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<document id="6D8F411456B4465857C9C2C502120559" ID-DOI="10.15407/zoo2022.02.091" ID-ISSN="2707-7268" ID-Zenodo-Dep="7175709" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665294511910" checkinUser="felipe" docAuthor="Glotov, S., Hushtan, K., Hushtan, H., Koval, N. &amp; Diedus, V." docDate="2022" docId="03E487DDFFA7FF93FF37086FFBCD8227" docLanguage="en" docName="Zoodiversity.56.1.91–110.pdf" docOrigin="Zoodiversity 56 (1)" docSource="http://dx.doi.org/10.15407/zoo2022.02.091" docStyle="DocumentStyle:65E6F982F72656C606C1639D3996B172.2:Zoodiversity.2020-.journal_article" docStyleId="65E6F982F72656C606C1639D3996B172" docStyleName="Zoodiversity.2020-.journal_article" docStyleVersion="2" docTitle="Atheta subtilis" docType="treatment" docVersion="3" lastPageNumber="105" masterDocId="FFDDFFA5FFAAFF9DFF990E30FFE2805E" masterDocTitle="The Genus Atheta (Coleoptera, Staphylinidae, Aleocharinae) In The Ukrainian Carpathians" masterLastPageNumber="110" masterPageNumber="91" pageNumber="104" updateTime="1699381295323" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-4.0">
<mods:mods id="C5C06A42E00A0852A6A59C317DE2ADD1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="542256D2A25088F850E4722ACBA07ABE">The Genus Atheta (Coleoptera, Staphylinidae, Aleocharinae) In The Ukrainian Carpathians</mods:title>
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<mods:namePart id="F922E878C0DAD401A1B7D45928DDCDDA">Glotov, S.</mods:namePart>
<mods:affiliation id="F1132CA78CF8FAACF47396C10E879029">State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine</mods:affiliation>
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<mods:namePart id="41C0C36DA097ABD14A29F84788522B3B">Hushtan, K.</mods:namePart>
<mods:affiliation id="CBE9095AA7BF8FE71E24D21848E35C12">State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine &amp; Ecological College of Lviv National Agrarian University, Zamarstynivska st., 167, Lviv, 79068 Ukraine</mods:affiliation>
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<mods:namePart id="C7D1B123768BB57E1194774D278E736A">Hushtan, H.</mods:namePart>
<mods:affiliation id="1164CF5DE323F171DF76CE6E7F531BD9">State Museum of Natural History NAS of Ukraine, Teatralna st., 18, Lviv, 79008 Ukraine</mods:affiliation>
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<mods:namePart id="5D7086A473A672780AAE6B747AD66D40">Koval, N.</mods:namePart>
<mods:affiliation id="5ECBDEFBAF52DFCC105D3EC808C6CDD8">Uzhgorod National University, A. Voloshyn st., 32, Uzhgorod, 88000 Ukraine &amp; Uzhansky National Nature Park, Nezalezhnosti st., 7, village Velykyi Bereznyi, Ukraine</mods:affiliation>
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</mods:role>
<mods:namePart id="321A4F03AE25E5D2643832E0D31B4D21">Diedus, V.</mods:namePart>
<mods:affiliation id="BD20DEC2BD81F16038510D3C7DDB9D25">Uzhgorod National University, A. Voloshyn st., 32, Uzhgorod, 88000 Ukraine</mods:affiliation>
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<mods:title id="7C48B2FA9D594141B3E2DDB82A6DC0A0">Zoodiversity</mods:title>
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<mods:date id="4C4BF87D13736306CBDE374917FF77E3">2022</mods:date>
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<mods:number id="076202561A14CBE426A386C17B1653E9">56</mods:number>
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<mods:start id="D13C2A8A19063F8E5E49C937C790E263">91</mods:start>
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<mods:identifier id="EB481620E42F439447055671E63120E2" type="DOI">10.15407/zoo2022.02.091</mods:identifier>
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<mods:identifier id="BC4394C5805AA8507EBA13BF67E8CB0F" type="Zenodo-Dep">7175709</mods:identifier>
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<treatment id="03E487DDFFA7FF93FF37086FFBCD8227" LSID="urn:lsid:plazi:treatment:03E487DDFFA7FF93FF37086FFBCD8227" httpUri="http://treatment.plazi.org/id/03E487DDFFA7FF93FF37086FFBCD8227" lastPageId="14" lastPageNumber="105" pageId="13" pageNumber="104">
<subSubSection id="C3576540FFA7FF90FF37086FFD9D8627" box="[174,639,1631,1658]" pageId="13" pageNumber="104" type="nomenclature">
<paragraph id="8BF236CBFFA7FF90FF37086FFD9D8627" blockId="13.[174,639,1631,1658]" box="[174,639,1631,1658]" pageId="13" pageNumber="104">
<heading id="D0BA81A7FFA7FF90FF37086FFD9D8627" box="[174,639,1631,1658]" fontSize="36" level="2" pageId="13" pageNumber="104" reason="3">
<taxonomicName id="4C4D4D48FFA7FF90FF37086FFDE48627" ID-CoL="5W8RZ" authority="(Scriba, 1866)" baseAuthorityName="Scriba" baseAuthorityYear="1866" box="[174,518,1631,1658]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="104" phylum="Arthropoda" rank="species" species="subtilis">
<emphasis id="B939EAD9FFA7FF90FF37086FFEBE8624" bold="true" box="[174,348,1631,1658]" italics="true" pageId="13" pageNumber="104">Atheta subtilis</emphasis>
(Scriba, 1866)
</taxonomicName>
(
<figureCitation id="13762A4EFFA7FF90FD8D086FFDAD8627" box="[532,591,1631,1657]" captionStart="Fig" captionStartId="4.[170,203,1668,1690]" captionTargetBox="[229,1172,250,1618]" captionTargetId="figure-13@4.[166,1205,244,1630]" captionTargetPageId="4" captionText="Fig. 1. Habitus: a — Atheta coriaria (Kraatz, 1856); b — A. divisa (Märkel, 1844); c — A. graminicola (Gravenhorst, 1806); d — A. oblita (Erichson, 1839); e — A. ravilla (Erichson, 1839); f — A. macrocera (Thomson, 1856); g — A. occulta (Erichson, 1837); h — A. longicornis (Gravenhorst, 1802); i — A. melanaria (Mannerheim, 1830); j — A. zosterae (Thomson, 1856); k — A. atramentaria (Gyllenhal, 1810); l — A. marcida (Erichson, 1837); m — A. subtilis (Scriba, 1866); n — A. amicula (Stephens, 1832); o — A. laticollis Stephens, 1832; p — A. liturata (Stephens, 1832); q — A. nigritula (Gravenhorst, 1802)." figureDoi="http://doi.org/10.5281/zenodo.7175711" httpUri="https://zenodo.org/record/7175711/files/figure.png" pageId="13" pageNumber="104">fig. 1</figureCitation>
, m)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3576540FFA7FF93FF7A08A8FD5D81D4" lastPageId="14" lastPageNumber="105" pageId="13" pageNumber="104" type="materials_examined">
<paragraph id="8BF236CBFFA7FF90FF7A08A8FB7F8697" blockId="13.[174,1232,1688,1737]" pageId="13" pageNumber="104">
<materialsCitation id="3B253C96FFA7FF90FF7A08A8FE6486F0" box="[227,390,1688,1710]" location="Material." pageId="13" pageNumber="104" specimenCount="2">
<location id="8E926010FFA7FF90FF7A08A8FEAC86F0" LSID="urn:lsid:plazi:treatment:03E487DDFFA7FF93FF37086FFBCD8227:8E926010FFA7FF90FF7A08A8FEAC86F0" box="[227,334,1688,1710]" name="Material." pageId="13" pageNumber="104">Material.</location>
<specimenCount id="9D4BFD42FFA7FF90FEC108A8FE6486F0" box="[344,390,1688,1710]" pageId="13" pageNumber="104" type="generic">2 ex.</specimenCount>
</materialsCitation>
<materialsCitation id="3B253C96FFA7FF90FE1208A8FCC986F0" box="[395,811,1688,1710]" location="Lviv" pageId="13" pageNumber="104" specimenCount="1" stateProvince="L'viv">
<emphasis id="B939EAD9FFA7FF90FE1208A8FE5D86F3" bold="true" box="[395,447,1688,1709]" pageId="13" pageNumber="104">LWI:</emphasis>
<collectingRegion id="4989F829FFA7FF90FE5C08A8FE0D86F0" box="[453,495,1688,1710]" country="Ukraine" name="L'viv" pageId="13" pageNumber="104">Lviv</collectingRegion>
[
<collectingRegion id="4989F829FFA7FF90FE6208A8FDD486F0" box="[507,566,1688,1710]" country="Ukraine" name="L'viv" pageId="13" pageNumber="104">Lwiw</collectingRegion>
], date not specified,
<specimenCount id="9D4BFD42FFA7FF90FD6508A8FCC986F0" box="[764,811,1688,1710]" pageId="13" pageNumber="104" type="generic">1 ex.</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B253C96FFA7FF90FCAC08A8FE7A8697" collectingDate="1917-04-06" collectionCode="SMNH" location="Pohulanka" pageId="13" pageNumber="104" specimenCount="1" stateProvince="L'viv">
same locality but,
<location id="8E926010FFA7FF90FC7C08A8FBAE86F0" LSID="urn:lsid:plazi:treatment:03E487DDFFA7FF93FF37086FFBCD8227:8E926010FFA7FF90FC7C08A8FBAE86F0" box="[997,1100,1688,1710]" name="Pohulanka" pageId="13" pageNumber="104">Pohulanka</location>
[Pohulanka],
<date id="FFF3100BFFA7FF90FF370883FEE78697" box="[174,261,1715,1737]" pageId="13" pageNumber="104" value="1917-04-06">
<collectingDate id="EFB7E9E3FFA7FF90FF370883FEE78697" box="[174,261,1715,1737]" pageId="13" pageNumber="104" value="1917-04-06">6.4. 1917</collectingDate>
</date>
,
<specimenCount id="9D4BFD42FFA7FF90FE970883FED98697" box="[270,315,1715,1737]" pageId="13" pageNumber="104" type="generic">1 ex.</specimenCount>
(
<collectionCode id="ED5CAE0EFFA7FF90FEDF0883FE738697" box="[326,401,1715,1737]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="13" pageNumber="104">SMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B253C96FFA7FF90FE390883FB7B8697" box="[416,1177,1715,1737]" collectingDate="2009-05-21" collectionCode="SMNH" collectorName="Yu. Kanarskyi" location="mountain Yavirnyk" pageId="13" pageNumber="104" specimenCount="1" stateProvince="L'viv">
<emphasis id="B939EAD9FFA7FF90FE390883FE358696" bold="true" box="[416,471,1715,1736]" pageId="13" pageNumber="104">ZAK:</emphasis>
<location id="8E926010FFA7FF90FE450883FD798697" LSID="urn:lsid:plazi:treatment:03E487DDFFA7FF93FF37086FFBCD8227:8E926010FFA7FF90FE450883FD798697" box="[476,667,1715,1737]" name="mountain Yavirnyk" pageId="13" pageNumber="104" stateProvince="L'viv">mountain Yavirnyk</location>
, 03.05–
<date id="FFF3100BFFA7FF90FD7D0883FCAC8697" box="[740,846,1715,1737]" pageId="13" pageNumber="104" value="2009-05-21">
<collectingDate id="EFB7E9E3FFA7FF90FD7D0883FCAC8697" box="[740,846,1715,1737]" pageId="13" pageNumber="104" value="2009-05-21">21.05.2009</collectingDate>
</date>
,
<specimenCount id="9D4BFD42FFA7FF90FCCF0883FC668697" box="[854,900,1715,1737]" pageId="13" pageNumber="104" type="generic">1 ex.</specimenCount>
, leg.
<collectorName id="26B8531DFFA7FF90FC2A0883FBD98697" box="[947,1083,1715,1737]" pageId="13" pageNumber="104">Yu. Kanarskyi</collectorName>
(
<collectionCode id="ED5CAE0EFFA7FF90FBD10883FB718697" box="[1096,1171,1715,1737]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="13" pageNumber="104">SMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BF236CBFFA7FF90FF7A08DAFB32875A" blockId="13.[174,1232,1770,1828]" box="[227,1232,1770,1797]" pageId="13" pageNumber="104">
Distribution. Europe,
<collectingCountry id="F35A765BFFA7FF90FD8908DAFD7F875A" box="[528,669,1770,1796]" name="Turkey" pageId="13" pageNumber="104">Asia Minor</collectingCountry>
,
<collectingCountry id="F35A765BFFA7FF90FD3408DAFCFD875A" box="[685,799,1770,1796]" name="Mongolia" pageId="13" pageNumber="104">Mongolia</collectingCountry>
(
<bibRefCitation id="EFDC4B3AFFA7FF90FCA808DAFB8D875B" author="Schulke, M. &amp; Smetana, A." box="[817,1135,1770,1797]" pageId="13" pageNumber="104" pagination="1 - 1702" refId="ref15821" refString="Schulke, M., Smetana, A. 2015. Staphylinidae Latreille 1802, 304 - 1134. In: Lobl, I. &amp; Lobl, D., eds. Catalogue of Palaearctic Coleoptera vols. 1 &amp; 2, Hydrophiloidea - Staphylinoidea, revised and updated edition. Brill, Leiden &amp; Boston, I - XXV, 1 - 1702." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
). In the
</paragraph>
<paragraph id="8BF236CBFFA7FF93FF37093AFCA680B7" blockId="13.[174,1232,1770,1828]" lastBlockId="14.[166,1225,175,330]" lastPageId="14" lastPageNumber="105" pageId="13" pageNumber="104">
Ukrainian Carpathians known from Chernivtsi Region, Chernivtsi, mountain Tsetsyno, Bila (
<bibRefCitation id="EFDC4B3AFFA4FF93FF780E9FFE958097" author="Marcu, O." box="[225,375,175,201]" pageId="14" pageNumber="105" pagination="56 - 83" refId="ref15458" refString="Marcu, O. 1936. Coleopterenfunde aus der Bucovina. Verhandlungen und Mitteilungen des Siebenburgischen Vereins fur Naturwissenschaften, 15 - 16, 56 - 83." type="journal article" year="1936">Marcu, 1936</bibRefCitation>
) and Transcarpathian Region, mountain Pоp Ivan (
<bibRefCitation id="EFDC4B3AFFA4FF93FC540E9FFB5E8094" author="Fleischer, J. &amp; Mazura, K." box="[973,1212,175,202]" pageId="14" pageNumber="105" pagination="82 - 83" refId="ref14572" refString="Fleischer, J., Mazura, K. 1925. Ctvrty entomologicky zajezd do Podkarpatske Rusi. Sbornik klubu prirodovedeckeho v Brne za rok 1924. Brno, 82 - 83." type="journal article" year="1925">Fleischer et al., 1925</bibRefCitation>
). “
<collectingCountry id="F35A765BFFA4FF93FF280EFFFEEA80B7" box="[177,264,207,233]" name="Poland" pageId="14" pageNumber="105">Poland</collectingCountry>
”, without indication of region (
<bibRefCitation id="EFDC4B3AFFA4FF93FDE00EFFFCD580B7" author="Lomnicki, M." box="[633,823,207,233]" pageId="14" pageNumber="105" pagination="21 - 155" refId="ref15390" refString="Lomnicki, M. 1913. Wykaz chrzaszczow czyli Tegopokrywych (Coleoptera) ziem polskich. (Catalogus coleopterorum Poloniae). Kosmos. Seria A Biologia, 21 - 155." type="book chapter" year="1913">Łomnicki, 1913</bibRefCitation>
).
</paragraph>
<paragraph id="8BF236CBFFA4FF93FF420EDFFC0F8117" blockId="14.[166,1225,175,330]" pageId="14" pageNumber="105">
Bionomics. The species occurs mainly in lowlands and foothills in deciduous and mixed forests in forest litter and in rotten wood, in coastal sediments, in rotten fungi, on fermented birch sap, in straw and on animal cadavers (
<bibRefCitation id="EFDC4B3AFFA4FF93FCB10F1FFC028117" author="Semenov, V. B." box="[808,992,303,329]" pageId="14" pageNumber="105" pagination="24 - 52" refId="ref15760" refString="Semenov, V. B. 2007. Stafilinidy podsemeistva Aleocharinae (Coleoptera: Staphylinidae) Moskovskoy oblasti. Chast 1. Triby Deinopsini - Athetini. The staphylinid beetles subfamily Aleocharinae (Coleoptera: Staphylinidae) of the Moscow Area. Part 1. The tribes Deinopsini - Athetini. Eversmannia, 11 - 12, 24 - 52." type="journal article" year="2007">Semenov, 2007</bibRefCitation>
).
</paragraph>
<paragraph id="8BF236CBFFA4FF93FF3F0F5FFD5D81D4" blockId="14.[166,703,367,426]" box="[166,703,367,394]" pageId="14" pageNumber="105">
<emphasis id="B939EAD9FFA4FF93FF3F0F5FFEC581D7" bold="true" box="[166,295,367,393]" pageId="14" pageNumber="105">Subgenus</emphasis>
<taxonomicName id="4C4D4D48FFA4FF93FEB70F40FD5D81D4" authority="Mulsant &amp; Rey, 1873" authorityName="Mulsant &amp; Rey" authorityYear="1873" box="[302,703,368,394]" class="Insecta" family="Staphylinidae" genus="Microdota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="B939EAD9FFA4FF93FEB70F40FE5781D4" bold="true" box="[302,437,368,394]" italics="true" pageId="14" pageNumber="105">Microdota</emphasis>
Mulsant &amp; Rey, 1873
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3576540FFA4FF93FF3F0FBFFBCD8227" pageId="14" pageNumber="105" type="description">
<paragraph id="8BF236CBFFA4FF93FF3F0FBFFE3E81F7" blockId="14.[166,703,367,426]" box="[166,476,399,426]" pageId="14" pageNumber="105">
<taxonomicName id="4C4D4D48FFA4FF93FF3F0FBFFE3E81F7" authority="(Heer, 1841)" baseAuthorityName="Heer" baseAuthorityYear="1841" box="[166,476,399,426]" class="Insecta" family="Staphylinidae" genus="Atheta" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="105" phylum="Arthropoda" rank="species" species="aegra">
<emphasis id="B939EAD9FFA4FF93FF3F0FBFFEA381F4" bold="true" box="[166,321,399,426]" italics="true" pageId="14" pageNumber="105">Atheta aegra</emphasis>
(Heer, 1841)
</taxonomicName>
</paragraph>
<paragraph id="8BF236CBFFA4FF93FF420FF8FDF68180" blockId="14.[219,532,456,478]" box="[219,532,456,478]" pageId="14" pageNumber="105">M a t e r i a l. Literature data only.</paragraph>
<paragraph id="8BF236CBFFA4FF93FF420C30FBDA8267" blockId="14.[166,1225,511,633]" pageId="14" pageNumber="105">
D i s t r i b u t i o n. Europe,
<collectingCountry id="F35A765BFFA4FF93FE620FCFFD638247" box="[507,641,511,537]" name="Turkey" pageId="14" pageNumber="105">Asia Minor</collectingCountry>
,
<collectingCountry id="F35A765BFFA4FF93FD100FCFFD268247" box="[649,708,511,537]" name="Syria" pageId="14" pageNumber="105">Syria</collectingCountry>
(
<bibRefCitation id="EFDC4B3AFFA4FF93FD490FCFFC1B8244" author="Schulke, M. &amp; Smetana, A." box="[720,1017,511,538]" pageId="14" pageNumber="105" pagination="1 - 1702" refId="ref15821" refString="Schulke, M., Smetana, A. 2015. Staphylinidae Latreille 1802, 304 - 1134. In: Lobl, I. &amp; Lobl, D., eds. Catalogue of Palaearctic Coleoptera vols. 1 &amp; 2, Hydrophiloidea - Staphylinoidea, revised and updated edition. Brill, Leiden &amp; Boston, I - XXV, 1 - 1702." type="journal article" year="2015">Schülke &amp;Smetana, 2015</bibRefCitation>
). In the Ukrainian Carpathians known from Transcarpathian Region, Kuzii tract (
<bibRefCitation id="EFDC4B3AFFA4FF93FC120C2FFBC98267" author="Roubal, J." box="[907,1067,543,569]" pageId="14" pageNumber="105" pagination="1 - 527" refId="ref15689" refString="Roubal, J. 1930. Katalog Coleopter (brouku) Slovenska a Podkarpatska. Praha, 1 - 527." type="book chapter" year="1930">Roubal, 1930</bibRefCitation>
).
</paragraph>
<paragraph id="8BF236CBFFA4FF93FF420C70FBCD8227" blockId="14.[166,1225,511,633]" pageId="14" pageNumber="105">
B i o n o m i c s. The species occurs mainly in lowlands and foothills, in deciduous, mostly oak and beech forests, in forest litter and in remains of plants (
<bibRefCitation id="EFDC4B3AFFA4FF93FC1A0C6FFBC18227" author="Roubal, J." box="[899,1059,607,633]" pageId="14" pageNumber="105" pagination="1 - 527" refId="ref15689" refString="Roubal, J. 1930. Katalog Coleopter (brouku) Slovenska a Podkarpatska. Praha, 1 - 527." type="book chapter" year="1930">Roubal, 1930</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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