emphasis id="BAB309209F10FFDAA7CEFE853646FE90" box="[485,610,350,372]" italics="true" pageId="28">G. pascuali</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4FC7AEB19F10FFDAA4BCFE85372FFE97" authorityName="Abello & Martin & Cardoso" authorityYear="2021" box="[663,779,351,372]" class="Mammalia" family="Caenolestidae" genus="Stilotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Paucituberculata" pageId="28" phylum="Chordata" rank="species" species="parvum">
|
|||
|
|
<emphasis id="BAB309209F10FFDAA4BCFE85372FFE97" box="[663,779,351,372]" italics="true" pageId="28">S. parvum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are the oldest records for the family.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|