treatments-xml/data/03/BE/87/03BE87CFE70FFFFAFF57FDBBFE9DFFB8.xml

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<mods:title id="EEA0965F6E3ACE590BE1FFEF762A3315">Dolichopus nimbatus Parent, 1927 revisited, with two new synonyms (Diptera: Dolichopodidae)</mods:title>
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<heading id="D0E081B5E70FFFFEFF57FDBBFE4BFF24" bold="true" box="[151,500,544,571]" fontSize="11" level="1" pageId="1" pageNumber="144" reason="1">
<taxonomicName id="4C174D5AE70FFFFEFF57FDBBFE4BFF24" ID-CoL="8BTXK" authority="Parent" authorityName="Parent" box="[151,500,544,571]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70FFFFEFF57FDBBFE4BFF24" bold="true" box="[151,500,544,571]" pageId="1" pageNumber="144">
<emphasis id="B963EACBE70FFFFEFF57FDBBFE26FF25" bold="true" box="[151,409,544,570]" italics="true" pageId="1" pageNumber="144">Dolichopus nimbatus</emphasis>
Parent
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<paragraph id="8BA836D9E70FFFFEFF57FDF3FD90FF61" blockId="1.[151,675,616,699]" box="[151,559,616,638]" pageId="1" pageNumber="144">
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<taxonomicName id="4C174D5AE70FFFFEFF57FDF3FD94FF61" ID-CoL="8BTXK" authority="Parent, 1927: 454" authorityName="Parent" authorityPageNumber="454" authorityYear="1927" box="[151,555,616,638]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70FFFFEFF57FDF3FED0FF61" box="[151,367,616,638]" italics="true" pageId="1" pageNumber="144">Dolichopus nimbatus</emphasis>
<treatmentCitation id="0AB610C8E70FFFFEFEB5FDF2FD94FF61" author="Parent" box="[373,555,616,638]" page="454" pageId="1" pageNumber="144" year="1927">
<bibRefCitation id="EF864B28E70FFFFEFEB5FDF2FE47FF61" author="Parent" box="[373,504,616,638]" pageId="1" pageNumber="144" refString="Parent, O. (1927) Contribution a la distribution geographique de quelques especes de Dolichopodides. Congres des Societes Savantes. Paris et de departements. Comptes rendus, 1926, 449 - 484." type="journal article" year="1927">Parent, 1927</bibRefCitation>
: 454
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.
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</paragraph>
<paragraph id="8BA836D9E70FFFFEFF57FD1DFD38FFA5" blockId="1.[151,675,616,699]" pageId="1" pageNumber="144">
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<taxonomicName id="4C174D5AE70FFFFEFF57FD1DFDFFFF83" authority="Knezy, 1929: 19" authorityName="Knezy" authorityPageNumber="19" authorityYear="1929" box="[151,576,646,668]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="thalhammeri">
<emphasis id="B963EACBE70FFFFEFF57FD1DFE2CFF83" box="[151,403,646,668]" italics="true" pageId="1" pageNumber="144">Dolichopus thalhammeri</emphasis>
<treatmentCitation id="0AB610C8E70FFFFEFE5AFD1DFDFFFF83" author="Knezy" box="[410,576,646,668]" page="19" pageId="1" pageNumber="144" year="1929">
<bibRefCitation id="EF864B28E70FFFFEFE5AFD1DFDA2FF83" author="Knezy" box="[410,541,646,668]" pageId="1" pageNumber="144" refString="Knezy, G. (1929) Legyeszeti tanulmanyok. Folia Societatis Entomologicae Hungaricae, II (1), 1 - 20." type="journal article" year="1929">Knézy, 1929</bibRefCitation>
: 19
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<emphasis id="B963EACBE70FFFFEFD86FD1CFD1CFF83" bold="true" box="[582,675,647,668]" pageId="1" pageNumber="144">syn. nov.</emphasis>
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<taxonomicName id="4C174D5AE70FFFFEFF57FD3EFD9CFFA4" ID-CoL="92R84" authority="Meuffels, 1981: 327" authorityName="Meuffels" authorityPageNumber="327" authorityYear="1981" box="[151,547,677,699]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70FFFFEFF57FD3EFEEFFFA4" box="[151,336,677,699]" italics="true" pageId="1" pageNumber="144">Dolichopus balius</emphasis>
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: 327
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<emphasis id="B963EACBE70FFFFEFDEAFD3EFD38FFA5" bold="true" box="[554,647,677,698]" pageId="1" pageNumber="144">syn. nov.</emphasis>
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</paragraph>
<paragraph id="8BA836D9E70FFFFEFF57FD7CFA8FFE57" blockId="1.[151,1437,743,2015]" pageId="1" pageNumber="144">
<emphasis id="B963EACBE70FFFFEFF57FD7CFE3BFE1F" bold="true" box="[151,388,743,768]" pageId="1" pageNumber="144">Notes on synonymy.</emphasis>
In order to establish if all species were conspecific, the female 
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of 
<taxonomicName id="4C174D5AE70FFFFEFB0FFD72FA8AFE1F" ID-CoL="92R84" box="[1231,1333,744,768]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70FFFFEFB0FFD72FA8AFE1F" box="[1231,1333,744,768]" italics="true" pageId="1" pageNumber="144">D. balius</emphasis>
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was first compared to the female 
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of 
<taxonomicName id="4C174D5AE70FFFFEFDEBFC96FD0CFE3B" ID-CoL="8BTXK" box="[555,691,780,804]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70FFFFEFDEBFC96FD0CFE3B" box="[555,691,780,804]" italics="true" pageId="1" pageNumber="144">D. nimbatus</emphasis>
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. Subsequently, features listed in the original description of the lost 
<taxonomicName id="4C174D5AE70FFFFEFF57FCAAFEF7FE57" box="[151,328,816,840]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="thalhammeri">
<emphasis id="B963EACBE70FFFFEFF57FCAAFEF7FE57" box="[151,328,816,840]" italics="true" pageId="1" pageNumber="144">D. thalhammeri</emphasis>
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were checked in a male 
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of 
<taxonomicName id="4C174D5AE70FFFFEFD23FCAAFCF6FE57" ID-CoL="92R84" box="[739,841,816,840]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70FFFFEFD23FCAAFCF6FE57" box="[739,841,816,840]" italics="true" pageId="1" pageNumber="144">D. balius</emphasis>
</taxonomicName>
. All specimens were examined thoroughly.
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<paragraph id="8BA836D9E70FFFFEFF07FCCFFAA5F95A" blockId="1.[151,1437,743,2015]" pageId="1" pageNumber="144">
All differences between the female 
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of 
<taxonomicName id="4C174D5AE70FFFFEFD21FCCEFCD5FE73" box="[737,874,852,876]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70FFFFEFD21FCCEFCD5FE73" box="[737,874,852,876]" italics="true" pageId="1" pageNumber="144">D. nimbatus</emphasis>
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and female 
<typeStatus id="54AC887BE70FFFFEFC36FCCEFBE9FE72" box="[1014,1110,853,877]" pageId="1" pageNumber="144" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C174D5AE70FFFFEFBBDFCCEFB5BFE73" box="[1149,1252,852,876]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70FFFFEFBBDFCCEFB5BFE73" box="[1149,1252,852,876]" italics="true" pageId="1" pageNumber="144">D. balius</emphasis>
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were within the range of intraspecific variability and all important diagnostic features were identical or very comparable in both specimens. Also, the male 
<typeStatus id="54AC887BE70FFFFEFE03FC06FD9CFEAA" box="[451,547,925,949]" pageId="1" pageNumber="144" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C174D5AE70FFFFEFD89FC06FD10FEAB" box="[585,687,924,948]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70FFFFEFD89FC06FD10FEAB" box="[585,687,924,948]" italics="true" pageId="1" pageNumber="144">D. balius</emphasis>
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entirely matched the characters described by 
<bibRefCitation id="EF864B28E70FFFFEFB70FC07FAF4FEAB" author="Knezy" box="[1200,1355,924,948]" pageId="1" pageNumber="144" refString="Knezy, G. (1929) Legyeszeti tanulmanyok. Folia Societatis Entomologicae Hungaricae, II (1), 1 - 20." type="journal article" year="1929">Knézy (1929)</bibRefCitation>
for the male of 
<taxonomicName id="4C174D5AE70FFFFEFF36FC5AFE15FEC7" box="[246,426,960,984]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="thalhammeri">
<emphasis id="B963EACBE70FFFFEFF36FC5AFE15FEC7" box="[246,426,960,984]" italics="true" pageId="1" pageNumber="144">D. thalhammeri</emphasis>
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, except for some minor discrepancies in the chaetotaxy of the legs. The latter, however, could be explained as the mere result of a slightly different interpretation of character states by the respective authors, and/or of a slightly different accuracy. Both males also shared the important diagnostic features listed below. It could therefore be concluded that 
<taxonomicName id="4C174D5AE70FFFFEFDBDFBB6FCB9F95B" box="[637,774,1068,1092]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70FFFFEFDBDFBB6FCB9F95B" box="[637,774,1068,1092]" italics="true" pageId="1" pageNumber="144">D. nimbatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C174D5AE70FFFFEFCD4FBB6FCC5F95B" box="[788,890,1068,1092]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70FFFFEFCD4FBB6FCC5F95B" box="[788,890,1068,1092]" italics="true" pageId="1" pageNumber="144">D. balius</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C174D5AE70FFFFEFC71FBB6FBDDF95B" box="[945,1122,1068,1092]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="thalhammeri">
<emphasis id="B963EACBE70FFFFEFC71FBB6FBDDF95B" box="[945,1122,1068,1092]" italics="true" pageId="1" pageNumber="144">D. thalhammeri</emphasis>
</taxonomicName>
are conspecific.
</paragraph>
</subSubSection>
<subSubSection id="C30D6552E70FFFFEFF07FBD4FF42F83A" pageId="1" pageNumber="144" type="diagnosis">
<paragraph id="8BA836D9E70FFFFEFF07FBD4FF42F83A" blockId="1.[151,1437,743,2015]" pageId="1" pageNumber="144">
<emphasis id="B963EACBE70FFFFEFF07FBD4FE85F977" bold="true" box="[199,314,1103,1128]" pageId="1" pageNumber="144">Diagnosis</emphasis>
(both sexes). Wing (
<figureCitation id="132C2A5CE70FFFFEFDE5FBD4FDD3F977" box="[549,620,1103,1128]" captionStart="FIGURE 1" captionStartId="2.[151,249,1956,1978]" captionTargetBox="[151,1436,968,1932]" captionTargetId="figure@2.[151,1436,968,1932]" captionTargetPageId="2" captionText="FIGURE 1. Dolichopus nimbatus Parent, holotype (female) (habitus)." httpUri="https://zenodo.org/record/345072/files/figure.png" pageId="1" pageNumber="144">Figs 1</figureCitation>
, 
<figureCitation id="132C2A5CE70FFFFEFDBAFBCBFD35F977" box="[634,650,1104,1128]" captionStart="FIGURE 2" captionStartId="3.[151,249,1178,1200]" captionTargetBox="[151,1436,193,1157]" captionTargetId="figure@3.[151,1436,193,1157]" captionTargetPageId="3" captionText="FIGURE 2. Dolichopus balius Meuffels, male paratype (habitus)." httpUri="https://zenodo.org/record/345073/files/figure.png" pageId="1" pageNumber="144">2</figureCitation>
, 
<figureCitation id="132C2A5CE70FFFFEFD57FBD4FD1AF977" box="[663,677,1103,1128]" captionStart="FIGURE 5" captionStartId="6.[151,249,1325,1347]" captionTargetBox="[151,1436,339,1303]" captionTargetId="figure@6.[151,1436,339,1303]" captionTargetPageId="6" captionText="FIGURE 5. Dolichopus balius Meuffels, female paratype (habitus)." httpUri="https://zenodo.org/record/345076/files/figure.png" pageId="1" pageNumber="144">5</figureCitation>
) distinctly infuscate on about apical half anteriorly, at bend of M1+2 and at crossvein dm-cu. Bend of M1+2 oblique, with two short hanging veins. Insertion points of dorsocentral and other mesonotal bristles (
<figureCitation id="132C2A5CE70FFFFEFE7DFB06FDBBF9A9" box="[445,516,1181,1206]" captionStart="FIGURE 3" captionStartId="4.[151,249,1469,1491]" captionTargetBox="[151,1436,484,1448]" captionTargetId="figure@4.[151,1436,484,1448]" captionTargetPageId="4" captionText="FIGURE 3. Dolichopus balius Meuffels, male paratype (head)." httpUri="https://zenodo.org/record/345074/files/figure.png" pageId="1" pageNumber="144">Fig. 3</figureCitation>
), and of ad and pd bristles on tibiae II and III (
<figureCitation id="132C2A5CE70FFFFEFBFEFB06FB34F9A9" box="[1086,1163,1181,1206]" captionStart="FIGURE 2" captionStartId="3.[151,249,1178,1200]" captionTargetBox="[151,1436,193,1157]" captionTargetId="figure@3.[151,1436,193,1157]" captionTargetPageId="3" captionText="FIGURE 2. Dolichopus balius Meuffels, male paratype (habitus)." httpUri="https://zenodo.org/record/345073/files/figure.png" pageId="1" pageNumber="144">Figs 2</figureCitation>
, 
<figureCitation id="132C2A5CE70FFFFEFB5BFB06FB16F9A9" box="[1179,1193,1181,1206]" captionStart="FIGURE 5" captionStartId="6.[151,249,1325,1347]" captionTargetBox="[151,1436,339,1303]" captionTargetId="figure@6.[151,1436,339,1303]" captionTargetPageId="6" captionText="FIGURE 5. Dolichopus balius Meuffels, female paratype (habitus)." httpUri="https://zenodo.org/record/345076/files/figure.png" pageId="1" pageNumber="144">5</figureCitation>
) distinctly infuscate. Antenna (
<figureCitation id="132C2A5CE70FFFFEFEC8FB5AFEF2F9C5" box="[264,333,1217,1242]" captionStart="FIGURE 3" captionStartId="4.[151,249,1469,1491]" captionTargetBox="[151,1436,484,1448]" captionTargetId="figure@4.[151,1436,484,1448]" captionTargetPageId="4" captionText="FIGURE 3. Dolichopus balius Meuffels, male paratype (head)." httpUri="https://zenodo.org/record/345074/files/figure.png" pageId="1" pageNumber="144">Fig. 3</figureCitation>
) with first flagellomere subcircular, about as wide as long. Male (
<figureCitation id="132C2A5CE70FFFFEFB90FB5AFB26F9C5" box="[1104,1177,1217,1242]" captionStart="FIGURE 4" captionStartId="5.[151,249,1721,1743]" captionTargetBox="[151,1436,735,1699]" captionTargetId="figure@5.[151,1436,735,1699]" captionTargetPageId="5" captionText="FIGURE 4. Dolichopus balius Meuffels, male paratype (fore tarsus, ventral view)." httpUri="https://zenodo.org/record/345075/files/figure.png" pageId="1" pageNumber="144">Fig. 4</figureCitation>
): tibia I with one pale curved ventral preapical bristle, and tarsus I with a dense white ventral pilosity from apex of tarsomere taI1 onwards.
</paragraph>
</subSubSection>
<subSubSection id="C30D6552E70FFFFDFF07FAABFCAEFE8E" lastPageId="2" lastPageNumber="145" pageId="1" pageNumber="144" type="description">
<paragraph id="8BA836D9E70FFFFDFF07FAABFCAEFE8E" blockId="1.[151,1437,743,2015]" lastBlockId="2.[151,1437,151,913]" lastPageId="2" lastPageNumber="145" pageId="1" pageNumber="144">
<emphasis id="B963EACBE70FFFFEFF07FAABFE00F856" bold="true" box="[199,447,1328,1353]" pageId="1" pageNumber="144">Redescription. Male.</emphasis>
Body size: 
<quantity id="4CEF9B3CE70FFFFEFD85FAABFD22F857" box="[581,669,1328,1353]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="144" unit="mm" value="4.6">4.6 mm</quantity>
, wing size: 
<quantity id="4CEF9B3CE70FFFFEFCE2FAAAFCC4F857" box="[802,891,1329,1353]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="1" pageNumber="144" unit="mm" value="4.9">4.9 mm</quantity>
. 
<emphasis id="B963EACBE70FFFFEFC46FAABFC79F856" bold="true" box="[902,966,1328,1353]" pageId="1" pageNumber="144">Head</emphasis>
. Face silvery white, nearly parallel-sided, 1.2x as long as postpedicel (MSSC). Palpus very small, half moon-shaped, pale yellow, with black pubescence and one black apical bristle. About upper 1/3 of postoculars black, lowermost 2/3 white. Antenna (
<figureCitation id="132C2A5CE70FFFFEFB0EFAE3FAADF88E" box="[1230,1298,1400,1425]" captionStart="FIGURE 3" captionStartId="4.[151,249,1469,1491]" captionTargetBox="[151,1436,484,1448]" captionTargetId="figure@4.[151,1436,484,1448]" captionTargetPageId="4" captionText="FIGURE 3. Dolichopus balius Meuffels, male paratype (head)." httpUri="https://zenodo.org/record/345074/files/figure.png" pageId="1" pageNumber="144">Fig. 3</figureCitation>
) with scape and pedicel yellow, scape slightly infuscate dorsally; postpedicel dark brown, subcircular, as long as wide; stylus dorsal, inserted at about apical 1/3 of postpedicel, 1.7x as long as first three antennal segments combined, with very short pubescence. 
<emphasis id="B963EACBE70FFFFEFEA7FA7FFE78F8E2" bold="true" box="[359,455,1508,1533]" pageId="1" pageNumber="144">Thorax.</emphasis>
Mesonotum mainly bronze, dusted, with 2 rather wide green longitudinal bands along dorsocentral bristles, with distinct bronze insertion points of bristles; scutellum with marginal fringe of minute white setae, and a few minute setae on disk near anterior margin. Over 10 acrostichal bristles, biserial, reaching beyond 4th dorsocentral bristles. Bristle arrangement otherwise typical of 
<taxonomicName id="4C174D5AE70FFFFEFC18F9C8FBE6FB74" box="[984,1113,1619,1643]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="B963EACBE70FFFFEFC18F9C8FBE6FB74" box="[984,1113,1619,1643]" italics="true" pageId="1" pageNumber="144">Dolichopus</emphasis>
</taxonomicName>
. 
<emphasis id="B963EACBE70FFFFEFBA8F9C8FB16FB73" bold="true" box="[1128,1193,1619,1644]" pageId="1" pageNumber="144">Wing</emphasis>
(
<figureCitation id="132C2A5CE70FFFFEFB79F9C8FABDFB74" box="[1209,1282,1619,1644]" captionStart="FIGURE 1" captionStartId="2.[151,249,1956,1978]" captionTargetBox="[151,1436,968,1932]" captionTargetId="figure@2.[151,1436,968,1932]" captionTargetPageId="2" captionText="FIGURE 1. Dolichopus nimbatus Parent, holotype (female) (habitus)." httpUri="https://zenodo.org/record/345072/files/figure.png" pageId="1" pageNumber="144">Figs 1</figureCitation>
, 
<figureCitation id="132C2A5CE70FFFFEFAD1F9C8FA9EFB74" box="[1297,1313,1619,1643]" captionStart="FIGURE 2" captionStartId="3.[151,249,1178,1200]" captionTargetBox="[151,1436,193,1157]" captionTargetId="figure@3.[151,1436,193,1157]" captionTargetPageId="3" captionText="FIGURE 2. Dolichopus balius Meuffels, male paratype (habitus)." httpUri="https://zenodo.org/record/345073/files/figure.png" pageId="1" pageNumber="144">2</figureCitation>
, 
<figureCitation id="132C2A5CE70FFFFEFAEEF9C8FA83FB73" box="[1326,1340,1619,1644]" captionStart="FIGURE 5" captionStartId="6.[151,249,1325,1347]" captionTargetBox="[151,1436,339,1303]" captionTargetId="figure@6.[151,1436,339,1303]" captionTargetPageId="6" captionText="FIGURE 5. Dolichopus balius Meuffels, female paratype (habitus)." httpUri="https://zenodo.org/record/345076/files/figure.png" pageId="1" pageNumber="144">5</figureCitation>
) mainly hyaline, distinctly infuscate on about apical half anteriorly, at bend of M1+2 and at crossvein dm-cu. Vein M1+2 with oblique bend, bearing two short hanging veins (as in species of the 
<taxonomicName id="4C174D5AE70FFFFEFC5DF905FAFDFBA9" authority="Wiedemann, 1824" authorityName="Wiedemann" authorityYear="1824" box="[925,1346,1693,1718]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="144" phylum="Arthropoda" rank="species" species="ziczac">
<emphasis id="B963EACBE70FFFFEFC5DF905FBD3FBA9" box="[925,1132,1694,1718]" italics="true" pageId="1" pageNumber="144">Dolichopus ziczac</emphasis>
Wiedemann, 1824
</taxonomicName>
species group) with anterior one distinctly shorter than posterior one, slightly converging with vein R4+5 at wing apex. Proximal section of M1+2 only slightly longer than apical section. Proximal section of CuA1 2.7x as long as apical section. CuA x ratio 1.6. No anal lobe. Halter yellow, calypteral fringe with short yellowish white posterobasal setae, and long black apical setae. 
<emphasis id="B963EACBE70FFFFEFDF4F8ADFDD3FA50" bold="true" box="[564,620,1846,1871]" pageId="1" pageNumber="144">Legs</emphasis>
mainly yellow to brownish yellow, except for coxae II and III, with insertion points of ad and pd bristles on tibiae II and III distinctly infuscate (
<figureCitation id="132C2A5CE70FFFFEFC2FF8C1FB87FA6C" box="[1007,1080,1882,1907]" captionStart="FIGURE 2" captionStartId="3.[151,249,1178,1200]" captionTargetBox="[151,1436,193,1157]" captionTargetId="figure@3.[151,1436,193,1157]" captionTargetPageId="3" captionText="FIGURE 2. Dolichopus balius Meuffels, male paratype (habitus)." httpUri="https://zenodo.org/record/345073/files/figure.png" pageId="1" pageNumber="144">Figs 2</figureCitation>
, 
<figureCitation id="132C2A5CE70FFFFEFB84F8C1FBEBFA6C" box="[1092,1108,1882,1907]" captionStart="FIGURE 5" captionStartId="6.[151,249,1325,1347]" captionTargetBox="[151,1436,339,1303]" captionTargetId="figure@6.[151,1436,339,1303]" captionTargetPageId="6" captionText="FIGURE 5. Dolichopus balius Meuffels, female paratype (habitus)." httpUri="https://zenodo.org/record/345076/files/figure.png" pageId="1" pageNumber="144">5</figureCitation>
). Coxa I mainly yellow, with about basal 1/3 infuscate, and frontal face dusted greyish; with less than basal ½ with white pubescence, and more than apical ½ with black bristles. Coxae II and III dark brown, sometimes with yellow apex; coxa III with one black bristle at basal 3/5, and small white setae. Trochanters I-III yellow. All bristles of legs black, unless otherwise mentioned. Femur I entirely dark yellow, with large pv zone with sparse white pubescence (MSSC). Femur II dark yellow, brownish ventrally along basal 2/3, with one large preapical ad bristle. Femur III dark yellow with dark brown knee, and extreme apex whitish yellow; with ventral ¼ (width) brownish along basal 2/3; with one ad preapical bristle and without erect basodorsal bristles. Tibia I yellow, with black apex on anterior face, with 4 strong ds and 2 vt bristles; with ad serration of short bristles mainly on apical ½; with one black ap and one curved pale (brownish at basis, white at apex) preapical av bristle (
<figureCitation id="132C2A5CE70CFFFDFCEAFED7FCD2FC7B" box="[810,877,332,357]" captionStart="FIGURE 4" captionStartId="5.[151,249,1721,1743]" captionTargetBox="[151,1436,735,1699]" captionTargetId="figure@5.[151,1436,735,1699]" captionTargetPageId="5" captionText="FIGURE 4. Dolichopus balius Meuffels, male paratype (fore tarsus, ventral view)." httpUri="https://zenodo.org/record/345075/files/figure.png" pageId="2" pageNumber="145">Fig. 4</figureCitation>
) (MSSC). Tibia II whitish yellow, with 4 ad, 2 pd, and 5 ap bristles, and 1 av bristle at apical 2/5. Tibia III whitish yellow, with apical 1/10 brown; with 5 ad and 4 pd bristles, one ds preapical and 2 ap bristles; with 3 small vt bristles. Tarsus I (
<figureCitation id="132C2A5CE70CFFFDFC28FE0FFB94FCB3" box="[1000,1067,404,429]" captionStart="FIGURE 4" captionStartId="5.[151,249,1721,1743]" captionTargetBox="[151,1436,735,1699]" captionTargetId="figure@5.[151,1436,735,1699]" captionTargetPageId="5" captionText="FIGURE 4. Dolichopus balius Meuffels, male paratype (fore tarsus, ventral view)." httpUri="https://zenodo.org/record/345075/files/figure.png" pageId="2" pageNumber="145">Fig. 4</figureCitation>
) yellow, darker from apex of taI1 onwards; with dense vt white pilosity on about apical ¼ of taI1, and on taI2-5 (MSSC). Tarsus II yellow with taII1-2 darker on apex, taII3 dark on apical 1/2, and taII4-5 entirely dark. Tarsus III brownish yellow, with taIII1-4 darkened on apex, and taIII5 entirely dark; taIII1 with 1 dorsal bristle. Size (length) ratio of femur/tibia/tarsomeres1-5, in leg I: 5.3/9.5/4.3/2.2/1.7/1.2/1, in leg II: 6.6/7.6/4.3/2.7/1.6/1/1, and in leg III: 6.7/9.9/3.3/3.3/2.1/1.4/1. 
<emphasis id="B963EACBE70CFFFDFB2DFDB4FADEFF57" bold="true" box="[1261,1377,559,584]" pageId="2" pageNumber="145">Abdomen</emphasis>
with tergites brilliantly green, mainly dusted laterally, coppery near anterior and posterior margins; sternites black to coppery, tergite I with small white setae laterally. Hypopygium rather small, reaching posterior 1/3 of tergite III; without conspicuous epandrial lobes or surstylar processes, cercus small, white with narrow dark brown margin, and simple marginal bristles (
<figureCitation id="132C2A5CE70CFFFDFE25FD24FD94FFC8" box="[485,555,703,728]" captionStart="FIGURE 2" captionStartId="3.[151,249,1178,1200]" captionTargetBox="[151,1436,193,1157]" captionTargetId="figure@3.[151,1436,193,1157]" captionTargetPageId="3" captionText="FIGURE 2. Dolichopus balius Meuffels, male paratype (habitus)." httpUri="https://zenodo.org/record/345073/files/figure.png" pageId="2" pageNumber="145">Fig. 2</figureCitation>
; see also 
<bibRefCitation id="EF864B28E70CFFFDFD58FD24FCFAFFC8" author="Meuffels" box="[664,837,703,728]" pageId="2" pageNumber="145" refString="Meuffels, H. J. G. (1981) Dolichopus balius et Dolichopus ptenopedilus nouveaux Dolichopodidae palearctiques (Diptera). Bulletin des Recherches Agronomiques de Gembloux, 16 (4), 327 - 334." type="journal article" year="1981">Meuffels, 1981</bibRefCitation>
, fig. 3). 
<emphasis id="B963EACBE70CFFFDFC66FD24FBBBFFC7" bold="true" box="[934,1028,703,728]" pageId="2" pageNumber="145">Female.</emphasis>
Body size: 
<quantity id="4CEF9B3CE70CFFFDFB4BFD24FB5BFFC8" box="[1163,1252,703,728]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="145" unit="mm" value="4.6">4.6 mm</quantity>
, wing size: 
<quantity id="4CEF9B3CE70CFFFDFAABFD24FF4EFFE4" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.8" metricValueMin="4.4" pageId="2" pageNumber="145" unit="mm" value="4.6" valueMax="4.8" valueMin="4.4">4.4– 4.8 mm</quantity>
. As male, except for: face 1.7x as wide as postpedicel, latter slightly wider than long. Femur I with small elongated vt bare area near base. Tibia I without curved preapical av bristle. Tarsus I pale yellow, dark brown from apical ½ of tarsomere 
<date id="FFA91019E70CFFFDFE4FFCB7FE1FFE57" box="[399,416,812,840]" pageId="2" pageNumber="145">I3</date>
onwards, without ventral pilosity. Tarsi II and III pale yellow, with ta1-4 slightly infuscate at apex, and ta5 dark brown. Size (length) ratio of femur/tibia/tarsomeres1-5, in leg I: 9.0/9.3/4.8/2.0/1.5/1/1.3, in leg II: 7.8/8.8/4.5/1.8/1.7/1/1, in leg III: 6.7/9.6/2.9/3.0/1.9/1/1.
</paragraph>
</subSubSection>
<caption id="DF686651E70CFFFDFF57F83FFCDFFAA5" box="[151,864,1956,1978]" httpUri="https://zenodo.org/record/345072/files/figure.png" pageId="2" pageNumber="145" targetBox="[151,1436,968,1932]" targetPageId="2">
<paragraph id="8BA836D9E70CFFFDFF57F83FFCDFFAA5" blockId="2.[151,864,1956,1978]" box="[151,864,1956,1978]" pageId="2" pageNumber="145">
<emphasis id="B963EACBE70CFFFDFF57F83FFEADFAA6" bold="true" box="[151,274,1956,1978]" pageId="2" pageNumber="145">FIGURE 1.</emphasis>
<taxonomicName id="4C174D5AE70CFFFDFED8F83FFD85FAA5" authority="Parent" authorityName="Parent" box="[280,570,1956,1978]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="145" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70CFFFDFED8F83FFE4EFAA5" box="[280,497,1956,1978]" italics="true" pageId="2" pageNumber="145">Dolichopus nimbatus</emphasis>
Parent
</taxonomicName>
, holotype (female) (habitus).
</paragraph>
</caption>
<caption id="DF686651E70DFFFCFF57FB01FC8AF9AF" box="[151,821,1178,1200]" httpUri="https://zenodo.org/record/345073/files/figure.png" pageId="3" pageNumber="146" targetBox="[151,1436,193,1157]" targetPageId="3">
<paragraph id="8BA836D9E70DFFFCFF57FB01FC8AF9AF" blockId="3.[151,821,1178,1200]" box="[151,821,1178,1200]" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFF57FB01FEADF9B0" bold="true" box="[151,274,1178,1200]" pageId="3" pageNumber="146">FIGURE 2.</emphasis>
<taxonomicName id="4C174D5AE70DFFFCFED8FB01FD89F9AF" authority="Meuffels" authorityName="Meuffels" box="[280,566,1178,1200]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70DFFFCFED8FB01FE32F9AF" box="[280,397,1178,1200]" italics="true" pageId="3" pageNumber="146">Dolichopus</emphasis>
<emphasis id="B963EACBE70DFFFCFE54FB01FE6DF9AF" box="[404,466,1178,1200]" italics="true" pageId="3" pageNumber="146">balius</emphasis>
Meuffels
</taxonomicName>
, male paratype (habitus).
</paragraph>
</caption>
<subSubSection id="C30D6552E70DFFFCFF07FB46FDA3FA61" pageId="3" pageNumber="146" type="materials_examined">
<paragraph id="8BA836D9E70DFFFCFF07FB46FBB4F821" blockId="3.[151,1437,1245,2030]" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFF07FB46FE50F9E9" bold="true" box="[199,495,1245,1270]" pageId="3" pageNumber="146">
<typeStatus id="54AC887BE70DFFFCFF07FB46FEBDF9E9" box="[199,258,1245,1270]" pageId="3" pageNumber="146">Type</typeStatus>
material examined.
</emphasis>
<taxonomicName id="4C174D5AE70DFFFCFE36FB46FD5BF9EA" box="[502,740,1245,1269]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70DFFFCFE36FB46FD5BF9EA" box="[502,740,1245,1269]" italics="true" pageId="3" pageNumber="146">Dolichopus nimbatus</emphasis>
</taxonomicName>
: 
<typeStatus id="54AC887BE70DFFFCFD34FB46FC2CF9E9" box="[756,915,1245,1270]" pageId="3" pageNumber="146" type="holotype">
<emphasis id="B963EACBE70DFFFCFD34FB46FC2CF9E9" bold="true" box="[756,915,1245,1270]" pageId="3" pageNumber="146">HOLOTYPE</emphasis>
</typeStatus>
<specimenCount id="9D11FD50E70DFFFCFC5AFB45FC10F9EA" box="[922,943,1246,1269]" pageId="3" pageNumber="146" type="female">♀</specimenCount>
[white], “ 
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” / “ 
<collectionCode id="ED06AE1CE70DFFFCFADEFB45FACFF9E9" box="[1310,1392,1246,1270]" pageId="3" pageNumber="146">CAMP</collectionCode>
DE 
<collectionCode id="ED06AE1CE70DFFFCFF57FA99FE97F806" box="[151,296,1282,1305]" pageId="3" pageNumber="146">ZEITENLIK</collectionCode>
[
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]” / “prês SALONIQUE” / “ 
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D’ORIENT” / “D’RIVET 1917”; [white] “OCTOBRE”; [white] “ 
<typeStatus id="54AC887BE70DFFFCFF39FABDFE8AF821" box="[249,309,1318,1342]" pageId="3" pageNumber="146">Type</typeStatus>
” / “ 
<taxonomicName id="4C174D5AE70DFFFCFE9FFABEFE5AF821" box="[351,485,1317,1342]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="genus">Dolichopus</taxonomicName>
” / “ 
<taxonomicName id="4C174D5AE70DFFFCFDCFFABEFD76F821" authority="Parent" authorityName="Parent" box="[527,713,1317,1342]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="nimbatus">nimbatus Parent</taxonomicName>
” / “det O. Parent (
<collectionCode id="ED06AE1CE70DFFFCFC57FABDFC4AF821" box="[919,1013,1318,1342]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="146">MNHN</collectionCode>
)”.
</paragraph>
<paragraph id="8BA836D9E70DFFFCFF07FAD2FB07F8B6" blockId="3.[151,1437,1245,2030]" pageId="3" pageNumber="146">
<materialsCitation id="3B7F3C84E70DFFFCFF07FAD2FCF6F899" ID-GBIF-Occurrence="1437764343" collectingDate="79-07-25" collectionCode="GARD, FSAG" collectorName="La Capelle &amp; H. G. J. Meuffels" country="France" location="France" pageId="3" pageNumber="146" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<taxonomicName id="4C174D5AE70DFFFCFF07FAD2FE2BF87E" box="[199,404,1353,1377]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70DFFFCFF07FAD2FE2BF87E" box="[199,404,1353,1377]" italics="true" pageId="3" pageNumber="146">Dolichopus balius</emphasis>
</taxonomicName>
: 
<specimenCount id="9D11FD50E70DFFFCFE64FAD1FE06F87E" box="[420,441,1354,1377]" pageId="3" pageNumber="146" type="male">♂</specimenCount>
[white], “ 
<collectionCode id="ED06AE1CE70DFFFCFDEAFAD2FDC3F87E" box="[554,636,1353,1377]" pageId="3" pageNumber="146">GARD</collectionCode>
[
<collectingCountry id="F3007649E70DFFFCFD4AFAD2FD62F87E" box="[650,733,1353,1377]" name="France" pageId="3" pageNumber="146">France</collectingCountry>
]”/ “ 
<collectorName id="26E2530FE70DFFFCFCC9FAD2FC37F87E" box="[777,904,1353,1377]" pageId="3" pageNumber="146">La Capelle</collectorName>
” / “ 
<collectingDate id="EFEDE9F1E70DFFFCFC74FAD2FBB0F87E" box="[948,1039,1353,1377]" pageId="3" pageNumber="146" value="79-07-25">
<date id="FFA91019E70DFFFCFC74FAD2FBB0F87E" box="[948,1039,1353,1377]" pageId="3" pageNumber="146" value="79-07-25">25.7.79</date>
</collectingDate>
”; [white] “ 
<taxonomicName id="4C174D5AE70DFFFCFB49FAD2FAB2F87E" box="[1161,1293,1353,1377]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="genus">Dolichopus</taxonomicName>
<specimenCount id="9D11FD50E70DFFFCFAD4FAD2FA95F87E" box="[1300,1322,1353,1377]" pageId="3" pageNumber="146" type="male">♂</specimenCount>
” / “ 
<taxonomicName id="4C174D5AE70DFFFCFA97FAD2FF41F899" authority="Meuffels" authorityName="Meuffels" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="balius">balius Meuffels</taxonomicName>
” / “det 
<collectorName id="26E2530FE70DFFFCFE99FAF5FDADF899" box="[345,530,1389,1414]" pageId="3" pageNumber="146">H.G.J. Meuffels</collectorName>
”; [red] “ 
<typeStatus id="54AC887BE70DFFFCFDB3FAF5FD69F899" box="[627,726,1390,1414]" pageId="3" pageNumber="146" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED06AE1CE70DFFFCFD27FAF5FC89F899" box="[743,822,1390,1414]" country="Belgium" httpUri="http://grbio.org/cool/ms79-qhaw" name="Faculte des Sciences Agronomiques de Gembloux" pageId="3" pageNumber="146">FSAG</collectionCode>
)”
</materialsCitation>
. ♀ [white], “GARD” / “La Capelle” / “ 
<date id="FFA91019E70DFFFCFAD1FAF6FA33F899" box="[1297,1420,1389,1414]" pageId="3" pageNumber="146" value="1980-04-20">20.iv.1980</date>
”; [white] “ 
<taxonomicName id="4C174D5AE70DFFFCFF39FA0AFEC2F8B5" box="[249,381,1425,1450]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="genus">Dolichopus</taxonomicName>
♀” / “ 
<taxonomicName id="4C174D5AE70DFFFCFE05FA0AFDC7F8B5" authority="Meuffels" authorityName="Meuffels" box="[453,632,1425,1450]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="balius">balius Meuffels</taxonomicName>
” / “det H.G.J. Meuffels”; [red] “paratype (FSAG)”.
</paragraph>
<paragraph id="8BA836D9E70DFFFCFF07FA2EFDFEFBB9" blockId="3.[151,1437,1245,2030]" pageId="3" pageNumber="146">
<materialsCitation id="3B7F3C84E70DFFFCFF07FA2EFC60FB09" ID-GBIF-Occurrence="1437764342" collectingDate="2004-10-02" collectingDateMax="2004-10-08" collectingDateMin="2004-10-02" collectionCode="HELI" collectorName="W. D'Haeseleer &amp; Pollet &amp; Belgium" country="Belgium" location="Honegem Nature Reserve" municipality="Province Oost-Vlaanderen" pageId="3" pageNumber="146" specimenCount="1" specimenCount-male="1">
<emphasis id="B963EACBE70DFFFCFF07FA2EFD6CF8D1" bold="true" box="[199,723,1461,1486]" pageId="3" pageNumber="146">
Additional material examined. 
<collectingCountry id="F3007649E70DFFFCFD8CFA2EFD6CF8D1" box="[588,723,1461,1486]" name="Belgium" pageId="3" pageNumber="146">BELGIUM</collectingCountry>
</emphasis>
: 
<specimenCount id="9D11FD50E70DFFFCFD28FA2DFCA5F8D2" box="[744,794,1462,1486]" pageId="3" pageNumber="146" type="male">1 ♂</specimenCount>
, 
<collectingMunicipality id="6BCCACA3E70DFFFCFCECFA2EFBE1F8D1" box="[812,1118,1461,1486]" pageId="3" pageNumber="146">Province Oost-Vlaanderen</collectingMunicipality>
, 
<location id="8EC86002E70DFFFCFBAFFA2DFA23F8D2" LSID="urn:lsid:plazi:treatment:03BE87CFE70FFFFAFF57FDBBFE9DFFB8:8EC86002E70DFFFCFBAFFA2DFA23F8D2" box="[1135,1436,1462,1486]" country="Belgium" municipality="Province Oost-Vlaanderen" name="Honegem Nature Reserve" pageId="3" pageNumber="146">Honegem Nature Reserve</location>
(
<locationDeviation id="9A1A2FFBE70DFFFCFF5EFA42FDB9F8ED" box="[158,518,1497,1522]" pageId="3" pageNumber="146">between Erpe-Mere and Lede</locationDeviation>
), sampling site 
<collectionCode id="ED06AE1CE70DFFFCFD0AFA41FCB5F8EE" box="[714,778,1498,1521]" pageId="3" pageNumber="146">HELI</collectionCode>
(WID/2004/03), ES7045, open marshland, 
<collectingDate id="EFEDE9F1E70DFFFCFAD7FA42FA27F8EE" box="[1303,1432,1497,1521]" pageId="3" pageNumber="146" value="2004-10-02" valueMax="2004-10-08" valueMin="2004-10-02">
<date id="FFA91019E70DFFFCFAD7FA42FA27F8EE" box="[1303,1432,1497,1521]" pageId="3" pageNumber="146" value="2004-10-02" valueMax="2004-10-08" valueMin="2004-10-02">2–8.x.2004</date>
</collectingDate>
, 
<collectingMethod id="52564ECEE70DFFFCFF57FA66FE9BFB0A" box="[151,292,1533,1558]" pageId="3" pageNumber="146">Malaise trap</collectingMethod>
, leg. 
<collectorName id="26E2530FE70DFFFCFEA0FA65FDB2FB09" box="[352,525,1533,1558]" pageId="3" pageNumber="146">W. D'Haeseleer</collectorName>
(
<bibRefCitation id="EF864B28E70DFFFCFDDBFA66FD21FB09" author="Pollet" box="[539,670,1533,1558]" pageId="3" pageNumber="146" refString="Pollet, M. (2012) ' Uitgestorven' slankpootvliegen, en nieuwe soorten voor Belgie, ontdekt in de Denderstreek. Dendriet, 11 (1), 22 - 27." type="journal article" year="2012">
<collectorName id="26E2530FE70DFFFCFDDBFA66FDE1FB09" box="[539,606,1533,1558]" pageId="3" pageNumber="146">Pollet</collectorName>
2012
</bibRefCitation>
)— 
<emphasis id="B963EACBE70DFFFCFD03FA66FC60FB09" bold="true" box="[707,991,1533,1558]" pageId="3" pageNumber="146">
first record for 
<collectorName id="26E2530FE70DFFFCFCBBFA66FC60FB09" box="[891,991,1533,1558]" pageId="3" pageNumber="146">Belgium</collectorName>
</emphasis>
</materialsCitation>
. 
<materialsCitation id="3B7F3C84E70DFFFCFC2DFA66FBB7FB41" ID-GBIF-Occurrence="1437764347" collectingDate="1997-06" collectingDateMax="1997-08-31" collectingDateMin="1997-06" collectorName="Departement du Nord &amp; Mare de Prussien &amp; C. Sibold" country="France" county="Departement du Nord" latitude="50.408108" location="Parc Naturel Regional Scarpe-Escaut" longLatPrecision="1" longitude="3.4495668" municipality="Raismes-Vicoigne" pageId="3" pageNumber="146" specimenCount="1" specimenCount-female="1" stateProvince="Region Nord-Pas-de-Calais">
<collectingCountry id="F3007649E70DFFFCFC2DFA66FBDEFB09" box="[1005,1121,1533,1558]" name="France" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFC2DFA66FBDEFB09" bold="true" box="[1005,1121,1533,1558]" pageId="3" pageNumber="146">FRANCE</emphasis>
</collectingCountry>
: 
<specimenCount id="9D11FD50E70DFFFCFBB0FA65FB25FB0A" box="[1136,1178,1534,1558]" pageId="3" pageNumber="146" type="female">1 ♀</specimenCount>
, 
<collectorName id="26E2530FE70DFFFCFB68FA65FA23FB09" box="[1192,1436,1533,1558]" pageId="3" pageNumber="146">
<collectingCounty id="62C94E55E70DFFFCFB68FA65FA23FB09" box="[1192,1436,1533,1558]" pageId="3" pageNumber="146">Département du Nord</collectingCounty>
</collectorName>
(
<collectingRegion id="49D3F83BE70DFFFCFF5EF9BAFE5BFB25" box="[158,484,1569,1594]" country="France" name="Nord-Pas-de-Calais" pageId="3" pageNumber="146">Région Nord-Pas-de-Calais</collectingRegion>
), 
<collectingMunicipality id="6BCCACA3E70DFFFCFDC1F9BAFD6EFB25" box="[513,721,1569,1594]" pageId="3" pageNumber="146">Raismes-Vicoigne</collectingMunicipality>
, 
<location id="8EC86002E70DFFFCFD26F9B9FB17FB26" LSID="urn:lsid:plazi:treatment:03BE87CFE70FFFFAFF57FDBBFE9DFFB8:8EC86002E70DFFFCFD26F9B9FB17FB26" box="[742,1192,1569,1594]" country="France" county="Departement du Nord" latitude="50.408108" longLatPrecision="1" longitude="3.4495668" municipality="Raismes-Vicoigne" name="Parc Naturel Regional Scarpe-Escaut" pageId="3" pageNumber="146" stateProvince="Region Nord-Pas-de-Calais">Parc Naturel Régional Scarpe-Escaut</location>
, 
<collectorName id="26E2530FE70DFFFCFB7EF9B9FA27FB25" box="[1214,1432,1569,1594]" pageId="3" pageNumber="146">
<location id="8EC86002E70DFFFCFB7EF9B9FA27FB25" LSID="urn:lsid:plazi:treatment:03BE87CFE70FFFFAFF57FDBBFE9DFFB8:8EC86002E70DFFFCFB7EF9B9FA27FB25" box="[1214,1432,1569,1594]" country="France" county="Departement du Nord" latitude="50.408108" longLatPrecision="1" longitude="3.4495668" municipality="Raismes-Vicoigne" name="Mare de Prussien" pageId="3" pageNumber="146" stateProvince="Region Nord-Pas-de-Calais">Mare de Prussien</location>
</collectorName>
, 
<geoCoordinate id="EE23501EE70DFFFCFF57F9DEFEFDFB41" box="[151,322,1605,1630]" direction="north" orientation="latitude" pageId="3" pageNumber="146" precision="1" value="50.408108">50°24′29.18″N</geoCoordinate>
, 
<geoCoordinate id="EE23501EE70DFFFCFE91F9DEFE54FB41" box="[337,491,1605,1630]" direction="east" orientation="longitude" pageId="3" pageNumber="146" precision="1" value="3.4495668">3°26′58.44″E</geoCoordinate>
, marshland, 
<collectingDate id="EFEDE9F1E70DFFFCFD42F9DEFCB3FB41" box="[642,780,1605,1630]" pageId="3" pageNumber="146" value="1997-06" valueMax="1997-08-31" valueMin="1997-06-00">
<date id="FFA91019E70DFFFCFD42F9DEFCB3FB41" box="[642,780,1605,1630]" pageId="3" pageNumber="146" value="1997-06" valueMax="1997-08-31" valueMin="1997-06-00">vi–viii.1997</date>
</collectingDate>
, YPT, leg. 
<collectorName id="26E2530FE70DFFFCFC56F9DDFBB7FB41" box="[918,1032,1605,1630]" pageId="3" pageNumber="146">C. Sibold</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B7F3C84E70DFFFCFBD6F9DEFDFEFBB9" ID-GBIF-Occurrence="1437764341" collectingDate="2008-03-10" collectingDateMax="2008-03-16" collectingDateMin="2008-03-10" collectionCode="MAPC" collectorName="G. Ramel" country="Greece" county="Province Serres" elevation="40" latitude="41.213" location="Lake Kerkini" longLatPrecision="1" longitude="23.095251" municipality="Kerkini" pageId="3" pageNumber="146" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F3007649E70DFFFCFBD6F9DEFB33FB41" box="[1046,1164,1605,1630]" name="Greece" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFBD6F9DEFB33FB41" bold="true" box="[1046,1164,1605,1630]" pageId="3" pageNumber="146">GREECE</emphasis>
</collectingCountry>
: 
<specimenCount id="9D11FD50E70DFFFCFB5FF9DDFB6FFB42" box="[1183,1232,1606,1630]" pageId="3" pageNumber="146" type="male">1 ♂</specimenCount>
, 
<collectingCounty id="62C94E55E70DFFFCFB20F9DEFA27FB41" box="[1248,1432,1605,1630]" pageId="3" pageNumber="146">Province Serres</collectingCounty>
, 
<collectingMunicipality id="6BCCACA3E70DFFFCFF57F9F2FF50FB9D" box="[151,239,1641,1666]" pageId="3" pageNumber="146">Kerkini</collectingMunicipality>
, 
<location id="8EC86002E70DFFFCFF3DF9F2FE28FB9D" LSID="urn:lsid:plazi:treatment:03BE87CFE70FFFFAFF57FDBBFE9DFFB8:8EC86002E70DFFFCFF3DF9F2FE28FB9D" box="[253,407,1641,1666]" country="Greece" county="Province Serres" latitude="41.213" longLatPrecision="1" longitude="23.095251" municipality="Kerkini" name="Lake Kerkini" pageId="3" pageNumber="146">Lake Kerkini</location>
, 
<location id="8EC86002E70DFFFCFE65F9F2FD93FB9D" LSID="urn:lsid:plazi:treatment:03BE87CFE70FFFFAFF57FDBBFE9DFFB8:8EC86002E70DFFFCFE65F9F2FD93FB9D" box="[421,556,1641,1666]" country="Greece" county="Province Serres" latitude="41.213" longLatPrecision="1" longitude="23.095251" municipality="Kerkini" name="Cafe Elodia" pageId="3" pageNumber="146">Café Elodia</location>
(sampling site), 
<geoCoordinate id="EE23501EE70DFFFCFD31F9F2FC31FB9D" box="[753,910,1641,1666]" direction="north" orientation="latitude" pageId="3" pageNumber="146" precision="1" value="41.213">41°12′46.8″N</geoCoordinate>
, 
<geoCoordinate id="EE23501EE70DFFFCFC5CF9F2FB8AFB9D" box="[924,1077,1641,1666]" direction="east" orientation="longitude" pageId="3" pageNumber="146" precision="1" value="23.095251">23°05′42.9″E</geoCoordinate>
, 
<elevation id="003AD1EAE70DFFFCFB83F9F2FB3EFB9E" box="[1091,1153,1641,1665]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="146" unit="m" value="40.0">
<quantity id="4CEF9B3CE70DFFFCFB83F9F2FB3EFB9E" box="[1091,1153,1641,1665]" metricMagnitude="1" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="146" unit="m" value="40.0">40 m</quantity>
</elevation>
, 
<collectingDate id="EFEDE9F1E70DFFFCFB50F9F2FA89FB9D" box="[1168,1334,1641,1666]" pageId="3" pageNumber="146" value="2008-03-10" valueMax="2008-03-16" valueMin="2008-03-10">
<date id="FFA91019E70DFFFCFB50F9F2FA89FB9D" box="[1168,1334,1641,1666]" pageId="3" pageNumber="146" value="2008-03-10" valueMax="2008-03-16" valueMin="2008-03-10">10–16.iii.2008</date>
</collectingDate>
, 
<collectingMethod id="52564ECEE70DFFFCFA83F9F2FF7AFBBA" pageId="3" pageNumber="146">Malaise trap</collectingMethod>
, leg. 
<collectorName id="26E2530FE70DFFFCFEC0F915FED7FBB9" box="[256,360,1677,1702]" pageId="3" pageNumber="146">G. Ramel</collectorName>
(all W; in 
<collectionCode id="ED06AE1CE70DFFFCFE20F915FD8AFBB9" box="[480,565,1678,1702]" pageId="3" pageNumber="146">MAPC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA836D9E70DFFFCFF07F92AFDA3FA61" blockId="3.[151,1437,1245,2030]" pageId="3" pageNumber="146">
<materialsCitation id="3B7F3C84E70DFFFCFF07F92AFA27FBD6" ID-GBIF-Occurrence="1437764348" box="[199,1432,1713,1738]" collectorName="Knezy" country="Bulgaria" location="Other" pageId="3" pageNumber="146" specimenCount="1" specimenCount-female="1">
<emphasis id="B963EACBE70DFFFCFF07F92AFEC9FBD5" bold="true" box="[199,374,1713,1738]" pageId="3" pageNumber="146">
<location id="8EC86002E70DFFFCFF07F92AFEB0FBD5" LSID="urn:lsid:plazi:treatment:03BE87CFE70FFFFAFF57FDBBFE9DFFB8:8EC86002E70DFFFCFF07F92AFEB0FBD5" box="[199,271,1713,1738]" country="Bulgaria" name="Other" pageId="3" pageNumber="146">Other</location>
records
</emphasis>
(specimens not examined). 
<collectingCountry id="F3007649E70DFFFCFD0AF92AFCDEFBD5" box="[714,865,1713,1738]" name="Bulgaria" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFD0AF92AFCDEFBD5" bold="true" box="[714,865,1713,1738]" pageId="3" pageNumber="146">BULGARIA</emphasis>
</collectingCountry>
: 
<specimenCount id="9D11FD50E70DFFFCFCB6F92AFC18FBD6" box="[886,935,1713,1737]" pageId="3" pageNumber="146" type="female">1 ♀</specimenCount>
, no additional information (
<bibRefCitation id="EF864B28E70DFFFCFB3BF92AFA2FFBD6" author="Knezy" box="[1275,1424,1713,1737]" pageId="3" pageNumber="146" refString="Knezy, G. (1929) Legyeszeti tanulmanyok. Folia Societatis Entomologicae Hungaricae, II (1), 1 - 20." type="journal article" year="1929">
<collectorName id="26E2530FE70DFFFCFB3BF92AFAF9FBD6" box="[1275,1350,1713,1737]" pageId="3" pageNumber="146">Knézy</collectorName>
1929
</bibRefCitation>
)
</materialsCitation>
. 
<materialsCitation id="3B7F3C84E70DFFFCFF57F94EFB7FFA0D" ID-GBIF-Occurrence="1437764344" collectorName="W. Emeis &amp; W. Emeis Collection" country="Germany" location="Schleswig-Holstein" pageId="3" pageNumber="146" specimenCount="1" stateProvince="Schleswig-Holstein">
<collectingCountry id="F3007649E70DFFFCFF57F94EFE97FBF1" box="[151,296,1749,1774]" name="Germany" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFF57F94EFE97FBF1" bold="true" box="[151,296,1749,1774]" pageId="3" pageNumber="146">GERMANY</emphasis>
</collectingCountry>
: 
<collectingRegion id="49D3F83BE70DFFFCFEFCF94EFDA8FBF1" box="[316,535,1749,1774]" country="Germany" name="Schleswig-Holstein" pageId="3" pageNumber="146">Schleswig-Holstein</collectingRegion>
(
<bibRefCitation id="EF864B28E70DFFFCFDEBF94EFB07FA0D" author="Tischler" pageId="3" pageNumber="146" refString="Tischler, W. (1948) Biozonotische Untersuchungen an Wallhecken (Ein Beitrag zur Agrarokologie). Zoologische Jahrbucher. Abteilung fur Systematik, Okologie und Geographie der Tiere, 77, 283 - 400." type="journal article" year="1948">
Tischler 1948; the specimen was identified by 
<collectorName id="26E2530FE70DFFFCFB9FF94DFB6CFBF1" box="[1119,1235,1749,1774]" pageId="3" pageNumber="146">W. Emeis</collectorName>
, and is probably currently present in 
<collectorName id="26E2530FE70DFFFCFEBCF961FDD7FA0D" box="[380,616,1785,1810]" pageId="3" pageNumber="146">W. Emeis Collection</collectorName>
in Flensburg, Germany, but could not be examined
</bibRefCitation>
)
</materialsCitation>
. 
<materialsCitation id="3B7F3C84E70DFFFCFB0CF962FE79FA2A" ID-GBIF-Occurrence="1437764345" collectorName="Kalocsa &amp; Pester Komitat" country="Hungary" location="HUNGARY" pageId="3" pageNumber="146" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F3007649E70DFFFCFB0CF962FAE6FA0D" box="[1228,1369,1785,1810]" name="Hungary" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFB0CF962FAE6FA0D" bold="true" box="[1228,1369,1785,1810]" pageId="3" pageNumber="146">HUNGARY</emphasis>
</collectingCountry>
: 
<specimenCount id="9D11FD50E70DFFFCFAA9F962FA28FA0E" box="[1385,1431,1785,1809]" pageId="3" pageNumber="146" type="male">1 ♂</specimenCount>
, 
<collectorName id="26E2530FE70DFFFCFF57F886FF4EFA2A" box="[151,241,1821,1845]" pageId="3" pageNumber="146">Kalocsa</collectorName>
(
<collectorName id="26E2530FE70DFFFCFEC7F886FE01FA2A" box="[263,446,1821,1845]" pageId="3" pageNumber="146">Pester Komitat</collectorName>
)
</materialsCitation>
; 1 ♀, Gyón (
<bibRefCitation id="EF864B28E70DFFFCFDB6F885FCB3FA29" author="Knezy" box="[630,780,1822,1846]" pageId="3" pageNumber="146" refString="Knezy, G. (1929) Legyeszeti tanulmanyok. Folia Societatis Entomologicae Hungaricae, II (1), 1 - 20." type="journal article" year="1929">Knézy 1929</bibRefCitation>
). 
<materialsCitation id="3B7F3C84E70DFFFCFCE8F886FDA3FA61" ID-GBIF-Occurrence="1437764346" collectingDate="2007-03-25" collectionCode="ZMUM" collectorName="N. Vikhrev" country="Turkey" location="Lake Titreyengol" municipality="Manavgat" pageId="3" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Antalya Province">
<collectingCountry id="F3007649E70DFFFCFCE8F886FC22FA29" box="[808,925,1821,1846]" name="Turkey" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFCE8F886FC22FA29" bold="true" box="[808,925,1821,1846]" pageId="3" pageNumber="146">TURKEY</emphasis>
</collectingCountry>
: 
<specimenCount id="9D11FD50E70DFFFCFC74F885FC57FA2A" box="[948,1000,1822,1846]" pageId="3" pageNumber="146" type="male">1 ♂</specimenCount>
, 
<collectingRegion id="49D3F83BE70DFFFCFC3CF886FB74FA29" box="[1020,1227,1821,1846]" country="Turkey" name="Antalya" pageId="3" pageNumber="146">Antalya Province</collectingRegion>
, 
<collectingMunicipality id="6BCCACA3E70DFFFCFB1EF885FAEEFA29" box="[1246,1361,1822,1846]" pageId="3" pageNumber="146">Manavgat</collectingMunicipality>
, 
<location id="8EC86002E70DFFFCFAA4F886FEA3FA45" LSID="urn:lsid:plazi:treatment:03BE87CFE70FFFFAFF57FDBBFE9DFFB8:8EC86002E70DFFFCFAA4F886FEA3FA45" country="Turkey" municipality="Manavgat" name="Lake Titreyengol" pageId="3" pageNumber="146" stateProvince="Antalya Province">Lake Titreyengöl</location>
, 36.757′N 31.455′E, 
<collectingDate id="EFEDE9F1E70DFFFCFDD2F8DAFD31FA45" box="[530,654,1857,1882]" pageId="3" pageNumber="146" value="2007-03-25">
<date id="FFA91019E70DFFFCFDD2F8DAFD31FA45" box="[530,654,1857,1882]" pageId="3" pageNumber="146" value="2007-03-25">25.iii.2007</date>
</collectingDate>
, 
<collectingMethod id="52564ECEE70DFFFCFD5EF8D9FCABFA46" box="[670,788,1858,1881]" pageId="3" pageNumber="146">sweep net</collectingMethod>
, leg. 
<collectorName id="26E2530FE70DFFFCFC97F8D9FC69FA45" box="[855,982,1857,1882]" pageId="3" pageNumber="146">N. Vikhrev</collectorName>
(
<collectionCode id="ED06AE1CE70DFFFCFC29F8DAFBFBFA46" box="[1001,1092,1857,1881]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="3" pageNumber="146">ZMUM</collectionCode>
) (see http://www.diptera.info/ photogallery.php?photo_id=1374).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30D6552E70DFFFBFF07F812FAD6FCB0" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146" type="discussion">
<paragraph id="8BA836D9E70DFFFBFF07F812FCF8FC95" blockId="3.[151,1437,1245,2030]" lastBlockId="4.[151,1436,151,431]" lastPageId="4" lastPageNumber="147" pageId="3" pageNumber="146">
<emphasis id="B963EACBE70DFFFCFF07F812FE84FABD" bold="true" box="[199,315,1929,1954]" pageId="3" pageNumber="146">Remarks.</emphasis>
From a morphological perspective, 
<taxonomicName id="4C174D5AE70DFFFCFD2AF811FCC7FABE" box="[746,888,1929,1953]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70DFFFCFD2AF811FCC7FABE" box="[746,888,1929,1953]" italics="true" pageId="3" pageNumber="146">D. nimbatus</emphasis>
</taxonomicName>
does not seem related to any of its European congeners, which confirms the conclusions of the previous authors (
<bibRefCitation id="EF864B28E70DFFFCFC6BF835FB85FAD9" author="Parent" box="[939,1082,1966,1990]" pageId="3" pageNumber="146" refString="Parent, O. (1927) Contribution a la distribution geographique de quelques especes de Dolichopodides. Congres des Societes Savantes. Paris et de departements. Comptes rendus, 1926, 449 - 484." type="journal article" year="1927">Parent 1927</bibRefCitation>
; 
<bibRefCitation id="EF864B28E70DFFFCFB88F835FB68FAD9" author="Knezy" box="[1096,1239,1966,1990]" pageId="3" pageNumber="146" refString="Knezy, G. (1929) Legyeszeti tanulmanyok. Folia Societatis Entomologicae Hungaricae, II (1), 1 - 20." type="journal article" year="1929">Knézy 1929</bibRefCitation>
; 
<bibRefCitation id="EF864B28E70DFFFCFB25F836FA2FFAD9" author="Meuffels" box="[1253,1424,1965,1990]" pageId="3" pageNumber="146" refString="Meuffels, H. J. G. (1981) Dolichopus balius et Dolichopus ptenopedilus nouveaux Dolichopodidae palearctiques (Diptera). Bulletin des Recherches Agronomiques de Gembloux, 16 (4), 327 - 334." type="journal article" year="1981">Meuffels 1981</bibRefCitation>
). Species like 
<taxonomicName id="4C174D5AE70DFFFCFEE4F849FE5CFAF5" authority="Fallen" authorityName="Fallen" box="[292,483,2001,2026]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="nitidus">
<emphasis id="B963EACBE70DFFFCFEE4F849FE2CFAF6" box="[292,403,2001,2025]" italics="true" pageId="3" pageNumber="146">D. nitidus</emphasis>
Fallén
</taxonomicName>
, 
<taxonomicName id="4C174D5AE70DFFFCFE2EF849FCBFFAF5" authority="Stannius" authorityName="Stannius" box="[494,768,2001,2026]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="griseipennis">
<emphasis id="B963EACBE70DFFFCFE2EF849FD26FAF6" box="[494,665,2002,2025]" italics="true" pageId="3" pageNumber="146">D. griseipennis</emphasis>
Stannius
</taxonomicName>
and 
<taxonomicName id="4C174D5AE70DFFFCFCF7F849FBA5FAF5" authority="Haliday" authorityName="Haliday" box="[823,1050,2001,2026]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="146" phylum="Arthropoda" rank="species" species="diadema">
<emphasis id="B963EACBE70DFFFCFCF7F849FC05FAF6" box="[823,954,2001,2025]" italics="true" pageId="3" pageNumber="146">D. diadema</emphasis>
Haliday
</taxonomicName>
also feature an oblique M1+2 bend, however, with only the posterior hanging vein. Species of the 
<taxonomicName id="4C174D5AE70AFFFBFC94FF03FB9DFDB0" box="[852,1058,152,176]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="ziczac">
<emphasis id="B963EACBE70AFFFBFC94FF03FB9DFDB0" box="[852,1058,152,176]" italics="true" pageId="4" pageNumber="147">Dolichopus ziczac</emphasis>
</taxonomicName>
species group have both hanging veins, but differ from 
<taxonomicName id="4C174D5AE70AFFFBFE5EFF26FD93FDCB" box="[414,556,188,212]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70AFFFBFE5EFF26FD93FDCB" box="[414,556,188,212]" italics="true" pageId="4" pageNumber="147">D. nimbatus</emphasis>
</taxonomicName>
in many other ways; e.g., different hypopygial structure (
<bibRefCitation id="EF864B28E70AFFFBFB19FF27FA28FDCB" author="Grichanov" box="[1241,1431,188,213]" pageId="4" pageNumber="147" refString="Grichanov, I. Y. (1998) Afrotropical species of the genus Lichtwardtia Enderlein (Diptera: Dolichopodidae). International Journal of Dipterological Research, 9 (3), 221 - 236." type="journal article" year="1998">Grichanov 1998</bibRefCitation>
; 
<bibRefCitation id="EF864B28E70AFFFBFF57FF78FED5FDE4" author="Zhang" box="[151,362,227,251]" pageId="4" pageNumber="147" refString="Zhang, L., Masunaga, K. &amp; Yang, D. (2009) Species of Lichtwardtia from China (Diptera: Dolichopodidae). Transactions of the American Entomological Society, 135 (1 + 2), 197 - 203." type="journal article" year="2009">
<emphasis id="B963EACBE70AFFFBFF57FF78FEA1FDE4" box="[151,286,227,251]" italics="true" pageId="4" pageNumber="147">Zhang et al</emphasis>
. 2009
</bibRefCitation>
; 
<bibRefCitation id="EF864B28E70AFFFBFEB9FF79FD3CFDE4" author="Selivanova" box="[377,643,226,251]" pageId="4" pageNumber="147" refString="Selivanova, O. V., Negrobov, O. P. &amp; Yang, D. (2010) Redescription of the holotype of Lichtwardtia formosana (Diptera: Dolichopodidae), with new data on its synonymy. Zoosystematica Rossica, 19 (1), 143 - 146." type="journal article" year="2010">
Selivanova 
<emphasis id="B963EACBE70AFFFBFDC0FF78FD8AFDE4" box="[512,565,227,251]" italics="true" pageId="4" pageNumber="147">et al</emphasis>
. 2010
</bibRefCitation>
). The short subcircular 1st flagellomere is not shared with any other European 
<taxonomicName id="4C174D5AE70AFFFBFECAFE9CFE34FC00" box="[266,395,263,287]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B963EACBE70AFFFBFECAFE9CFE34FC00" box="[266,395,263,287]" italics="true" pageId="4" pageNumber="147">Dolichopus</emphasis>
</taxonomicName>
species, which also holds true for the ventral pilosity of tarsus I. A strongly supported clade of 
<taxonomicName id="4C174D5AE70AFFFBFF76FEB0FE88FC5C" box="[182,311,299,323]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="genus">
<emphasis id="B963EACBE70AFFFBFF76FEB0FE88FC5C" box="[182,311,299,323]" italics="true" pageId="4" pageNumber="147">Dolichopus</emphasis>
</taxonomicName>
species does show a short erect ventral pubescence on the fore tarsus (Bernasconi 
<emphasis id="B963EACBE70AFFFBFB26FEB0FAA6FC5C" box="[1254,1305,299,323]" italics="true" pageId="4" pageNumber="147">et al</emphasis>
. 2006), but this is most likely not homologous with the pilosity in 
<taxonomicName id="4C174D5AE70AFFFBFCD7FECBFC1CFC78" box="[791,931,335,359]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70AFFFBFCD7FECBFC1CFC78" box="[791,931,335,359]" italics="true" pageId="4" pageNumber="147">D. nimbatus</emphasis>
</taxonomicName>
. Moreover, all species of this clade share a dorsal bristle on metatarus II, which is absent in 
<taxonomicName id="4C174D5AE70AFFFBFD77FEE8FCFFFC94" box="[695,832,371,395]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70AFFFBFD77FEE8FCFFFC94" box="[695,832,371,395]" italics="true" pageId="4" pageNumber="147">D. nimbatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BA836D9E70AFFFBFF07FE0DFAD6FCB0" blockId="4.[151,1436,151,431]" box="[199,1385,406,431]" pageId="4" pageNumber="147">
<bibRefCitation id="EF864B28E70AFFFBFF07FE0DFEC2FCB0" author="Meuffels" box="[199,381,406,431]" pageId="4" pageNumber="147" refString="Meuffels, H. J. G. (1981) Dolichopus balius et Dolichopus ptenopedilus nouveaux Dolichopodidae palearctiques (Diptera). Bulletin des Recherches Agronomiques de Gembloux, 16 (4), 327 - 334." type="journal article" year="1981">Meuffels (1981)</bibRefCitation>
mentions another three males of 
<taxonomicName id="4C174D5AE70AFFFBFD32FE03FCE7FCB0" box="[754,856,407,431]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70AFFFBFD32FE03FCE7FCB0" box="[754,856,407,431]" italics="true" pageId="4" pageNumber="147">D. balius</emphasis>
</taxonomicName>
from the same site collected on 
<date id="FFA91019E70AFFFBFB04FE0CFADAFCB0" box="[1220,1381,406,431]" pageId="4" pageNumber="147" value="1980-04-20">20 April 1980</date>
.
</paragraph>
</subSubSection>
<caption id="DF686651E70AFFFBFF57FA26FCA3F8CC" box="[151,796,1469,1491]" httpUri="https://zenodo.org/record/345074/files/figure.png" pageId="4" pageNumber="147" targetBox="[151,1436,484,1448]" targetPageId="4">
<paragraph id="8BA836D9E70AFFFBFF57FA26FCA3F8CC" blockId="4.[151,796,1469,1491]" box="[151,796,1469,1491]" pageId="4" pageNumber="147">
<emphasis id="B963EACBE70AFFFBFF57FA26FEADF8CD" bold="true" box="[151,274,1469,1491]" pageId="4" pageNumber="147">FIGURE 3.</emphasis>
<taxonomicName id="4C174D5AE70AFFFBFED8FA26FD89F8CC" authority="Meuffels" authorityName="Meuffels" box="[280,566,1469,1491]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70AFFFBFED8FA26FE32F8CC" box="[280,397,1469,1491]" italics="true" pageId="4" pageNumber="147">Dolichopus</emphasis>
<emphasis id="B963EACBE70AFFFBFE54FA26FE6DF8CC" box="[404,466,1469,1491]" italics="true" pageId="4" pageNumber="147">balius</emphasis>
Meuffels
</taxonomicName>
, male paratype (head).
</paragraph>
</caption>
<subSubSection id="C30D6552E70AFFFAFF07FA61FE9DFFB8" lastPageId="5" lastPageNumber="148" pageId="4" pageNumber="147" type="distribution">
<paragraph id="8BA836D9E70AFFFBFF07FA61FD81FBBC" blockId="4.[151,1436,1530,2023]" pageId="4" pageNumber="147">
<emphasis id="B963EACBE70AFFFBFF07FA61FEDFFB0C" bold="true" box="[199,352,1530,1555]" pageId="4" pageNumber="147">Distribution.</emphasis>
The website Fauna Europaea (
<bibRefCitation id="EF864B28E70AFFFBFD03FA61FCF7FB0C" author="Pollet" box="[707,840,1530,1555]" pageId="4" pageNumber="147" refString="Pollet, M. (2011) Fauna Europaea: Dolichopodidae. In: Pape, T. &amp; Beuk, P. (Eds.), Fauna Europaea: Diptera, Brachycera. Fauna Europaea version 2.4. Available from: http: // www. faunaeur. org (accessed 10 November 2014)" type="journal article" year="2011">Pollet 2011</bibRefCitation>
) currently only displays the presence of 
<taxonomicName id="4C174D5AE70AFFFBFADBFA60FF41FB29" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70AFFFBFADBFA60FF41FB29" italics="true" pageId="4" pageNumber="147">Dolichopus nimbatus</emphasis>
</taxonomicName>
in mainland 
<collectingCountry id="F3007649E70AFFFBFE4EF984FE5EFB28" box="[398,481,1567,1591]" name="Greece" pageId="4" pageNumber="147">Greece</collectingCountry>
. The following countries can be added now: 
<collectingCountry id="F3007649E70AFFFBFC13F985FB89FB28" box="[979,1078,1566,1591]" name="Belgium" pageId="4" pageNumber="147">Belgium</collectingCountry>
, 
<collectingCountry id="F3007649E70AFFFBFB81F984FB2FFB29" box="[1089,1168,1567,1590]" name="France" pageId="4" pageNumber="147">France</collectingCountry>
, 
<collectingCountry id="F3007649E70AFFFBFB5AF984FABDFB28" box="[1178,1282,1567,1591]" name="Germany" pageId="4" pageNumber="147">Germany</collectingCountry>
(?), 
<collectingCountry id="F3007649E70AFFFBFAF5F984FA27FB28" box="[1333,1432,1567,1591]" name="Hungary" pageId="4" pageNumber="147">Hungary</collectingCountry>
, 
<collectingCountry id="F3007649E70AFFFBFF57F9D9FF47FB44" box="[151,248,1602,1627]" name="Bulgaria" pageId="4" pageNumber="147">Bulgaria</collectingCountry>
and 
<collectingCountry id="F3007649E70AFFFBFEEEF9D9FE3FFB44" box="[302,384,1602,1627]" name="Turkey" pageId="4" pageNumber="147">Turkey</collectingCountry>
. Although these records suggest a primarily southeastern European distribution, I assume that the species occurs all over Europe (excl. Fennoscandia), but that other factors have prevented a more extensive and representative recording (see further).
</paragraph>
<paragraph id="8BA836D9E70AFFFAFF07F935FEF7FC20" blockId="4.[151,1436,1530,2023]" lastBlockId="5.[151,1436,151,680]" lastPageId="5" lastPageNumber="148" pageId="4" pageNumber="147">
<emphasis id="B963EACBE70AFFFBFF07F935FE94FBD8" bold="true" box="[199,299,1710,1735]" pageId="4" pageNumber="147">Ecology.</emphasis>
In 
<collectingCountry id="F3007649E70AFFFBFE91F935FE0BFBD8" box="[337,436,1710,1735]" name="Belgium" pageId="4" pageNumber="147">Belgium</collectingCountry>
, 
<taxonomicName id="4C174D5AE70AFFFBFE00F934FDF6FBD9" box="[448,585,1710,1734]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="nimbatus">
<emphasis id="B963EACBE70AFFFBFE00F934FDF6FBD9" box="[448,585,1710,1734]" italics="true" pageId="4" pageNumber="147">D. nimbatus</emphasis>
</taxonomicName>
has been collected in an open swamp within an agricultural area. The Mare de Prussien sampling site in 
<collectingCountry id="F3007649E70AFFFBFE2AF948FD89FBF5" box="[490,566,1747,1770]" name="France" pageId="4" pageNumber="147">France</collectingCountry>
comprises wetlands bordering a lake with vegetation composed of 
<taxonomicName id="4C174D5AE70AFFFBFA8EF948FF5AFA10" class="Liliopsida" family="Juncaceae" genus="Juncus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="147" phylum="Tracheophyta" rank="species" species="effusus">
<emphasis id="B963EACBE70AFFFBFA8EF948FF5AFA10" italics="true" pageId="4" pageNumber="147">Juncus effusus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C174D5AE70AFFFBFF33F96DFE62FA11" box="[243,477,1782,1806]" class="Liliopsida" family="Poaceae" genus="Phragmites" kingdom="Plantae" order="Poales" pageId="4" pageNumber="147" phylum="Tracheophyta" rank="species" species="australis">
<emphasis id="B963EACBE70AFFFBFF33F96DFE62FA11" box="[243,477,1782,1806]" italics="true" pageId="4" pageNumber="147">Phragmites australis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C174D5AE70AFFFBFE2BF96DFD4EFA11" box="[491,753,1782,1806]" class="Liliopsida" family="Poaceae" genus="Deschampsia" kingdom="Plantae" order="Poales" pageId="4" pageNumber="147" phylum="Tracheophyta" rank="species" species="cespitosa">
<emphasis id="B963EACBE70AFFFBFE2BF96DFD4EFA11" box="[491,753,1782,1806]" italics="true" pageId="4" pageNumber="147">Deschampsia cespitosa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C174D5AE70AFFFBFD3FF96CFC04FA11" box="[767,955,1782,1806]" class="Magnoliopsida" family="Asteraceae" genus="Cirsium" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="147" phylum="Tracheophyta" rank="species" species="palustre">
<emphasis id="B963EACBE70AFFFBFD3FF96CFC04FA11" box="[767,955,1782,1806]" italics="true" pageId="4" pageNumber="147">Cirsium palustre</emphasis>
</taxonomicName>
, and 
<emphasis id="B963EACBE70AFFFBFC38F96CFB43FA11" box="[1016,1276,1782,1806]" italics="true" pageId="4" pageNumber="147">Eupatoria cannabinum</emphasis>
. This site was bordered by 
<taxonomicName id="4C174D5AE70AFFFBFEEEF880FED9FA2C" box="[302,358,1819,1843]" class="Ascidiacea" family="Polycitoridae" genus="Salix" kingdom="Animalia" order="Enterogona" pageId="4" pageNumber="147" phylum="Chordata" rank="genus">
<emphasis id="B963EACBE70AFFFBFEEEF880FED9FA2C" box="[302,358,1819,1843]" italics="true" pageId="4" pageNumber="147">Salix</emphasis>
</taxonomicName>
sp. and 
<taxonomicName id="4C174D5AE70AFFFBFE12F880FDADFA2C" box="[466,530,1819,1843]" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="147" phylum="Tracheophyta" rank="genus">
<emphasis id="B963EACBE70AFFFBFE12F880FDADFA2C" box="[466,530,1819,1843]" italics="true" pageId="4" pageNumber="147">Alnus</emphasis>
</taxonomicName>
sp. along the edge of a 
<emphasis id="B963EACBE70AFFFBFCFEF880FBCDFA2C" box="[830,1138,1819,1843]" italics="true" pageId="4" pageNumber="147">
<taxonomicName id="4C174D5AE70AFFFBFCFEF880FC23FA2D" box="[830,924,1819,1842]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="147" phylum="Tracheophyta" rank="genus">Quercus</taxonomicName>
– Betula pendula
</emphasis>
forest, and flooded from November to May (Cédrik Sibold, pers. comm.). Also in La Capelle (
<typeStatus id="54AC887BE70AFFFBFC78F8A4FC55FA48" box="[952,1002,1855,1879]" pageId="4" pageNumber="147">type</typeStatus>
locality of 
<taxonomicName id="4C174D5AE70AFFFBFBB1F8A4FB66FA49" box="[1137,1241,1854,1878]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="147" phylum="Arthropoda" rank="species" species="balius">
<emphasis id="B963EACBE70AFFFBFBB1F8A4FB66FA49" box="[1137,1241,1854,1878]" italics="true" pageId="4" pageNumber="147">D. balius</emphasis>
</taxonomicName>
), the habitat is a natural shallow lake that dries up in summer (
<bibRefCitation id="EF864B28E70AFFFBFD63F8F9FCF1FA64" author="Meuffels" box="[675,846,1890,1915]" pageId="4" pageNumber="147" refString="Meuffels, H. J. G. (1981) Dolichopus balius et Dolichopus ptenopedilus nouveaux Dolichopodidae palearctiques (Diptera). Bulletin des Recherches Agronomiques de Gembloux, 16 (4), 327 - 334." type="journal article" year="1981">Meuffels 1981</bibRefCitation>
). In 
<collectingCountry id="F3007649E70AFFFBFC43F8F8FC6DFA64" box="[899,978,1891,1915]" name="Greece" pageId="4" pageNumber="147">Greece</collectingCountry>
(Kerkini), it was collected in an area of rough grassland alongside the marshes of an old river bed (Gordon Ramel, pers. comm.). The Malaise trap that produced the sample was immediately adjacent to reeds and a willow tree, about 
<quantity id="4CEF9B3CE70AFFFBFBFDF830FBD5FADD" box="[1085,1130,1963,1987]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="147" unit="m" value="2.0">2 m</quantity>
from the water surface. In 
<collectingCountry id="F3007649E70AFFFBFF57F855FF57FAF8" box="[151,232,1998,2023]" name="Turkey" pageId="4" pageNumber="147">Turkey</collectingCountry>
, the species was swept (possibly several specimens) from grassy vegetation in a marshy site near a lake (Titreyengöl), with pastures extensively grazed by horses, and with brashes (ruderal sites with broken rocks and rubbish), wetland, and temporary pools (in March), surrounded by suburban area and pine forests on sandy dunes (Nikita Vikhrev, pers. comm.). In summary, the habitat of the species might be described as open grassland or marshland mostly bordering large stagnant water bodies, often flooded during winter and spring, and drying out during summer.
</paragraph>
<paragraph id="8BA836D9E70BFFFAFF07FED6FE9DFFB8" blockId="5.[151,1436,151,680]" pageId="5" pageNumber="148">
It is unclear why this seemingly widespread species has not been collected more often. One explanation might be that it is extremely stenotopic and entirely confined to its habitat which implies that this specific habitat must be investigated to encounter the species. Swamps that are flooded most of the year are not the most appealing collecting sites for dipterists. Another factor that most certainly contributes to its scarcity in collections is its seasonal activity that differs considerably from that of any other 
<taxonomicName id="4C174D5AE70BFFFAFCA9FE47FC55FCEB" box="[873,1002,476,500]" class="Insecta" family="Dolichopodidae" genus="Dolichopus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B963EACBE70BFFFAFCA9FE47FC55FCEB" box="[873,1002,476,500]" italics="true" pageId="5" pageNumber="148">Dolichopus</emphasis>
</taxonomicName>
species in 
<collectingRegion id="49D3F83BE70BFFFAFBA5FE46FB7FFCEB" box="[1125,1216,477,500]" country="American Samoa" name="Western" pageId="5" pageNumber="148">Western</collectingRegion>
Europe. Not taking climatological effects at different latitudes into account, collection dates vary from March-April over July to October, with most specimens encountered in (very) early spring. As most dolichopodid workers conduct collecting mostly from late spring to late summer (May-August), activity peaks of this species might thus be missed. For that reason, it might be quite rewarding to investigate grassy habitats along lakes in early spring and late autumn.
</paragraph>
</subSubSection>
</treatment>
</document>