treatments-xml/data/03/EB/70/03EB700FFFB3FFACA796FEC2FEAAF36F.xml

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<mods:title id="79E70D262D9918A0FFCC286C5B052574">Taxonomic notes on the genus Satyrium Scudder, 1876 (Lepidoptera, Lycaenidae) of Afghanistan with description of two new taxa</mods:title>
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<paragraph id="8BFDC119FFB3FFACA796FEC2FD82F75A" blockId="2.[151,565,295,322]" box="[151,565,295,322]" pageId="2" pageNumber="423">
<heading id="D0B57675FFB3FFACA796FEC2FD82F75A" bold="true" box="[151,565,295,322]" fontSize="11" level="1" pageId="2" pageNumber="423" reason="1">
<emphasis id="B9361D0BFFB3FFACA796FEC2FD82F75A" bold="true" box="[151,565,295,322]" pageId="2" pageNumber="423">
Subgenus 
<taxonomicName id="4C42BA9AFFB3FFACA616FECDFD82F75A" ID-CoL="9CM5M" authority="Strand, 1910" authorityName="Strand" authorityYear="1910" box="[279,565,295,322]" class="Insecta" family="Lycaenidae" genus="Superflua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B9361D0BFFB3FFACA616FECDFE38F759" bold="true" box="[279,399,295,321]" italics="true" pageId="2" pageNumber="423">Superflua</emphasis>
Strand, 1910
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFDC119FFB3FFACA796FE84FDB6F447" blockId="2.[151,1436,366,1399]" pageId="2" pageNumber="423">
All collected 
<taxonomicName id="4C42BA9AFFB3FFACA63BFE84FE10F79E" box="[314,423,366,390]" class="Insecta" family="Lycaenidae" genus="Superflua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B9361D0BFFB3FFACA63BFE84FE10F79E" box="[314,423,366,390]" italics="true" pageId="2" pageNumber="423">Superflua</emphasis>
</taxonomicName>
specimens belong to the widely distributed 
<taxonomicName id="4C42BA9AFFB3FFACA4C0FE85FF62F7B3" authority="Kollar, 1849" authorityName="Kollar" authorityYear="1849" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="sassanides" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA4C0FE85FB91F79E" box="[961,1062,367,390]" italics="true" pageId="2" pageNumber="423">Satyrium</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA33DFE84FB1EF79E" box="[1084,1193,366,390]" italics="true" pageId="2" pageNumber="423">Superflua</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA3C1FE84FA8FF79E" box="[1216,1336,366,390]" italics="true" pageId="2" pageNumber="423">sassanides</emphasis>
(
<bibRefCitation id="EFD3BCE8FFB3FFACA24CFE84FF79F7B3" author="Kollar" pageId="2" pageNumber="423" refString="Kollar, V. ([1849]) Lepidoptera. In: Kollar, V. &amp; Redtenbacher, L. (Eds.), Ueber den Charakter der Insecten-Fauna von Sudpersien. Denkschrifteb der Kaiserlichen Akademie der Wissenschaften, Bd. 1, S. 42 - 53. [Wien]" type="book chapter" year="1849">Kollar, 1849</bibRefCitation>
)
</taxonomicName>
—species group which includes tailed butterflies with well-developed white postdiscal line on hindwing underside: complex of species from Iranian Highland— 
<taxonomicName id="4C42BA9AFFB3FFACA415FE5DFC6CF7D6" box="[788,987,438,463]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="sassanides" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA415FE5DFC9EF7D6" box="[788,809,439,462]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA43DFE5DFCE6F7D6" box="[828,849,439,462]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA462FE5CFC6CF7D6" box="[867,987,438,462]" italics="true" pageId="2" pageNumber="423">sassanides</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C42BA9AFFB3FFACA314FE5DFAE5F7D7" authority="Eckweiler" authorityName="Eckweiler" box="[1045,1362,438,463]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="persepolis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA314FE5DFB9DF7D6" box="[1045,1066,439,462]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA33CFE5DFBE5F7D6" box="[1085,1106,439,462]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA365FE5CFB60F7D6" box="[1124,1239,438,462]" italics="true" pageId="2" pageNumber="423">persepolis</emphasis>
Eckweiler
</taxonomicName>
&amp; ten Hagen, 2003, recently described 
<taxonomicName id="4C42BA9AFFB3FFACA51FFE31FB9DF7EB" authority="Churkin &amp; Pletnev, 2010" authorityName="Churkin &amp; Pletnev" authorityYear="2010" box="[542,1066,474,499]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="turkmanica" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA51FFE31FD84F7EA" box="[542,563,475,498]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA548FE31FDE8F7EA" box="[585,607,475,498]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA575FE31FD44F7EB" box="[628,755,475,499]" italics="true" pageId="2" pageNumber="423">turkmanica</emphasis>
<bibRefCitation id="EFD3BCE8FFB3FFACA400FE30FB9DF7EB" author="Churkin" box="[769,1066,474,499]" pageId="2" pageNumber="423" refString="Churkin, S. V. &amp; Pletnev, V. A. (2010) A review of the Satyrium (Superflua) sassanides (Kollar, 1849) species group from &quot; Russian &quot; Central Asia (Lepidoptera, Lycaenidae). Atalanta, 41 (1 / 2), 207 - 220." type="journal article" year="2010">Churkin &amp; Pletnev, 2010</bibRefCitation>
</taxonomicName>
from Turkmenian Kopet Dagh mountains, Middle and Central Asian 
<taxonomicName id="4C42BA9AFFB3FFACA54BFE15FC75F40F" authority="Erschoff, 1874" authorityName="Erschoff" authorityYear="1874" box="[586,962,510,535]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA54BFE15FDE8F40E" box="[586,607,511,534]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA570FE15FD32F40E" box="[625,645,511,534]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA599FE14FD4AF40E" box="[664,765,510,534]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
(
<bibRefCitation id="EFD3BCE8FFB3FFACA40CFE14FC0DF40F" author="Erschoff" box="[781,954,510,535]" pageId="2" pageNumber="423" refString="Erschoff, N. G. (1874) Lepidoptera. In: Fedtschenko, A. P. (Ed.), Voyage in Turkestan (1874 - 1879). Memoires de la Societe des amis de sciences naturelles Moscou, 11 (2), pp. [1 - 4] + I - VI + 1 - 127. [in Russian]" type="journal article" year="1874">Erschoff, 1874</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="4C42BA9AFFB3FFACA4CEFE15FA2BF40F" authority="Churkin &amp; Pletnev, 2010" authorityName="Churkin &amp; Pletnev" authorityYear="2010" box="[975,1436,510,535]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="zabirovi" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA4CEFE15FC53F40E" box="[975,996,511,534]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA4F4FE15FBBDF40E" box="[1013,1034,511,534]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA31DFE14FBC0F40E" box="[1052,1143,510,534]" italics="true" pageId="2" pageNumber="423">zabirovi</emphasis>
<bibRefCitation id="EFD3BCE8FFB3FFACA381FE14FA2BF40F" author="Churkin" box="[1152,1436,510,535]" pageId="2" pageNumber="423" refString="Churkin, S. V. &amp; Pletnev, V. A. (2010) A review of the Satyrium (Superflua) sassanides (Kollar, 1849) species group from &quot; Russian &quot; Central Asia (Lepidoptera, Lycaenidae). Atalanta, 41 (1 / 2), 207 - 220." type="journal article" year="2010">Churkin &amp; Pletnev, 2010</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C42BA9AFFB3FFACA7C8FDC9FD1AF423" authority="Churkin &amp; Pletnev, 2010" authorityName="Churkin &amp; Pletnev" authorityYear="2010" box="[201,685,546,571]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="muksuria" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA7C8FDC9FF69F422" box="[201,222,547,570]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA7F0FDC9FEB2F422" box="[241,261,547,570]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA618FDC9FE36F423" box="[281,385,547,571]" italics="true" pageId="2" pageNumber="423">muksuria</emphasis>
<bibRefCitation id="EFD3BCE8FFB3FFACA68AFDC8FD1AF423" author="Churkin" box="[395,685,546,571]" pageId="2" pageNumber="423" refString="Churkin, S. V. &amp; Pletnev, V. A. (2010) A review of the Satyrium (Superflua) sassanides (Kollar, 1849) species group from &quot; Russian &quot; Central Asia (Lepidoptera, Lycaenidae). Atalanta, 41 (1 / 2), 207 - 220." type="journal article" year="2010">Churkin &amp; Pletnev, 2010</bibRefCitation>
</taxonomicName>
, and Himalayan 
<taxonomicName id="4C42BA9AFFB3FFACA470FDC9FB1BF423" authority="Moore, 1865" authorityName="Moore" authorityYear="1865" box="[881,1196,546,571]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="deria" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA470FDC9FC30F422" box="[881,903,547,570]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA498FDC9FC18F422" box="[921,943,547,570]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA4C0FDC9FC4BF423" box="[961,1020,547,571]" italics="true" pageId="2" pageNumber="423">deria</emphasis>
(
<bibRefCitation id="EFD3BCE8FFB3FFACA30FFDC9FB13F423" author="Moore" box="[1038,1188,546,571]" pageId="2" pageNumber="423" refString="Moore, F. (1865) List of Diurnal Lepidoptera collected by Capt. A. M. Lang in the N. W. Himalayas. Proceedings, Zoological Society of London, 2, 485 - 511. http: // dx. doi. org / 10.1111 / j. 1469 - 7998.1865. tb 02378. x" type="journal article" year="1865">Moore, 1865</bibRefCitation>
)
</taxonomicName>
(
<bibRefCitation id="EFD3BCE8FFB3FFACA3BCFDC8FF64F447" author="Weidenhoffer" pageId="2" pageNumber="423" refString="Weidenhoffer, Z., Bozano, G. C. &amp; Churkin, S. (2004) Guide to the butterflies of the Palearctic Region. Lycaenidae part II. Omnes Artes, Milano, 94 pp." type="book" year="2004">
Weidenhoffer 
<emphasis id="B9361D0BFFB3FFACA260FDC9FA22F423" box="[1377,1429,547,571]" italics="true" pageId="2" pageNumber="423">et al</emphasis>
. 2004
</bibRefCitation>
; 
<bibRefCitation id="EFD3BCE8FFB3FFACA7DCFDACFE42F447" author="Churkin" box="[221,501,582,607]" pageId="2" pageNumber="423" refString="Churkin, S. V. &amp; Pletnev, V. A. (2010) A review of the Satyrium (Superflua) sassanides (Kollar, 1849) species group from &quot; Russian &quot; Central Asia (Lepidoptera, Lycaenidae). Atalanta, 41 (1 / 2), 207 - 220." type="journal article" year="2010">Churkin &amp; Pletnev 2010</bibRefCitation>
).
</paragraph>
<paragraph id="8BFDC119FFB3FFACA7C6FD80FEFBF52F" blockId="2.[151,1436,366,1399]" pageId="2" pageNumber="423">
Specimens from the two 
<taxonomicName id="4C42BA9AFFB3FFACA6E0FD81FDF9F49B" box="[481,590,619,643]" class="Insecta" family="Lycaenidae" genus="Superflua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B9361D0BFFB3FFACA6E0FD81FDF9F49B" box="[481,590,619,643]" italics="true" pageId="2" pageNumber="423">Superflua</emphasis>
</taxonomicName>
populations collected in the foothills near Kabul (Qargha and Paghman) at mid altitude (
<quantity id="4CBA6CFCFFB3FFACA62DFD65FE34F4BF" box="[300,387,655,679]" metricMagnitude="3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="423" unit="m" value="1900.0">1900 m</quantity>
) at the end of May–June 
<date id="FFFCE7D9FFB3FFACA59FFD65FCABF4BF" box="[670,796,655,679]" pageId="2" pageNumber="423" value="2010" valueMax="2012">2010–2012</date>
have traits of rather typical 
<taxonomicName id="4C42BA9AFFB3FFACA357FD65FAB2F4BE" box="[1110,1285,654,679]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA357FD65FBD3F4BE" box="[1110,1124,655,678]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA37AFD65FB3EF4BE" box="[1147,1161,655,678]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA3A1FD64FAB2F4BE" box="[1184,1285,654,678]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
</taxonomicName>
. This species is widely distributed in Middle and Central Asia and was formerly treated as a subspecies of 
<taxonomicName id="4C42BA9AFFB3FFACA3A9FD59FADDF4D3" box="[1192,1386,691,715]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="sassanides" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA3A9FD59FB0AF4D2" box="[1192,1213,691,714]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA3CCFD59FB55F4D2" box="[1229,1250,691,714]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA3F3FD59FADDF4D3" box="[1266,1386,691,715]" italics="true" pageId="2" pageNumber="423">sassanides</emphasis>
</taxonomicName>
, but studies conducted by 
<bibRefCitation id="EFD3BCE8FFB3FFACA695FD3CFD7EF4F7" box="[404,713,726,751]" pageId="2" pageNumber="423" refString="Weidenhoffer, Z., Bozano, G. C. &amp; Churkin, S. (2004) Guide to the butterflies of the Palearctic Region. Lycaenidae part II. Omnes Artes, Milano, 94 pp." type="book">
Weidenhoffer 
<emphasis id="B9361D0BFFB3FFACA539FD3DFDC3F4F6" box="[568,628,726,750]" italics="true" pageId="2" pageNumber="423">et al.</emphasis>
(2004)
</bibRefCitation>
proved its specific status. Studies by Eckweiler &amp; ten Hagen (2003) and 
<bibRefCitation id="EFD3BCE8FFB3FFACA61BFD10FDF2F50B" box="[282,581,762,787]" pageId="2" pageNumber="423" refString="Weidenhoffer, Z., Bozano, G. C. &amp; Churkin, S. (2004) Guide to the butterflies of the Palearctic Region. Lycaenidae part II. Omnes Artes, Milano, 94 pp." type="book">
Weidenhoffer 
<emphasis id="B9361D0BFFB3FFACA6BBFD11FE44F50B" box="[442,499,763,787]" italics="true" pageId="2" pageNumber="423">et al.</emphasis>
(2004)
</bibRefCitation>
outlined distribution of 
<taxonomicName id="4C42BA9AFFB3FFACA458FD11FBA9F50B" box="[857,1054,763,787]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="sassanides" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA458FD11FCD0F50A" box="[857,871,763,786]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA47EFD11FC3AF50A" box="[895,909,763,786]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA4A7FD11FBA9F50B" box="[934,1054,763,787]" italics="true" pageId="2" pageNumber="423">sassanides</emphasis>
</taxonomicName>
by mountains of South-West 
<collectingCountry id="F3558189FFB3FFACA26EFD11FA2BF50A" box="[1391,1436,763,786]" name="Iran" pageId="2" pageNumber="423">Iran</collectingCountry>
(
<collectingCountry id="F3558189FFB3FFACA79FFCF5FF67F52F" box="[158,208,799,823]" name="Iran" pageId="2" pageNumber="423">Fars</collectingCountry>
Province).
</paragraph>
</subSubSection>
<subSubSection id="C3589292FFB3FFACA7C6FCA9FEAAF36F" pageId="2" pageNumber="423" type="reference_group">
<paragraph id="8BFDC119FFB3FFACA7C6FCA9FAA4F2FE" blockId="2.[151,1436,366,1399]" pageId="2" pageNumber="423">
<taxonomicName id="4C42BA9AFFB3FFACA7C6FCA9FEC0F543" box="[199,375,835,859]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA7C6FCA9FF6BF542" box="[199,220,835,858]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA7EDFCA9FEB6F542" box="[236,257,835,858]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA613FCA9FEC0F543" box="[274,375,835,859]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
</taxonomicName>
was already reported for 
<collectingCountry id="F3558189FFB3FFACA599FCA8FC96F543" box="[664,801,834,859]" name="Afghanistan" pageId="2" pageNumber="423">Afghanistan</collectingCountry>
by 
<bibRefCitation id="EFD3BCE8FFB3FFACA44DFCA8FC6BF543" author="Sakai" box="[844,988,834,859]" pageId="2" pageNumber="423" refString="Sakai, S. (1981) Butterflies of Afghanistan. Koudansha, Tokyo, 272 pp. [in Japanese]" type="book" year="1981">Sakai (1981)</bibRefCitation>
(under the name 
<taxonomicName id="4C42BA9AFFB3FFACA3A0FCA9FAAEF543" box="[1185,1305,835,859]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="sassanides">
<emphasis id="B9361D0BFFB3FFACA3A0FCA9FAAEF543" box="[1185,1305,835,859]" italics="true" pageId="2" pageNumber="423">sassanides</emphasis>
</taxonomicName>
) but in fact only two specimens pictured in the book belong to 
<taxonomicName id="4C42BA9AFFB3FFACA5EEFC8DFC12F566" box="[751,933,870,895]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA5EEFC8DFD4AF566" box="[751,765,871,894]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA416FC8DFC92F566" box="[791,805,871,894]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA441FC8CFC12F566" box="[832,933,870,894]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
</taxonomicName>
(col. plate 46, figs. 3, 6), while two others (specimens from Bala Quran, NE Hindu Kush Mts., Badakhshan Prov.) clearly belong to 
<taxonomicName id="4C42BA9AFFB3FFACA3A1FC61FA9CF5BB" box="[1184,1323,907,931]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="deria" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA3A1FC61FB19F5BA" box="[1184,1198,907,930]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA3C9FC61FB61F5BA" box="[1224,1238,907,930]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA3F1FC61FA9CF5BB" box="[1264,1323,907,931]" italics="true" pageId="2" pageNumber="423">deria</emphasis>
</taxonomicName>
, which is characterized by the absence of androconial patch on the male forewing and in the colour pattern on the hindwing underside. 
<bibRefCitation id="EFD3BCE8FFB3FFACA612FC38FDF7F5F3" box="[275,576,978,1003]" pageId="2" pageNumber="423" refString="Weidenhoffer, Z., Bozano, G. C. &amp; Churkin, S. (2004) Guide to the butterflies of the Palearctic Region. Lycaenidae part II. Omnes Artes, Milano, 94 pp." type="book">
Weidenhoffer 
<emphasis id="B9361D0BFFB3FFACA6B5FC39FE5AF5F3" box="[436,493,979,1003]" italics="true" pageId="2" pageNumber="423">et al.</emphasis>
(2004)
</bibRefCitation>
also mentioned the latter species for Bala Quran. One specimen pictured in the book on butterflies of 
<collectingCountry id="F3558189FFB3FFACA6B6FC1CFDF7F217" box="[439,576,1014,1039]" name="Afghanistan" pageId="2" pageNumber="423">Afghanistan</collectingCountry>
(
<bibRefCitation id="EFD3BCE8FFB3FFACA54FFC1CFD79F217" author="Sakai" box="[590,718,1014,1039]" pageId="2" pageNumber="423" refString="Sakai, S. (1981) Butterflies of Afghanistan. Koudansha, Tokyo, 272 pp. [in Japanese]" type="book" year="1981">Sakai 1981</bibRefCitation>
) (col. plate 46, fig. 3) was mentioned by 
<bibRefCitation id="EFD3BCE8FFB3FFACA398FC1CFADCF217" author="Charmeaux" box="[1177,1387,1014,1039]" pageId="2" pageNumber="423" refString="Charmeaux, J. F. (2004) Une nouvelle espece du sous-genre Superflua du Pakistan. Phegea, 32, 9 - 16." type="journal article" year="2004">Charmeaux (2004)</bibRefCitation>
as a “ 
<typeStatus id="54F97FBBFFB3FFACA7A0FBF1FEB1F22B" box="[161,262,1051,1075]" pageId="2" pageNumber="423" type="paratype">paratype</typeStatus>
” of 
<taxonomicName id="4C42BA9AFFB3FFACA635FBF1FD1EF22B" authority="Charmeaux, 2004" authorityName="Charmeaux" authorityYear="2004" box="[308,681,1050,1075]" class="Insecta" family="Lycaenidae" genus="Superflua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="khowari" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA635FBF1FEF5F22A" box="[308,322,1051,1074]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA658FBF1FED0F22A" box="[345,359,1051,1074]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA67CFBF1FE6EF22B" box="[381,473,1051,1075]" italics="true" pageId="2" pageNumber="423">khowari</emphasis>
<bibRefCitation id="EFD3BCE8FFB3FFACA6E1FBF0FD1EF22B" author="Charmeaux" box="[480,681,1050,1075]" pageId="2" pageNumber="423" refString="Charmeaux, J. F. (2004) Une nouvelle espece du sous-genre Superflua du Pakistan. Phegea, 32, 9 - 16." type="journal article" year="2004">Charmeaux, 2004</bibRefCitation>
</taxonomicName>
, which he described from Chitral (North 
<collectingCountry id="F3558189FFB3FFACA372FBF0FB60F22B" box="[1139,1239,1050,1075]" name="Pakistan" pageId="2" pageNumber="423">Pakistan</collectingCountry>
). 
<bibRefCitation id="EFD3BCE8FFB3FFACA3E8FBF0FEBFF24F" pageId="2" pageNumber="423" refString="Weidenhoffer, Z., Bozano, G. C. &amp; Churkin, S. (2004) Guide to the butterflies of the Palearctic Region. Lycaenidae part II. Omnes Artes, Milano, 94 pp." type="book">
Weidenhoffer 
<emphasis id="B9361D0BFFB3FFACA289FBF1FF03F24E" italics="true" pageId="2" pageNumber="423">et al.</emphasis>
(2004)
</bibRefCitation>
synonymized the latter species with 
<taxonomicName id="4C42BA9AFFB3FFACA5B7FBD5FCDCF24E" box="[694,875,1086,1111]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA5B7FBD5FD73F24E" box="[694,708,1087,1110]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA5DFFBD5FD5BF24E" box="[734,748,1087,1110]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA407FBD4FCDCF24E" box="[774,875,1086,1110]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
</taxonomicName>
on a base of external and genital similarity of a 
<typeStatus id="54F97FBBFFB3FFACA796FB89FF40F263" box="[151,247,1123,1147]" pageId="2" pageNumber="423" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C42BA9AFFB3FFACA61EFB89FE7FF263" box="[287,456,1123,1147]" class="Insecta" family="Lycaenidae" genus="Superflua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="khowari" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA61EFB89FE9AF262" box="[287,301,1123,1146]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA644FB89FEE4F262" box="[325,339,1123,1146]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA66DFB89FE7FF263" box="[364,456,1123,1147]" italics="true" pageId="2" pageNumber="423">khowari</emphasis>
</taxonomicName>
with specimens of 
<taxonomicName id="4C42BA9AFFB3FFACA5A4FB89FCE0F263" box="[677,855,1123,1147]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA5A4FB89FD04F262" box="[677,691,1123,1146]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA5CDFB89FD6DF262" box="[716,730,1123,1146]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA5F3FB89FCE0F263" box="[754,855,1123,1147]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
</taxonomicName>
, but 
<bibRefCitation id="EFD3BCE8FFB3FFACA493FB88FB0CF263" author="Churkin" box="[914,1211,1122,1147]" pageId="2" pageNumber="423" refString="Churkin, S. V. &amp; Pletnev, V. A. (2010) A review of the Satyrium (Superflua) sassanides (Kollar, 1849) species group from &quot; Russian &quot; Central Asia (Lepidoptera, Lycaenidae). Atalanta, 41 (1 / 2), 207 - 220." type="journal article" year="2010">Churkin &amp; Pletnev (2010)</bibRefCitation>
suppose that 
<taxonomicName id="4C42BA9AFFB3FFACA256FB89FF44F286" class="Insecta" family="Lycaenidae" genus="Superflua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="khowari" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA256FB89FAD2F262" box="[1367,1381,1123,1146]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA27CFB89FA3CF262" box="[1405,1419,1123,1146]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA796FB6CFF44F286" box="[151,243,1158,1182]" italics="true" pageId="2" pageNumber="423">khowari</emphasis>
</taxonomicName>
is not synonymous with 
<taxonomicName id="4C42BA9AFFB3FFACA511FB6DFD76F286" box="[528,705,1158,1183]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA511FB6DFDA9F286" box="[528,542,1159,1182]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA537FB6DFDF3F286" box="[566,580,1159,1182]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA55DFB6CFD76F286" box="[604,705,1158,1182]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
</taxonomicName>
. Analysis of the genitalia of the specimens collected near Kabul showed their similarity with typical 
<taxonomicName id="4C42BA9AFFB3FFACA52DFB41FD6CF2DB" box="[556,731,1195,1219]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA52DFB41FD8DF2DA" box="[556,570,1195,1218]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA550FB41FDE8F2DA" box="[593,607,1195,1218]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA577FB41FD6CF2DB" box="[630,731,1195,1219]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
</taxonomicName>
from Middle Asia. Thus there are two described species of the subgenus 
<taxonomicName id="4C42BA9AFFB3FFACA607FB24FEC4F2FE" ID-CoL="9CM5M" box="[262,371,1230,1254]" class="Insecta" family="Lycaenidae" genus="Superflua" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B9361D0BFFB3FFACA607FB24FEC4F2FE" box="[262,371,1230,1254]" italics="true" pageId="2" pageNumber="423">Superflua</emphasis>
</taxonomicName>
known from the territory of 
<collectingCountry id="F3558189FFB3FFACA5B4FB24FC89F2FF" box="[693,830,1230,1255]" name="Afghanistan" pageId="2" pageNumber="423">Afghanistan</collectingCountry>
so far— 
<taxonomicName id="4C42BA9AFFB3FFACA49EFB25FBF8F2FE" box="[927,1103,1230,1255]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="mirabilis" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA49EFB25FC1AF2FE" box="[927,941,1231,1254]" italics="true" pageId="2" pageNumber="423">S</emphasis>
. (
<emphasis id="B9361D0BFFB3FFACA4C4FB25FC64F2FE" box="[965,979,1231,1254]" italics="true" pageId="2" pageNumber="423">S</emphasis>
.) 
<emphasis id="B9361D0BFFB3FFACA4EBFB24FBF8F2FE" box="[1002,1103,1230,1254]" italics="true" pageId="2" pageNumber="423">mirabilis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C42BA9AFFB3FFACA387FB25FABCF2FE" box="[1158,1291,1230,1255]" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="species" species="deria" subGenus="Superflua">
<emphasis id="B9361D0BFFB3FFACA387FB25FB2CF2FE" box="[1158,1179,1231,1254]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA3AAFB25FB77F2FE" box="[1195,1216,1231,1254]" italics="true" pageId="2" pageNumber="423">S.</emphasis>
) 
<emphasis id="B9361D0BFFB3FFACA3D1FB24FABCF2FE" box="[1232,1291,1230,1254]" italics="true" pageId="2" pageNumber="423">deria</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFDC119FFB3FFACA7C6FB18FEAAF36F" blockId="2.[151,1436,366,1399]" pageId="2" pageNumber="423">
Specimens collected in Band-e Amir (Bamyan Province) at higher altitudes (about 
<quantity id="4CBA6CFCFFB3FFACA384FB19FB69F313" box="[1157,1246,1267,1291]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="423" unit="m" value="3000.0">3000 m</quantity>
) in July–August 
<date id="FFFCE7D9FFB3FFACA796FAFDFEAFF337" box="[151,280,1303,1327]" pageId="2" pageNumber="423" value="2009" valueMax="2013">2009–2013</date>
, along with a single female from Koh-i-Baba Mts., have unique combination of external characters and genitalia, which allow to describe this population as a new species belonging to Iranian complex of 
<taxonomicName id="4C42BA9AFFB3FFACA236FAD1FEAFF36F" ID-CoL="9CLZF" authority="(Superflua)" baseAuthorityName="Superflua" class="Insecta" family="Lycaenidae" genus="Satyrium" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="423" phylum="Arthropoda" rank="genus">
<emphasis id="B9361D0BFFB3FFACA236FAD1FA2BF34A" box="[1335,1436,1339,1362]" italics="true" pageId="2" pageNumber="423">Satyrium</emphasis>
(
<emphasis id="B9361D0BFFB3FFACA7A1FAB4FEBAF36E" box="[160,269,1374,1398]" italics="true" pageId="2" pageNumber="423">Superflua</emphasis>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>