282 lines
36 KiB
XML
282 lines
36 KiB
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<document id="8458783A5566156BE4B21692B5B58544" ID-CLB-Dataset="3085" ID-DOI="10.15553/c2016v711a17" ID-GBIF-Dataset="c3268c84-33ed-4d8f-9fb6-cb50dbbde93e" ID-ISSN="2235-3658" ID-Zenodo-Dep="5721447" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637683481210" checkinUser="carolina" docAuthor="Rakotondrainibe, France & Jouy, Alain" docDate="2016" docId="03AA99569501CB29FCD31542FE74F945" docLanguage="fr" docName="Candollea.71.1.143-148.pdf" docOrigin="Candollea 71 (1)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Selaginella rasoloheryi Rakotondr. 2016, spec. nova" docType="treatment" docVersion="9" lastPageNumber="146" masterDocId="FF93E12E9500CB2AFFF6175FFFE2FFD4" masterDocTitle="Une espèce nouvelle et une synonymie nouvelle dans le genre Selaginella P. Beauv. (Lycopodiophyta, Selaginellaceae) à Madagascar" masterLastPageNumber="148" masterPageNumber="143" pageNumber="144" updateTime="1699469561603" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="B0D3E8E7B275666C5656C1AD05CD81FB">
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<mods:title id="23C123257F83970BA7F8D013E78D75E1">Une espèce nouvelle et une synonymie nouvelle dans le genre Selaginella P. Beauv. (Lycopodiophyta, Selaginellaceae) à Madagascar</mods:title>
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</mods:titleInfo>
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<mods:name id="A39B0FA4B03EA35FD8B6BC0118DAE48B" type="personal">
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<mods:roleTerm id="3C54141458A240BA50999CA3202EA405">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D14F0C2941648132E5A04DECCB76608E">Rakotondrainibe, France</mods:namePart>
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<mods:affiliation id="9CEF6D9E2E79862905E5F38710E958F2">Institut de Systématique, Evolution et Biodiversité (UMR 7205 – CNRS MNHN UPMC EPHE), Muséum national d’Histoire naturelle, rue Cuvier 57, C. P. 39, 7531 Paris CEDEX 05, France.</mods:affiliation>
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<mods:nameIdentifier id="4A5D0694A176E4DA2693AAF443D52F4D" type="email">rakotond@mnhn.fr</mods:nameIdentifier>
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</mods:name>
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</mods:role>
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<mods:namePart id="9CB254CA7A0D19A88FEFDB543535D54E">Jouy, Alain</mods:namePart>
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<mods:affiliation id="0228922414D1AFD1B513B3293071B290">avenue de la Bourdonnais 36, 75007 Paris, France.</mods:affiliation>
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<mods:typeOfResource id="4DFC8D339964C223B267FF2E434747FD">text</mods:typeOfResource>
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<mods:title id="9D35AC207EA34FE871C4C7D978D135C8">Candollea</mods:title>
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</mods:titleInfo>
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<mods:part id="16840AA4BC7086F9DF10ACFC6CB182C6">
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<mods:date id="77ACFB0B9614DCEA6ABCBA51F3FC443E">2016</mods:date>
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<mods:number id="1D9ED8BF0C6ACB19D4D36AB850473FBC">2016-04-29</mods:number>
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</mods:detail>
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<mods:number id="7F4F19CF0C5FFCC36A50EA81F757A536">71</mods:number>
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</mods:detail>
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<mods:detail id="8BA91FDF2FF0F623835C630A1629385E" type="issue">
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<mods:number id="726CF23D292C6F74B8D01D4D1F6101CD">1</mods:number>
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<mods:classification id="762608A431DF2E8EF4A90CC003E3F9D3">journal article</mods:classification>
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<mods:identifier id="523AA8FA6372D5F5A89A00D9662D5144" type="DOI">10.15553/c2016v711a17</mods:identifier>
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<treatment id="03AA99569501CB29FCD31542FE74F945" ID-DOI="http://doi.org/10.5281/zenodo.5764256" ID-GBIF-Taxon="191197849" ID-Zenodo-Dep="5764256" LSID="urn:lsid:plazi:treatment:03AA99569501CB29FCD31542FE74F945" httpUri="http://treatment.plazi.org/id/03AA99569501CB29FCD31542FE74F945" lastPageId="3" lastPageNumber="146" pageId="1" pageNumber="144">
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<subSubSection id="C3197BCB9501CB2BFCD31542FA83FDE2" box="[805,1377,541,566]" pageId="1" pageNumber="144" type="nomenclature">
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<paragraph id="8BBC28409501CB2BFCD31542FA83FDE2" blockId="1.[805,1377,541,566]" box="[805,1377,541,566]" pageId="1" pageNumber="144">
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<heading id="D0F49F2C9501CB2BFCD31542FA83FDE2" box="[805,1377,541,566]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="144" reason="2">
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<taxonomicName id="4C0353C39501CB2BFCD31542FB73FDE1" ID-CoL="6YC45" authority="Rakotondr." authorityName="Rakotondr." authorityYear="2016" box="[805,1169,541,565]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="144" phylum="Tracheophyta" rank="species" species="rasoloheryi" status="sp. nov.">
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<emphasis id="B977F4529501CB2BFCD31542FBF3FDE1" bold="true" box="[805,1041,541,565]" italics="true" pageId="1" pageNumber="144">Selaginella rasoloheryi</emphasis>
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Rakotondr.
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</taxonomicName>
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,
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<emphasis id="B977F4529501CB2BFB6D1541FAEFFDE2" bold="true" box="[1179,1293,542,566]" pageId="1" pageNumber="144">
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<taxonomicNameLabel id="A24449299501CB2BFB6D1541FAEFFDE2" box="[1179,1293,542,566]" pageId="1" pageNumber="144" rank="species">spec. nova</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="133834C59501CB2BFAED1542FAB3FDE2" box="[1307,1361,541,566]" captionStart="Fig" captionStartId="2.[173,200,1859,1878]" captionTargetBox="[170,1454,314,1770]" captionTargetId="figure-80@2.[151,1469,952,1782]" captionTargetPageId="2" captionText="Fig. 1. – Selaginella rasoloheryi Rakotondr. A. Plante entière; B. Détail d’une partie stérile de la tige feuillée, face dorsale, avec les feuilles médianes de petites taille, les feuilles latérales plus grandes et une feuille axillaire à la base de la ramification de la tige; C. Strobile. [Rakotondrainibe & Rasolohery 6425, P] [Dessin: A. Jouy]" figureDoi="http://doi.org/10.5281/zenodo.5764222" httpUri="https://zenodo.org/record/5764222/files/figure.png" pageId="1" pageNumber="144">fig. 1</figureCitation>
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).
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3197BCB9501CB2BFCBC1516FBD0FD36" pageId="1" pageNumber="144" type="materials_examined">
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<paragraph id="8BBC28409501CB2BFCBC1516FBD0FD36" blockId="1.[842,1456,584,738]" pageId="1" pageNumber="144">
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<materialsCitation id="3B6B221D9501CB2BFCBC1516FBCCFD36" ID-GBIF-Occurrence="3415109302" collectingDate="2001-10-28" collectionCode="P, TAN" collectorName="de Marojejy & Rakotondrainibe & Rasolohery" country="Madagascar" elevation="1150" latitude="-14.4366665" location="Prov" longLatPrecision="21" longitude="49.62" pageId="1" pageNumber="144" specimenCode="P00248501, P01313265" specimenCount="1" typeStatus="holotype">
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<emphasis id="B977F4529501CB2BFCBC1516FAA1FDB5" bold="true" box="[842,1347,584,609]" pageId="1" pageNumber="144">
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<typeStatus id="54B896E29501CB2BFCBC1516FC71FDB5" box="[842,915,585,609]" pageId="1" pageNumber="144">Typus</typeStatus>
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:
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<collectingCountry id="F31468D09501CB2BFC511516FBACFD8B" box="[935,1102,585,609]" name="Madagascar" pageId="1" pageNumber="144">MADAGASCAR</collectingCountry>
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.
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<location id="8EDC7E9B9501CB2BFBAB1516FB74FDB5" LSID="urn:lsid:plazi:treatment:03AA99569501CB29FCD31542FE74F945:8EDC7E9B9501CB2BFBAB1516FB74FDB5" box="[1117,1174,585,609]" country="Madagascar" name="Prov" pageId="1" pageNumber="144" stateProvince="Typus">Prov</location>
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.
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<collectingRegion id="49C7E6A29501CB2BFB551517FAD5FDB4" box="[1187,1335,584,608]" country="Madagascar" name="Antsiranana" pageId="1" pageNumber="144">Antsiranana</collectingRegion>
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:
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</emphasis>
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<location id="8EDC7E9B9501CB2BFAB81517FA4EFDB4" LSID="urn:lsid:plazi:treatment:03AA99569501CB29FCD31542FE74F945:8EDC7E9B9501CB2BFAB81517FA4EFDB4" box="[1358,1452,584,608]" country="Madagascar" latitude="-14.4366665" longLatPrecision="21" longitude="49.62" name="Andapa" pageId="1" pageNumber="144" stateProvince="Antsiranana">Andapa</location>
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, réserve
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<collectorName id="26F64D969501CB2BFC6D1536FBF8FD55" box="[923,1050,617,641]" pageId="1" pageNumber="144">de Marojejy</collectorName>
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, à
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<quantity id="4CFB85A59501CB2BFBC01536FB98FD55" box="[1078,1146,617,641]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="144" unit="km" value="13.0">13 km</quantity>
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au
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<locationDeviation id="9A0E31629501CB2BFB561536FACDFD55" box="[1184,1327,617,641]" location="Doany" pageId="1" pageNumber="144">SE de Doany</locationDeviation>
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,
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<geoCoordinate id="EE374E879501CB2BFACF1536FA4DFD55" box="[1337,1455,617,641]" degrees="14" direction="south" minutes="26" orientation="latitude" pageId="1" pageNumber="144" precision="15" seconds="12" value="-14.4366665">14°26’12’’S</geoCoordinate>
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<geoCoordinate id="EE374E879501CB2BFCBC15D6FC32FD75" box="[842,976,649,673]" degrees="49" direction="east" minutes="37" orientation="longitude" pageId="1" pageNumber="144" precision="15" seconds="12" value="49.62">49°37’12’’E</geoCoordinate>
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,
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<quantity id="4CFB85A59501CB2BFC1615D6FBD9FD75" box="[992,1083,649,673]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="144" unit="m" value="1150.0">
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<elevation id="002ECF739501CB2BFC1615D6FBD9FD75" box="[992,1083,649,673]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="1" pageNumber="144" unit="m" value="1150.0">1150 m</elevation>
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</quantity>
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,
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<date id="FFBD0E809501CB2BFBBC15D6FB24FD75" box="[1098,1222,649,673]" pageId="1" pageNumber="144" value="2001-10-28">
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<collectingDate id="EFF9F7689501CB2BFBBC15D6FB24FD75" box="[1098,1222,649,673]" pageId="1" pageNumber="144" value="2001-10-28">28.X.2001</collectingDate>
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</date>
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,
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<emphasis id="B977F4529501CB2BFB2315D6FBE9FD16" italics="true" pageId="1" pageNumber="144">
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<collectorName id="26F64D969501CB2BFB2315D6FA6CFD75" box="[1237,1422,649,673]" pageId="1" pageNumber="144">Rakotondrainibe</collectorName>
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&
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<collectorName id="26F64D969501CB2BFCBC15F5FC27FD16" box="[842,965,682,706]" pageId="1" pageNumber="144">Rasolohery</collectorName>
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6425
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</emphasis>
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(holo-:
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<collectionCode id="ED12B0859501CB2BFB8D15F5FB69FD15" box="[1147,1163,682,705]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="144" type="Herbarium">P</collectionCode>
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[
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<specimenCode id="DBA5803B9501CB2BFB6915F5FAD6FD16" box="[1183,1332,682,706]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="144" type="Herbarium">P00248501</specimenCode>
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]!; iso-:
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<collectionCode id="ED12B0859501CB2BFA6815F5FA4FFD15" box="[1438,1453,682,705]" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="144" type="Herbarium">P</collectionCode>
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[
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<specimenCode id="DBA5803B9501CB2BFCA71595FC34FD36" box="[849,982,714,738]" collectionCode="P" country="France" lsid="urn:lsid:biocol.org:col:15763" name="Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants" pageId="1" pageNumber="144" type="Herbarium">P01313265</specimenCode>
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]!,
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<collectionCode id="ED12B0859501CB2BFC1F1595FBC4FD36" box="[1001,1062,714,738]" country="Madagascar" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13842" lsid="urn:lsid:biocol.org:col:13842" name="Parc de Tsimbazaza" pageId="1" pageNumber="144" type="Herbarium">TAN</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3197BCB9501CB29FCBC15AAFE39FE66" lastPageId="3" lastPageNumber="146" pageId="1" pageNumber="144" type="description">
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<paragraph id="8BBC28409501CB2BFCBC15AAFAD8FC24" blockId="1.[842,1456,757,1008]" pageId="1" pageNumber="144">
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<emphasis id="B977F4529501CB2BFCBC15AAFAD8FC24" italics="true" pageId="1" pageNumber="144">
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With its sterile leaves and sporophylls distinctly dimorphic,
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<taxonomicName id="4C0353C39501CB2BFCBD1449FB22FCFA" authority="Rakotondr." authorityName="Rakotondr." box="[843,1216,790,814]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="144" phylum="Tracheophyta" rank="species" species="rasoloheryi">Selaginella rasoloheryi Rakotondr.</taxonomicName>
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belongs to
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<taxonomicName id="4C0353C39501CB2BFACF1449FA4CFCFA" box="[1337,1454,790,814]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="144" phylum="Tracheophyta" rank="genus">Selaginella</taxonomicName>
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subgenus
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<taxonomicName id="4C0353C39501CB2BFC5D1469FB26FC99" authority="Baker. Among" authorityName="Baker. Among" box="[939,1220,822,846]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="1" pageNumber="144" phylum="Tracheophyta" rank="subGenus" subGenus="Heterostachys">Heterostachys Baker. Among</taxonomicName>
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the species belonging to this subgenus, the combination of the following morphological characters is original: stem prostrate, creeping, up to
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<quantity id="4CFB85A59501CB2BFA9C1428FA52FC5B" box="[1386,1456,887,911]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="1" pageNumber="144" unit="cm" value="5.5">5,5 cm</quantity>
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long, not articulate, without rhizome, with short, not branched ramifications; rhizophores not branched, ciliform, up to
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<quantity id="4CFB85A59501CB2BFA9014E7FA48FC04" box="[1382,1450,952,976]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="144" unit="cm" value="1.7">1,7 cm</quantity>
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, in branch axils throughout the length of the plant.
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</emphasis>
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</paragraph>
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<paragraph id="8BBC28409501CB2BFCBC1346FA9DF849" blockId="1.[804,1456,1047,1949]" pageId="1" pageNumber="144">
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<emphasis id="B977F4529501CB2BFCBC1346FC6CFBE5" box="[842,910,1049,1073]" italics="true" pageId="1" pageNumber="144">Plante</emphasis>
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|
|
prostr, de petite taille, dépourvue de rhizome, de sobole et de tubercule.
|
|||
|
|
<emphasis id="B977F4529501CB2BFBBE1365FB05FB86" box="[1096,1255,1082,1106]" italics="true" pageId="1" pageNumber="144">Tige principale</emphasis>
|
|||
|
|
rampante, longue de
|
|||
|
|
<quantity id="4CFB85A59501CB2BFCB01305FC5DFBA5" box="[838,959,1114,1138]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="1" pageNumber="144" unit="cm" value="5.0" valueMax="5.5" valueMin="4.5">4,5-5,5 cm</quantity>
|
|||
|
|
, non articulée, ramifiée sur toute sa longueur; 7-8 ramifications simples, espacées de
|
|||
|
|
<quantity id="4CFB85A59501CB2BFB3D1324FACDFB46" box="[1227,1327,1147,1171]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="1" pageNumber="144" unit="cm" value="0.65" valueMax="1.0" valueMin="0.3">0,3-1 cm</quantity>
|
|||
|
|
, longues de
|
|||
|
|
<quantity id="4CFB85A59501CB2BFCD313C4FC7AFB66" box="[805,920,1179,1203]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" metricValueMax="2.2" metricValueMin="0.2" pageId="1" pageNumber="144" unit="cm" value="1.2" valueMax="2.2" valueMin="0.2">0,2-2,2 cm</quantity>
|
|||
|
|
; ensemble de la tige principale et des ramifications à contour ± linéaire.
|
|||
|
|
<emphasis id="B977F4529501CB2BFBF513E4FB63FB07" box="[1027,1153,1211,1235]" italics="true" pageId="1" pageNumber="144">Rhizophores</emphasis>
|
|||
|
|
situés sur toute la longueur de la tige principale, jaune paille, simples, ciliformes, env.
|
|||
|
|
<quantity id="4CFB85A59501CB2BFCD213A3FC95FAC0" box="[804,887,1276,1300]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="144" unit="mm" value="0.1">0,1 mm</quantity>
|
|||
|
|
de diamètre, longs de
|
|||
|
|
<quantity id="4CFB85A59501CB2BFB9213A3FB2AFAC0" box="[1124,1224,1276,1300]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.7" metricValueMin="0.8" pageId="1" pageNumber="144" unit="mm" value="12.5" valueMax="17.0" valueMin="8.0">8-17 mm</quantity>
|
|||
|
|
, naissant à la base des ramifications de la tige, face ventrale, à l’aisselle des feuilles axillaires.
|
|||
|
|
<emphasis id="B977F4529501CB2BFC781262FC39FA81" box="[910,987,1341,1365]" italics="true" pageId="1" pageNumber="144">Feuilles</emphasis>
|
|||
|
|
dimorphes, membraneuses, vert clair sur le frais, espacées sur la tige principale et contiguës ou légèrement imbriquées à l’extrémité des tiges avec une nervure médiane non apparente;
|
|||
|
|
<emphasis id="B977F4529501CB2BFC2612C1FB90FA62" box="[976,1138,1438,1462]" italics="true" pageId="1" pageNumber="144">feuilles latérales</emphasis>
|
|||
|
|
inéquilatérales, de 1,3-1,6 ×
|
|||
|
|
<quantity id="4CFB85A59501CB2BFCD312E0FC45FA02" box="[805,935,1471,1495]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="1" pageNumber="144" unit="mm" value="0.7" valueMax="0.8" valueMin="0.6">0,6-0,8 mm</quantity>
|
|||
|
|
, à apex aigu à courtement acuminé, base obtuse, marge entière à denticulée, la moitié acroscopique semielliptique, la moitié basiscopique oblongue;
|
|||
|
|
<emphasis id="B977F4529501CB2BFAF212A0FA4DF9C3" box="[1284,1455,1535,1559]" italics="true" pageId="1" pageNumber="144">feuilles axillaires</emphasis>
|
|||
|
|
semblables aux latérales mais équilatérales;
|
|||
|
|
<emphasis id="B977F4529501CB2BFAF5117FFA4CF9EC" box="[1283,1454,1568,1592]" italics="true" pageId="1" pageNumber="144">feuilles médianes</emphasis>
|
|||
|
|
subéquilatérales, ovées, de 0,6-0,8 ×
|
|||
|
|
<quantity id="4CFB85A59501CB2BFB29111EFA8CF98C" box="[1247,1390,1601,1625]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="144" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0,2-0,3 mm</quantity>
|
|||
|
|
, base obtuse à subcor擩e, marge dentée-ciliée, apex brusquement et longuement acuminé, acumen aussi long que le limbe ou 2/3 de sa longueur.
|
|||
|
|
<emphasis id="B977F4529501CB2BFBDE11FDFB60F96E" box="[1064,1154,1698,1722]" italics="true" pageId="1" pageNumber="144">Strobiles</emphasis>
|
|||
|
|
solitaires, à l’extrémité des ramifications, résupinés, longs de 1,2-3,0 mm; sporophylles dimorphes:
|
|||
|
|
<emphasis id="B977F4529501CB2BFC5011BDFBFAF92E" box="[934,1048,1762,1786]" italics="true" pageId="1" pageNumber="144">les latérales</emphasis>
|
|||
|
|
, formant un angle droit avec l’axe du strobile, ovées-lancéolées, de 1,0-1,2 ×
|
|||
|
|
<quantity id="4CFB85A59501CB2BFB2E105CFABCF8CE" box="[1240,1374,1795,1819]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="1" pageNumber="144" unit="mm" value="0.4" valueMax="0.5" valueMin="0.3">0,3-0,5 mm</quantity>
|
|||
|
|
, à apex aigu à courtement acuminé, base obtuse, marge lâchement denticulée, cars, à carène étroite, finement dentée;
|
|||
|
|
<emphasis id="B977F4529501CB2BFA61101BFC63F8A8" italics="true" pageId="1" pageNumber="144">les médianes</emphasis>
|
|||
|
|
ovées, de 1,0-1,2 (acumen compris) ×
|
|||
|
|
<quantity id="4CFB85A59501CB2BFADE103BFA48F8A8" box="[1320,1450,1892,1916]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="1" pageNumber="144" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0,3-0,4 mm</quantity>
|
|||
|
|
, non cars, apex brusquement et longuement acuminé, base
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF7C78C89502CB28FF5B101CFD69F85D" ID-DOI="http://doi.org/10.5281/zenodo.5764222" ID-Zenodo-Dep="5764222" httpUri="https://zenodo.org/record/5764222/files/figure.png" pageId="2" pageNumber="145" startId="2.[173,200,1859,1878]" targetBox="[170,1454,314,1770]" targetPageId="2">
|
|||
|
|
<paragraph id="8BBC28409502CB28FF5B101CFD69F85D" blockId="2.[173,1376,1859,1929]" pageId="2" pageNumber="145">
|
|||
|
|
<emphasis id="B977F4529502CB28FF5B101CFF0FF882" bold="true" box="[173,237,1859,1878]" pageId="2" pageNumber="145">Fig. 1. –</emphasis>
|
|||
|
|
<taxonomicName id="4C0353C39502CB28FF07101CFD96F882" authority="Rakotondr. A. Plante" box="[241,628,1859,1878]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="2" pageNumber="145" phylum="Tracheophyta" rank="species" species="rasoloheryi">
|
|||
|
|
<emphasis id="B977F4529502CB28FF07101CFE5BF882" box="[241,441,1859,1878]" italics="true" pageId="2" pageNumber="145">Selaginella rasoloheryi</emphasis>
|
|||
|
|
Rakotondr.
|
|||
|
|
<emphasis id="B977F4529502CB28FDD2101CFDD5F882" bold="true" box="[548,567,1859,1878]" pageId="2" pageNumber="145">A.</emphasis>
|
|||
|
|
Plante
|
|||
|
|
</taxonomicName>
|
|||
|
|
entière;
|
|||
|
|
<emphasis id="B977F4529502CB28FD35101CFD37F882" bold="true" box="[707,725,1859,1878]" pageId="2" pageNumber="145">B.</emphasis>
|
|||
|
|
Détail d’une partie stérile de la tige feuillée, face dorsale, avec les feuilles médianes de petites taille, les feuilles latérales plus grandes et une feuille axillaire à la base de la ramification de la tige;
|
|||
|
|
<emphasis id="B977F4529502CB28FB201003FB0BF8BB" bold="true" box="[1238,1257,1884,1903]" pageId="2" pageNumber="145">C.</emphasis>
|
|||
|
|
Strobile. [
|
|||
|
|
<emphasis id="B977F4529502CB28FF441029FE37F85D" box="[178,469,1910,1929]" italics="true" pageId="2" pageNumber="145">Rakotondrainibe & Rasolohery 6425</emphasis>
|
|||
|
|
, P] [Dessin: A. Jouy]
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BBC28409503CB29FF821645FE39FE66" blockId="3.[116,767,282,434]" pageId="3" pageNumber="146">obtuse, marge ciliée; 2-3 macrosporanges à la base du strobile, contenant chacun 4 macrospores blanches, globuleuses, trilètes, à paroi lisse à finement rugueuse-verruqueuse, non réticulée; microsporanges distaux contenant de nombreuses microspores rouge-orangé, tétrariques, trilètes.</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3197BCB9503CB29FF6C1685FDF1FD86" pageId="3" pageNumber="146" type="etymology">
|
|||
|
|
<paragraph id="8BBC28409503CB29FF6C1685FDF1FD86" blockId="3.[116,768,474,594]" pageId="3" pageNumber="146">
|
|||
|
|
<emphasis id="B977F4529503CB29FF6C1685FEFDFE26" box="[154,287,474,498]" italics="true" pageId="3" pageNumber="146">Etymologie.</emphasis>
|
|||
|
|
– L’épithète spécifique a été choisie en l’honneur d’Andriambolantsoa Rasolohery, botaniste malgache, qui a participé à l’expédition dans le Marojejy durant laquelle l’espèce nouvelle a été récoltée.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3197BCB9503CB29FF6C1525FD47FCE6" pageId="3" pageNumber="146" type="distribution">
|
|||
|
|
<paragraph id="8BBC28409503CB29FF6C1525FD47FCE6" blockId="3.[116,768,634,818]" pageId="3" pageNumber="146">
|
|||
|
|
<emphasis id="B977F4529503CB29FF6C1525FDDFFD46" box="[154,573,634,658]" italics="true" pageId="3" pageNumber="146">Distribution géographique et écologie</emphasis>
|
|||
|
|
. – Cette espèce endémique de
|
|||
|
|
<collectingCountry id="F31468D09503CB29FEED15C5FE43FD66" box="[283,417,666,690]" name="Madagascar" pageId="3" pageNumber="146">Madagascar</collectingCountry>
|
|||
|
|
n’est connue que par une seule récolte en provenance de la partie occidentale du massif de Marojejy (Nord-Est) où elle a⃩té observée dans le sous-bois de la forêt sempervirente, à
|
|||
|
|
<quantity id="4CFB85A59503CB29FE6815A5FE0DFCC5" box="[414,495,762,786]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="3" pageNumber="146" unit="m" value="1150.0">1150 m</quantity>
|
|||
|
|
d’altitude, en tapis dense sur le sommet d’un rocher, dans une vallée encaissée.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3197BCB9503CB29FF6C1404FE74F945" pageId="3" pageNumber="146" type="discussion">
|
|||
|
|
<paragraph id="8BBC28409503CB29FF6C1404FE74F945" blockId="3.[116,769,856,1682]" pageId="3" pageNumber="146">
|
|||
|
|
<emphasis id="B977F4529503CB29FF6C1404FF33FCA6" box="[154,209,859,882]" italics="true" pageId="3" pageNumber="146">Notes</emphasis>
|
|||
|
|
. – La monophylie des
|
|||
|
|
<taxonomicName id="4C0353C39503CB29FE341405FDB4FCA6" box="[450,598,858,882]" class="Lycopodiopsida" family="Selaginellaceae" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="146" phylum="Tracheophyta" rank="family">
|
|||
|
|
<emphasis id="B977F4529503CB29FE341405FDB4FCA6" box="[450,598,858,882]" italics="true" pageId="3" pageNumber="146">Selaginellaceae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
a confir淩e par plusieurs analyses moléculaires (
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FDE11425FD16FC46" author="Korall, P. & P. Kenrick & P. Therrien" box="[535,756,890,914]" journalOrPublisher="Int. J. Plant Sci." pageId="3" pageNumber="146" pagination="585 - 594" part="160" refId="ref4298" refString="Korall, P., P. Kenrick & J. P. Therrien (1999). Phylogeny of Selaginellaceae, evaluation of generic / subgeneric relationships based on rbcL gene sequences. Int. J. Plant Sci. 160: 585 - 594." title="Phylogeny of Selaginellaceae, evaluation of generic / subgeneric relationships based on rbcL gene sequences" type="journal article" year="1999">Korall et al. 1999</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FF8214C5FE4DFC66" author="Wikstrom, N. & P. Kenrick" box="[116,431,922,946]" journalOrPublisher="Int. J. Plant Sci." pageId="3" pageNumber="146" pagination="862 - 871" part="158" refId="ref4657" refString="Wikstrom, N. & P. Kenrick (1997). Phylogeny of Lycopodiaceae (Lycopsida) and the relationships of Phylloglossum drumondii Kunze based on rbcL sequences. Int. J. Plant Sci. 158: 862 - 871." title="Phylogeny of Lycopodiaceae (Lycopsida) and the relationships of Phylloglossum drumondii Kunze based on rbcL sequences" type="journal article" year="1997">Wikström & Kenrick, 1997</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FE4F14C5FD91FC66" author="Zhou, X. - M. & C. J. Rothfels & L. Zhang & Z. - R. He & T. Le Pechon & H. He & N. T. Lu & R. Knapp & D. Lorence & X. - J. He & X. - F. Gao & L. - B. Zhang" box="[441,627,922,946]" journalOrPublisher="Cladistics" pageId="3" pageNumber="146" refId="ref4700" refString="Zhou, X. - M., C. J. Rothfels, L. Zhang, Z. - R. He, T. Le Pechon, H. He, N. T. Lu, R. Knapp, D. Lorence, X. - J. He, X. - F. Gao & L. - B. Zhang (2015). A large-scale phylogeny of the lycophyte genus Selaginella (Selaginellaceae: Lycopodiopsida) based on plastid and nuclear loci. Cladistics [http: // dx. doi. org / 10.1111 / cla. 12136]." title="A large-scale phylogeny of the lycophyte genus Selaginella (Selaginellaceae: Lycopodiopsida) based on plastid and nuclear loci" type="book" year="2015">Zhou & al. 2015</bibRefCitation>
|
|||
|
|
). La majorité des spécialistes s’accordent à reconnaître un seul genre dans la famille,
|
|||
|
|
<taxonomicName id="4C0353C39503CB29FF301485FED2FC26" box="[198,304,986,1010]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="146" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis id="B977F4529503CB29FF301485FED2FC26" box="[198,304,986,1010]" italics="true" pageId="3" pageNumber="146">Selaginella</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, subdivisé en quatre (
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FDFB1485FD70FC26" author="Baker, J. G." box="[525,658,986,1010]" journalOrPublisher="J. Bot." pageId="3" pageNumber="146" pagination="1 - 5" refId="ref4110" refString="Baker, J. G. (1883). A synopsis of the genus Selaginella. J. Bot. 1883: 1 - 5, 42 - 46, 80 - 84." title="A synopsis of the genus Selaginella" type="journal article" year="1883">Baker, 1883</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FD6A1485FECEFBC6" author="Walton, J. & A. H. G. Alston" editor="Verdoorn, F." journalOrPublisher="Nijhoff, The Hage" pageId="3" pageNumber="146" pagination="500 - 506" refId="ref4567" refString="Walton, J. & A. H. G. Alston (1938). Lycopodiinae. In: Verdoorn, F. (ed.), Manual of pteridology: 500 - 506. Nijhoff, The Hage." title="Lycopodiinae" type="book chapter" volumeTitle="Manual of pteridology" year="1938">Walton & Alston, 1938</bibRefCitation>
|
|||
|
|
), cinq (
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FE7614A5FDE4FBC6" author="Jermy, A. C." box="[384,518,1018,1042]" journalOrPublisher="Fern Gaz." pageId="3" pageNumber="146" pagination="2" part="13" refId="ref4226" refString="Jermy, A. C. (1986). Subgeneric names in Selaginella. Fern Gaz. 13 (2)." title="Subgeneric names in Selaginella" type="journal article" year="1986">Jermy, 1986</bibRefCitation>
|
|||
|
|
), six (
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FDB314A5FF49FBE6" author="Zhou, X-M. & L. - B. Zhang" journalOrPublisher="Taxon" pageId="3" pageNumber="146" pagination="1117 - 1140" part="64" refId="ref4810" refString="Zhou, X-M. & L. - B. Zhang (2015). A classification of Selaginella (Selaginellaceae) based on molecular (chloroplast and nuclear), macromorphological, and spore features. Taxon 64: 1117 - 1140." title="A classification of Selaginella (Selaginellaceae) based on molecular (chloroplast and nuclear), macromorphological, and spore features" type="journal article" year="2015">Zhou & Zhang, 2015</bibRefCitation>
|
|||
|
|
) ou sept sous-genres (
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FE681345FD11FBE6" author="Weststrand, S. & P. Korall" box="[414,755,1050,1074]" journalOrPublisher="Next Generation Pteridology Conference" pageId="3" pageNumber="146" refId="ref4609" refString="Weststrand, S. & P. Korall (2015). There is some morphology there after all! Interpreting morphology in Selaginella in the context of phylogeny. Poster. Next Generation Pteridology Conference, 1 - 5 June 2015, Washington D. C." title="There is some morphology there after all! Interpreting morphology in Selaginella in the context of phylogeny" type="proceedings paper" year="2015">Weststrand & Korall, 2015</bibRefCitation>
|
|||
|
|
). La nouvelle espèce,
|
|||
|
|
<emphasis id="B977F4529503CB29FEA21365FE07FB86" box="[340,485,1082,1106]" italics="true" pageId="3" pageNumber="146">
|
|||
|
|
<taxonomicName id="4C0353C39503CB29FEA21365FE03FB86" box="[340,481,1082,1106]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="146" phylum="Tracheophyta" rank="species" species="rasoloheryi">S. rasoloheryi</taxonomicName>
|
|||
|
|
,
|
|||
|
|
</emphasis>
|
|||
|
|
appartient au sous-genre
|
|||
|
|
<taxonomicName id="4C0353C39503CB29FF801305FEABFBA6" authority="Baker" authorityName="Baker" box="[118,329,1114,1138]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="subGenus" subGenus="Heterostachys">
|
|||
|
|
<emphasis id="B977F4529503CB29FF801305FEE3FBA6" box="[118,257,1114,1138]" italics="true" pageId="3" pageNumber="146">Heterostachys</emphasis>
|
|||
|
|
Baker
|
|||
|
|
</taxonomicName>
|
|||
|
|
selon la classification de
|
|||
|
|
<bibRefCitation id="EF9255B19503CB29FD921305FD19FBA6" author="Jermy, A. C." box="[612,763,1114,1138]" journalOrPublisher="Fern Gaz." pageId="3" pageNumber="146" pagination="2" part="13" refId="ref4226" refString="Jermy, A. C. (1986). Subgeneric names in Selaginella. Fern Gaz. 13 (2)." title="Subgeneric names in Selaginella" type="journal article" year="1986">Jermy (1986)</bibRefCitation>
|
|||
|
|
. Malgré le fait que le sous-genre
|
|||
|
|
<taxonomicName id="4C0353C39503CB29FE201325FDBDFB46" box="[470,607,1146,1170]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="subGenus" subGenus="Heterostachys">
|
|||
|
|
<emphasis id="B977F4529503CB29FE201325FDBDFB46" box="[470,607,1146,1170]" italics="true" pageId="3" pageNumber="146">Heterostachys</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
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|
ainsi défini ne soit pas monophylétique, nous suivons ici la classification de
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<bibRefCitation id="EF9255B19503CB29FF8313E5FEE6FB06" author="Jermy, A. C." box="[117,260,1210,1234]" journalOrPublisher="Fern Gaz." pageId="3" pageNumber="146" pagination="2" part="13" refId="ref4226" refString="Jermy, A. C. (1986). Subgeneric names in Selaginella. Fern Gaz. 13 (2)." title="Subgeneric names in Selaginella" type="journal article" year="1986">Jermy (1986)</bibRefCitation>
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basée uniquement sur la morphologie des taxons car, dans la phylogénie moléculaire (
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<bibRefCitation id="EF9255B19503CB29FE071385FD04FB26" author="Zhou, X-M. & L. - B. Zhang" box="[497,742,1242,1266]" journalOrPublisher="Taxon" pageId="3" pageNumber="146" pagination="1117 - 1140" part="64" refId="ref4810" refString="Zhou, X-M. & L. - B. Zhang (2015). A classification of Selaginella (Selaginellaceae) based on molecular (chloroplast and nuclear), macromorphological, and spore features. Taxon 64: 1117 - 1140." title="A classification of Selaginella (Selaginellaceae) based on molecular (chloroplast and nuclear), macromorphological, and spore features" type="journal article" year="2015">Zhou & Zhang, 2015</bibRefCitation>
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) à l’origine de la classification de
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<bibRefCitation id="EF9255B19503CB29FE4A13A5FD65FAC6" author="Zhou, X. - M. & C. J. Rothfels & L. Zhang & Z. - R. He & T. Le Pechon & H. He & N. T. Lu & R. Knapp & D. Lorence & X. - J. He & X. - F. Gao & L. - B. Zhang" box="[444,647,1274,1298]" journalOrPublisher="Cladistics" pageId="3" pageNumber="146" refId="ref4700" refString="Zhou, X. - M., C. J. Rothfels, L. Zhang, Z. - R. He, T. Le Pechon, H. He, N. T. Lu, R. Knapp, D. Lorence, X. - J. He, X. - F. Gao & L. - B. Zhang (2015). A large-scale phylogeny of the lycophyte genus Selaginella (Selaginellaceae: Lycopodiopsida) based on plastid and nuclear loci. Cladistics [http: // dx. doi. org / 10.1111 / cla. 12136]." title="A large-scale phylogeny of the lycophyte genus Selaginella (Selaginellaceae: Lycopodiopsida) based on plastid and nuclear loci" type="book" year="2015">Zhou et al. (2015)</bibRefCitation>
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, le nombre d’espèces de
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<collectingCountry id="F31468D09503CB29FEF51245FE6AFAE6" box="[259,392,1306,1330]" name="Madagascar" pageId="3" pageNumber="146">Madagascar</collectingCountry>
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pris en compte est trop restreint. Les caractéristiques morphologiques qui justifient le classement de
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<taxonomicName id="4C0353C39503CB29FEE31205FE7CFAA6" box="[277,414,1370,1394]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="146" phylum="Tracheophyta" rank="species" species="rasoloheryi">
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<emphasis id="B977F4529503CB29FEE31205FE7CFAA6" box="[277,414,1370,1394]" italics="true" pageId="3" pageNumber="146">S. rasoloheryi</emphasis>
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</taxonomicName>
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dans le sous-genre
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<taxonomicName id="4C0353C39503CB29FD831205FCE2FAA6" box="[629,768,1370,1394]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="subGenus" subGenus="Heterostachys">
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<emphasis id="B977F4529503CB29FD831205FCE2FAA6" box="[629,768,1370,1394]" italics="true" pageId="3" pageNumber="146">Heterostachys</emphasis>
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</taxonomicName>
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sont les suivantes: tige non articulée et absence de rhizome; enracinement de la tige principale par des rhizophores naissant sur la face ventrale, à l’aisselle des feuilles axillaires; feuilles stériles et fertiles dimorphes disposées sur 4 rangs; strobiles bilatéraux, aplatis dorsi-ventralement. Le sousgenre
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<taxonomicName id="4C0353C39503CB29FF4A1145FEAFF9E6" box="[188,333,1562,1586]" class="Lycopodiopsida" family="Selaginellaceae" genus="Selaginella" kingdom="Plantae" order="Selaginellales" pageId="3" pageNumber="144" phylum="Tracheophyta" rank="subGenus" subGenus="Heterostachys">
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<emphasis id="B977F4529503CB29FF4A1145FEAFF9E6" box="[188,333,1562,1586]" italics="true" pageId="3" pageNumber="146">Heterostachys</emphasis>
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</taxonomicName>
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est représenté à
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<collectingCountry id="F31468D09503CB29FDE41145FD79F9E6" box="[530,667,1562,1586]" name="Madagascar" pageId="3" pageNumber="146">Madagascar</collectingCountry>
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par huit espèces qui peuvent se distinguer de la façon suivante (c泩 de détermination basée en grande partie sur celle de Stefanovᔉ et al. (1997):
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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