211 lines
25 KiB
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211 lines
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<document id="40F6413A11C0544F7D43E4DE11157559" ID-CLB-Dataset="9277" ID-DOI="10.11646/zootaxa.4895.2.3" ID-GBIF-Dataset="020059ec-0af5-44c5-8eb0-6a2874d9197d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4322473" ID-ZooBank="77117CFB-AD8E-4066-819F-FA81D3D589B7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1608025080741" checkinUser="plazi" docAuthor="Lawrence, John F." docDate="2020" docId="3735BF36A0045D78FF4695D070EAFBC8" docLanguage="en" docName="zootaxa.4895.2.3.pdf" docOrigin="Zootaxa 4895 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Atractocerinae Laporte 1840" docType="treatment" docVersion="8" lastPageNumber="223" masterDocId="CB0CC74EA0085D74FFD195477560FF91" masterDocTitle="The Australian Lymexylidae (Coleoptera: Tenerionoidea) with one new genus and two genera new to Australia" masterLastPageNumber="238" masterPageNumber="211" pageNumber="223" updateTime="1698847576668" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BBED937EFE16E8665A2A6B2307D950FC">The Australian Lymexylidae (Coleoptera: Tenerionoidea) with one new genus and two genera new to Australia</mods:title>
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<mods:namePart id="C1E2A11575696D28C28A4024697A788A">Lawrence, John F.</mods:namePart>
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<mods:title id="E2663DA8BF194EA97D3E37866007DEBA">Zootaxa</mods:title>
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<mods:part id="CCA849C8A6C01CD06BED1A50E0D8C250">
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<mods:date id="A8906899FBE2B198313208887F42F500">2020</mods:date>
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<mods:number id="B77E766816C1D1DA975CA957EA13564E">2020-12-15</mods:number>
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<mods:number id="2C7E853FF61BA495F9528D7130E292E6">4895</mods:number>
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<mods:number id="1662B7A782A203DC146F73E93FECDDC6">2</mods:number>
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<mods:identifier id="CE57F3CFB9FF59FADA6C09E08BD6677C" type="DOI">10.11646/zootaxa.4895.2.3</mods:identifier>
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<treatment id="3735BF36A0045D78FF4695D070EAFBC8" ID-DOI="http://doi.org/10.5281/zenodo.4361844" ID-GBIF-Taxon="176092565" ID-Zenodo-Dep="4361844" LSID="urn:lsid:plazi:treatment:3735BF36A0045D78FF4695D070EAFBC8" httpUri="http://treatment.plazi.org/id/3735BF36A0045D78FF4695D070EAFBC8" lastPageNumber="223" pageId="12" pageNumber="223">
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<subSubSection id="F7865DABA0045D78FF4695D07769FF20" box="[151,521,151,177]" pageId="12" pageNumber="223" type="nomenclature">
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<paragraph id="BF230E20A0045D78FF4695D07769FF20" blockId="12.[151,521,151,177]" box="[151,521,151,177]" pageId="12" pageNumber="223">
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<heading id="E46BB94CA0045D78FF4695D07769FF20" bold="true" box="[151,521,151,177]" fontSize="11" level="1" pageId="12" pageNumber="223" reason="1">
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<taxonomicName id="789C75A3A0045D78FF4695D07769FF20" authority="Laporte, 1840" authorityName="Laporte" authorityYear="1840" box="[151,521,151,177]" class="Insecta" family="Lymexylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="subFamily" subFamily="Atractocerinae">
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<emphasis id="8DE8D232A0045D78FF4695D07769FF20" bold="true" box="[151,521,151,177]" pageId="12" pageNumber="223">
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Atractocerinae
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<bibRefCitation id="DB0D73D1A0045D78FE8B95D07769FF20" author="Laporte, F. L. N. C. de" box="[346,521,151,177]" pageId="12" pageNumber="223" refId="ref28327" refString="Laporte, F. L. N. C. de (1840) Histoire naturelle des insectes Coleopteres; avec une introduction renfermant l'anatomie et la physiologie des animaux articules, par M. Brulle. Tome deuxieme. Histoire naturelle des animaux articules, annelides, crustaces, arachnides, myriapodes et insectes. Tome Troisieme. P. Duymenil, Paris, 563 pp., 38 pls." type="book" year="1840">Laporte, 1840</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="F7865DABA0045D78FF46959874A0FD38" pageId="12" pageNumber="223" type="reference_group">
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<paragraph id="BF230E20A0045D78FF4695987637FD8B" blockId="12.[151,1437,223,1114]" pageId="12" pageNumber="223">
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The
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<taxonomicName id="789C75A3A0045D78FF199598740DFF68" authorityName="Laporte" authorityYear="1840" box="[200,365,223,249]" class="Insecta" family="Lymexylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="subFamily" subFamily="Atractocerinae">Atractocerinae</taxonomicName>
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are unique among the
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<taxonomicName id="789C75A3A0045D78FDBE9598778AFF68" box="[623,746,223,249]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="223" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
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in many respects, but notably in the extreme reduction of the elytra combined with the lack of transverse folds in the hind wings, which are almost completely exposed, lying over the abdomen. Based on the wing character alone, this subfamily was treated as a beetle suborder, Aplicalae, by
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<bibRefCitation id="DB0D73D1A0045D78FF46940C7445FEF7" author="King, E. W." box="[151,293,331,358]" pageId="12" pageNumber="223" pagination="65 - 74" refId="ref27976" refString="King, E. W. (1955) The phylogenetic position of Atractocerus Palis. The Coleopterists Bulletin, 5, 65 - 74." type="journal article" year="1955">King (1955)</bibRefCitation>
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; but
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<bibRefCitation id="DB0D73D1A0045D78FE8A940C776FFEF7" author="Selander, R. B." box="[347,527,331,358]" pageId="12" pageNumber="223" pagination="1 - 5" refId="ref29239" refString="Selander, R. B. (1959) Additional remarks on wing structure in Atractocerus. The Coleopterists Bulletin, 13, 1 - 5." type="journal article" year="1959">Selander (1959)</bibRefCitation>
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demonstrated that this wing
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<typeStatus id="6027B082A0045D78FC84940B76E5FEF7" box="[853,901,332,358]" pageId="12" pageNumber="223">type</typeStatus>
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could be derived from that found in the lymexylid genus
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<taxonomicName id="789C75A3A0045D78FEC3942874F9FE18" ID-CoL="5NSR" authorityName="Murray" authorityYear="1867" box="[274,409,367,393]" class="Insecta" family="Lymexylidae" genus="Melittomma" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="8DE8D232A0045D78FEC3942874F9FE18" box="[274,409,367,393]" italics="true" pageId="12" pageNumber="223">Melittomma</emphasis>
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</taxonomicName>
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. Likewise,
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<bibRefCitation id="DB0D73D1A0045D78FDCD942877D0FE1B" author="Baehr, M." box="[540,688,367,394]" pageId="12" pageNumber="223" pagination="39 - 58" refId="ref26935" refString="Baehr, M. (1976) Das Prothorakalskelett von Atractocerus (Lymexylonidae) und seine Bedeutung fur die Phylogenie der Cole- opteren, besonders der Polyphagen (Insecta: Coleoptera) Zoomorphologie, 85, 39 - 58. https: // doi. org / 10.1007 / BF 00996064" type="journal article" year="1976">Baehr (1976)</bibRefCitation>
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in his detailed descriptive paper on the prothoracic morphology of “
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<taxonomicName id="789C75A3A0045D78FF7594D47455FE3C" ID-CoL="359N" authorityName="Palisot de Beauvois" authorityYear="1801" box="[164,309,403,429]" class="Insecta" family="Lymexylidae" genus="Atractocerus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="8DE8D232A0045D78FF7594D47455FE3C" box="[164,309,403,429]" italics="true" pageId="12" pageNumber="223">Atractocerus</emphasis>
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</taxonomicName>
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sp.” commented on several structures (e.g., spinasternum, dorsal cervical sclerites, karepisternum, which “have been lost” in all other
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<taxonomicName id="789C75A3A0045D78FDE194F077CFFE40" box="[560,687,439,465]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="12" pageNumber="223" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
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. Although this group appears to be derived from an ancestor well within the Polyphaga, the family
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<taxonomicName id="789C75A3A0045D78FDC3949C77C4FE64" ID-CoL="C9T" authorityName="Fleming" authorityYear="1821" box="[530,676,475,501]" class="Insecta" family="Lymexylidae" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="family">Lymexylidae</taxonomicName>
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being part of or sister to the large superfamily
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<taxonomicName id="789C75A3A0045D78FB6C949C700EFE64" authorityName="Latreille" authorityYear="1802" box="[1213,1390,475,501]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="12" pageNumber="223" phylum="Tracheophyta" rank="superFamily" superFamily="Tenebrionoidea">Tenebrionoidea</taxonomicName>
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, the adult has certainly undergone major changes in body structure.
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</paragraph>
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<paragraph id="BF230E20A0045D78FF16976474A0FD38" blockId="12.[151,1437,223,1114]" pageId="12" pageNumber="223">
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The ideas on relationships of atractocerines to other lymexylids have not really changed from those of
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<bibRefCitation id="DB0D73D1A0045D78FAEC97647584FDF3" author="Wheeler, Q. D." pageId="12" pageNumber="223" pagination="113 - 210" refId="ref29469" refString="Wheeler, Q. D. (1986) Revision of the genera of Lymexylidae (Coleoptera: Cucujiformia). Bulletin of the American Museum of Natural History, 183, 113 - 210." type="journal article" year="1986">Wheeler (1986)</bibRefCitation>
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, with the Palaearctic genus
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<taxonomicName id="789C75A3A0045D78FDC49700779BFDF0" ID-CoL="5HQ8" authority="Fabricius" authorityName="Fabricius" authorityYear="1775" box="[533,763,583,609]" class="Insecta" family="Lymexylidae" genus="Lymexylon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="8DE8D232A0045D78FDC4970077EDFDF0" box="[533,653,583,609]" italics="true" pageId="12" pageNumber="223">Lymexylon</emphasis>
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Fabricius
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</taxonomicName>
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as the sister group. It may be time, however, for a worldwide analysis to be performed based on adult and larval morphology and molecular data. This, however, is beyond the scope of the present study.
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</paragraph>
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</subSubSection>
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<subSubSection id="F7865DABA0045D78FF1697F470EAFBC8" pageId="12" pageNumber="223" type="description">
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<paragraph id="BF230E20A0045D78FF1697F470EAFBC8" blockId="12.[151,1437,223,1114]" pageId="12" pageNumber="223">
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As noted in the Introduction, the subfamily now includes six extant genera:
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<taxonomicName id="789C75A3A0045D78FBE797F470F7FD5F" authority="Kurosawa (1985)" authorityName="Kurosawa" authorityYear="1985" box="[1078,1431,691,718]" class="Insecta" family="Lymexylidae" genus="Arractocetus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="8DE8D232A0045D78FBE797F471A7FD5C" box="[1078,1223,691,717]" italics="true" pageId="12" pageNumber="223">Arractocetus</emphasis>
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<bibRefCitation id="DB0D73D1A0045D78FB0197F370F7FD5F" author="Kurosawa, Y." box="[1232,1431,691,718]" pageId="12" pageNumber="223" pagination="109 - 119" refId="ref28184" refString="Kurosawa, Y. (1985) Revisional notes on the family Lymexylonidae (Coleoptera) in eastern and southeastern Asia. Bulletin of the National Science Museum, Tokyo, (A), 11, 109 - 119." type="journal article" year="1985">Kurosawa (1985)</bibRefCitation>
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</taxonomicName>
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, with ten species from
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<collectingCountry id="C78B4EB0A0045D78FE5D979F74AFFD63" box="[396,463,728,754]" name="Japan" pageId="12" pageNumber="223">Japan</collectingCountry>
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,
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<collectingCountry id="C78B4EB0A0045D78FE089790774EFD60" box="[473,558,727,753]" name="Taiwan" pageId="12" pageNumber="223">Taiwan</collectingCountry>
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, Southeast Asia, the
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<collectingCountry id="C78B4EB0A0045D78FCC2979076F4FD60" box="[787,916,727,753]" name="Philippines" pageId="12" pageNumber="223">Philippines</collectingCountry>
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,
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<collectingCountry id="C78B4EB0A0045D78FC4E979076BBFD60" box="[927,987,727,753]" name="India" pageId="12" pageNumber="223">India</collectingCountry>
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,
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<collectingCountry id="C78B4EB0A0045D78FC3497907135FD60" box="[997,1109,727,753]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
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, New
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<collectingCountry id="C78B4EB0A0045D78FB4A9790718CFD60" box="[1179,1260,727,753]" name="Guinea-Bissau" pageId="12" pageNumber="223">Guinea</collectingCountry>
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and the
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<collectingCountry id="C78B4EB0A0045D78FA9B979075DEFC87" name="Indonesia" pageId="12" pageNumber="223">Moluccas</collectingCountry>
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;
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<taxonomicName id="789C75A3A0045D78FF1897BC775DFC84" authority="de Beauvois (1801)" authorityName="Palisot de Beauvois" authorityYear="1801" box="[201,573,763,789]" class="Insecta" family="Lymexylidae" genus="Atractocerus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="8DE8D232A0045D78FF1897BC743AFC84" box="[201,346,763,789]" italics="true" pageId="12" pageNumber="223">Atractocerus</emphasis>
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de Beauvois (1801)
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</taxonomicName>
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, with eight species from Central and South America, the West Indies, tropical Africa,
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<collectingCountry id="C78B4EB0A0045D78FF3A96587414FCA8" box="[235,372,799,825]" name="Madagascar" pageId="12" pageNumber="223">Madagascar</collectingCountry>
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,
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<collectingCountry id="C78B4EB0A0045D78FEAF965874DAFCA8" box="[382,442,799,825]" name="India" pageId="12" pageNumber="223">India</collectingCountry>
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,
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<collectingCountry id="C78B4EB0A0045D78FE1496587754FCA8" box="[453,564,799,825]" name="Sri Lanka" pageId="12" pageNumber="223">Sri Lanka</collectingCountry>
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, Sumatra and southern
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<collectingCountry id="C78B4EB0A0045D78FCE49658761BFCA8" box="[821,891,799,825]" name="China" pageId="12" pageNumber="223">China</collectingCountry>
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;
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<taxonomicName id="789C75A3A0045D78FC57966771CEFCAB" authority="Philippi (1866)" authorityName="Philippi" authorityYear="1866" box="[902,1198,799,826]" class="Insecta" family="Lymexylidae" genus="Fusicornis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
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<emphasis id="8DE8D232A0045D78FC579667769EFCA8" box="[902,1022,800,825]" italics="true" pageId="12" pageNumber="223">Fusicornis</emphasis>
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<bibRefCitation id="DB0D73D1A0045D78FBD2965871CEFCAB" author="Philippi, R. A." box="[1027,1198,799,826]" pageId="12" pageNumber="223" pagination="109 - 117" refId="ref29150" refString="Philippi, R. A. (1866) Einige Insekten von Chile. Stettiner Entomologische Zeitung, 27, 109 - 117." type="journal article" year="1866">Philippi (1866)</bibRefCitation>
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</taxonomicName>
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, with a single species from
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<collectingCountry id="C78B4EB0A0045D78FF0496047475FCCC" box="[213,277,835,861]" name="Chile" pageId="12" pageNumber="223">Chile</collectingCountry>
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;
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<taxonomicName id="789C75A3A0045D78FEF3960477E5FCCF" authority="Kurosawa (1985)" authorityName="Kurosawa" authorityYear="1985" box="[290,645,835,862]" class="Insecta" family="Lymexylidae" genus="Hymaloxylon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="8DE8D232A0045D78FEF3960474D7FCCC" box="[290,439,835,861]" italics="true" pageId="12" pageNumber="223">Hymaloxylon</emphasis>
|
||
|
|
<bibRefCitation id="DB0D73D1A0045D78FE6E960377E5FCCF" author="Kurosawa, Y." box="[447,645,835,862]" pageId="12" pageNumber="223" pagination="109 - 119" refId="ref28184" refString="Kurosawa, Y. (1985) Revisional notes on the family Lymexylonidae (Coleoptera) in eastern and southeastern Asia. Bulletin of the National Science Museum, Tokyo, (A), 11, 109 - 119." type="journal article" year="1985">Kurosawa (1985)</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
, with two species from
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FC45960476B0FCCC" box="[916,976,835,861]" name="India" pageId="12" pageNumber="223">India</collectingCountry>
|
||
|
|
,
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FC0D9604717FFCCC" box="[988,1055,835,861]" name="Nepal" pageId="12" pageNumber="223">Nepal</collectingCountry>
|
||
|
|
and western
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FB679604719CFCCC" box="[1206,1276,835,861]" name="China" pageId="12" pageNumber="223">China</collectingCountry>
|
||
|
|
;
|
||
|
|
<taxonomicName id="789C75A3A0045D78FAD99603743DFC13" authority="Kurasawa (1985)" authorityName="Kurasawa" authorityYear="1985" class="Insecta" family="Lymexylidae" genus="Raractocetus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="8DE8D232A0045D78FAD9960370FCFCCC" box="[1288,1436,836,861]" italics="true" pageId="12" pageNumber="223">Raractocetus</emphasis>
|
||
|
|
Kurasawa (1985)
|
||
|
|
</taxonomicName>
|
||
|
|
, with two species from
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FDA2962077CFFC10" box="[627,687,871,897]" name="India" pageId="12" pageNumber="223">India</collectingCountry>
|
||
|
|
, the
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FD3996207609FC10" box="[744,873,871,897]" name="Philippines" pageId="12" pageNumber="223">Philippines</collectingCountry>
|
||
|
|
,
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FCA4962076AAFC10" box="[885,970,871,897]" name="Taiwan" pageId="12" pageNumber="223">Taiwan</collectingCountry>
|
||
|
|
, Southeast Asia,
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FB419620719DFC10" box="[1168,1277,871,897]" name="Indonesia" pageId="12" pageNumber="223">Indonesia</collectingCountry>
|
||
|
|
and
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FAE7962070FDFC10" box="[1334,1437,871,897]" name="Australia" pageId="12" pageNumber="223">Australia</collectingCountry>
|
||
|
|
(introduced with
|
||
|
|
<taxonomicName id="789C75A3A0045D78FE8696CC74B3FC34" box="[343,467,907,933]" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" kingdom="Plantae" order="Myrtales" pageId="12" pageNumber="223" phylum="Tracheophyta" rank="genus">
|
||
|
|
<emphasis id="8DE8D232A0045D78FE8696CC74B3FC34" box="[343,467,907,933]" italics="true" pageId="12" pageNumber="223">Eucalyptus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
into
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FDDA96CB77C1FC34" box="[523,673,907,934]" name="New Zealand" pageId="12" pageNumber="223">New Zealand</collectingCountry>
|
||
|
|
and
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FD0796CC7677FC37" box="[726,791,907,934]" name="Chile" pageId="12" pageNumber="223">Chile</collectingCountry>
|
||
|
|
); and
|
||
|
|
<taxonomicName id="789C75A3A0045D78FC8896CB715CFC34" authority="Paulus (2004)" authorityName="Paulus" authorityYear="2004" box="[857,1084,907,933]" class="Insecta" family="Lymexylidae" genus="Urtea" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="8DE8D232A0045D78FC8896CB76FAFC34" box="[857,922,908,933]" italics="true" pageId="12" pageNumber="223">Urtea</emphasis>
|
||
|
|
<bibRefCitation id="DB0D73D1A0045D78FC7196CC715CFC34" author="Paulus, H. F." box="[928,1084,907,933]" pageId="12" pageNumber="223" pagination="277 - 290" refId="ref29085" refString="Paulus, H. F. (2004) Urtrea graeca nov. gen. et nov. spec., der erste Vertreter der tropischen Atractocerinae in Europa sowie eine Beschreibung von Homaloxylon aspoecki nov. spec. aus Yunnan (China) (Coleoptera, Cucujiformia, Lymexylidae, Atractocerinae nov. status). Denisia, 13, 277 - 290." type="journal article" year="2004">Paulus (2004)</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
with one species from northern
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FF4696E8758AFC58" box="[151,234,943,969]" name="Greece" pageId="12" pageNumber="223">Greece</collectingCountry>
|
||
|
|
. In addition, the following fossil
|
||
|
|
<taxonomicName id="789C75A3A0045D78FDA296E87678FC58" authorityName="Laporte" authorityYear="1840" box="[627,792,943,969]" class="Insecta" family="Lymexylidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="subFamily" subFamily="Atractocerinae">Atractocerinae</taxonomicName>
|
||
|
|
have been reported:
|
||
|
|
<emphasis id="8DE8D232A0045D78FBC396F77031FC58" box="[1042,1361,943,970]" italics="true" pageId="12" pageNumber="223">Cretoatractocerus grimaldii</emphasis>
|
||
|
|
WolfSchwenninger (2011), the earliest member of the group from the Lower Cretaceous (Upper Aptian) Crato Formation of
|
||
|
|
<collectingCountry id="C78B4EB0A0045D78FF3996B0744FFB80" box="[232,303,1015,1041]" name="Brazil" pageId="12" pageNumber="223">Brazil</collectingCountry>
|
||
|
|
, plus
|
||
|
|
<emphasis id="8DE8D232A0045D78FEA196BF77FFFB80" box="[368,671,1015,1041]" italics="true" pageId="12" pageNumber="223">Vetatractocerus burmiticus</emphasis>
|
||
|
|
Yamamoto,(2019),
|
||
|
|
<taxonomicName id="789C75A3A0045D78FCAE96BF702EFB80" authority="Yamamoto (2019)" authorityName="Yamamoto" authorityYear="2019" box="[895,1358,1015,1042]" class="Insecta" family="Lymexylidae" genus="Raractocetus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="species" species="extinctus">
|
||
|
|
<emphasis id="8DE8D232A0045D78FCAE96BF711DFB80" box="[895,1149,1016,1041]" italics="true" pageId="12" pageNumber="223">Raractocetus extinctus</emphasis>
|
||
|
|
<bibRefCitation id="DB0D73D1A0045D78FB5296BF702EFB80" author="Yamamoto, S." box="[1155,1358,1015,1042]" pageId="12" pageNumber="223" pagination="1 - 10" refId="ref29555" refString="Yamamoto, S. (2019) Fossil evidence of elytral reduction in ship-timber beetles. Nature, Scientific Reports, 9 (4938), 1 - 10. https: // doi. org / 10.1038 / s 41598 - 019 - 41310 - 1" type="journal article" year="2019">Yamamoto (2019)</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName id="789C75A3A0045D78FA5596BF74D8FBA4" authority="Yamamoto (2019)" authorityName="Yamamoto" authorityYear="2019" class="Insecta" family="Lymexylidae" genus="Raractocetus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="species" species="fossilis">
|
||
|
|
<emphasis id="8DE8D232A0045D78FA5596BF7585FBA4" italics="true" pageId="12" pageNumber="223">R. fossilis</emphasis>
|
||
|
|
<bibRefCitation id="DB0D73D1A0045D78FF3D915B74D8FBA4" author="Yamamoto, S." box="[236,440,1051,1078]" pageId="12" pageNumber="223" pagination="1 - 10" refId="ref29555" refString="Yamamoto, S. (2019) Fossil evidence of elytral reduction in ship-timber beetles. Nature, Scientific Reports, 9 (4938), 1 - 10. https: // doi. org / 10.1038 / s 41598 - 019 - 41310 - 1" type="journal article" year="2019">Yamamoto (2019)</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
from the mid-Cretaceous Burmese amber (earliest Cenomanian),
|
||
|
|
<taxonomicName id="789C75A3A0045D78FB72915B7582FBC8" authority="Yamamoto (2019)" authorityName="Yamamoto" authorityYear="2019" class="Insecta" family="Lymexylidae" genus="Raractocetus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="species" species="balticus">
|
||
|
|
<emphasis id="8DE8D232A0045D78FB72915B707CFBA4" box="[1187,1308,1051,1077]" italics="true" pageId="12" pageNumber="223">R. balticus</emphasis>
|
||
|
|
<bibRefCitation id="DB0D73D1A0045D78FAF2915B7582FBC8" author="Yamamoto, S." pageId="12" pageNumber="223" pagination="1 - 10" refId="ref29555" refString="Yamamoto, S. (2019) Fossil evidence of elytral reduction in ship-timber beetles. Nature, Scientific Reports, 9 (4938), 1 - 10. https: // doi. org / 10.1038 / s 41598 - 019 - 41310 - 1" type="journal article" year="2019">Yamamoto (2019)</bibRefCitation>
|
||
|
|
</taxonomicName>
|
||
|
|
from mid-Eocene Baltic amber and
|
||
|
|
<taxonomicName id="789C75A3A0045D78FDAA9178766CFBC8" authorityName="Palisot de Beauvois" authorityYear="1801" box="[635,780,1087,1113]" class="Insecta" family="Lymexylidae" genus="Atractocerus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="223" phylum="Arthropoda" rank="genus">
|
||
|
|
<emphasis id="8DE8D232A0045D78FDAA9178766CFBC8" box="[635,780,1087,1113]" italics="true" pageId="12" pageNumber="223">Atractocerus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
sp. from Miocene Dominican amber (
|
||
|
|
<bibRefCitation id="DB0D73D1A0045D78FB699178701EFBC8" author="Yamamoto, S." box="[1208,1406,1087,1113]" pageId="12" pageNumber="223" pagination="1 - 10" refId="ref29555" refString="Yamamoto, S. (2019) Fossil evidence of elytral reduction in ship-timber beetles. Nature, Scientific Reports, 9 (4938), 1 - 10. https: // doi. org / 10.1038 / s 41598 - 019 - 41310 - 1" type="journal article" year="2019">Yamamoto, 2019</bibRefCitation>
|
||
|
|
).
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|