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<document id="0F41F0E71A623128D78D5DCB7FF472B7" ID-DOI="10.11646/zootaxa.1161.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5058330" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625190197324" checkinUser="felipe" docAuthor="Hippa, Heikki" docDate="2006" docId="5D4C87852855FFBE5C1A0C4FFD24F8E1" docLanguage="en" docName="zt01161p049.pdf" docOrigin="Zootaxa 1161 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1161.1.1" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Manota angustata Hippa 2006, sp. n." docType="treatment" docVersion="4" lastPageNumber="18" masterDocId="A175FFFD2844FFAF5D120E76FFC5FFE7" masterDocTitle="Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species" masterLastPageNumber="49" masterPageNumber="1" pageNumber="18" updateTime="1698983105622" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="BDFF4569059F62BD334D7F0D35DB6A33">Diversity of Manota Williston (Diptera: Mycetophilidae) in a Malaysian rainforest: description of twenty-seven new sympatric species</mods:title>
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<mods:namePart id="732F58DDE413DA3049555EE039BFDEAF">Hippa, Heikki</mods:namePart>
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<mods:date id="95B7A8453C1E0D2F7FAE26DBEF5FC13E">2006</mods:date>
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<mods:number id="0A67C2798A44602DE85A188248577E1D">2006-03-30</mods:number>
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<mods:number id="9817D3DBF9B35496CAA601A5517B230C">1161</mods:number>
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<mods:identifier id="D6514579C4D3F17D87EC3F2F3FA37305" type="DOI">10.11646/zootaxa.1161.1.1</mods:identifier>
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<treatment id="5D4C87852855FFBE5C1A0C4FFD24F8E1" ID-DOI="http://doi.org/10.5281/zenodo.5059292" ID-Zenodo-Dep="5059292" LSID="urn:lsid:plazi:treatment:5D4C87852855FFBE5C1A0C4FFD24F8E1" httpUri="http://treatment.plazi.org/id/5D4C87852855FFBE5C1A0C4FFD24F8E1" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="9DFF65182855FFBE5C1A0C4FFDE8FDB4" box="[264,557,569,595]" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="D55A36932855FFBE5C1A0C4FFDE8FDB4" blockId="17.[264,557,569,630]" box="[264,557,569,595]" pageId="17" pageNumber="18">
<heading id="8E1281FF2855FFBE5C1A0C4FFDE8FDB4" bold="true" box="[264,557,569,595]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<emphasis id="E791EA812855FFBE5C1A0C4FFDE8FDB4" bold="true" box="[264,557,569,595]" pageId="17" pageNumber="18">
<taxonomicName id="12E54D102855FFBE5C1A0C4FFE26FDB4" ID-CoL="6R6JX" authority="Hippa, 2006" authorityName="Hippa" authorityYear="2006" box="[264,483,569,595]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="angustata" status="sp. nov.">
<emphasis id="E791EA812855FFBE5C1A0C4FFE26FDB4" bold="true" box="[264,483,569,595]" italics="true" pageId="17" pageNumber="18">Manota angustata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FCA257FA2855FFBE5CF90C4FFDE8FDB4" box="[491,557,569,595]" pageId="17" pageNumber="18" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9DFF65182855FFBE5C1A0C2AFD24F8E1" pageId="17" pageNumber="18" type="description">
<paragraph id="D55A36932855FFBE5C1A0C2AFE2AFD91" blockId="17.[264,557,569,630]" box="[264,495,604,630]" pageId="17" pageNumber="18">
(
<figureCitation id="4DDE2A162855FFBE5C030C2AFEB3FD91" box="[273,374,604,630]" captionStart="FIGURE 1" captionStartId="4.[264,368,1691,1715]" captionTargetBox="[300,1268,323,1675]" captionTargetId="figure-116@4.[285,1310,298,1688]" captionTargetPageId="4" captionText="FIGURE 1. Left palp, dorsal view (A); apical part of palpomere 3, lateral view (B, C, D); and apical part of tibia 1 with basal part of tarsomere 1, prolateral view (E). A, C. Manota heptacantha sp. n (paratypes). B. Manota pappi sp. n. (paratype). D. Manota spadix sp. n. (paratype). E. Manota angustata sp. n. (paratype). Scale 0.1 mm. 1 = palpomere 1, 2 = palpomere 2, 3 = palpomere 3, 4 = palpomere 4, 5 = palpomere 5, 6 = sensory pit, 7 = curved sensillum, 8 = bluntended sensillum, 9 = seta, 10 = parasegment." figureDoi="http://doi.org/10.5281/zenodo.5058332" httpUri="https://zenodo.org/record/5058332/files/figure.png" pageId="17" pageNumber="18">Figs. 1E</figureCitation>
;
<figureCitation id="4DDE2A162855FFBE5C910C2AFE2DFD91" box="[387,488,604,630]" captionStart="FIGURE 5" captionStartId="16.[264,367,1792,1816]" captionTargetBox="[322,1296,318,1750]" captionTargetId="figure-73@16.[288,1296,311,1771]" captionTargetPageId="16" captionText="FIGURE 5. Antennal flagellomere 4, lateral view (A, D); hypopygium, ventral (B, E) and dorsal view (C, F). A, B, C. Manota ovata sp. n. (A, C paratype; B holotype). D, E, F. Manota angustata sp. n. (D, E holotype; F paratype). Scale for A and D 0.05; for B, C, E, and F 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058344" httpUri="https://zenodo.org/record/5058344/files/figure.png" pageId="17" pageNumber="18">5D, E, F</figureCitation>
)
</paragraph>
<paragraph id="D55A36932855FFBE5C1A0CD9FDD6FD2E" blockId="17.[264,1323,687,1393]" box="[264,531,687,713]" pageId="17" pageNumber="18">
A small­sized
<taxonomicName id="12E54D102855FFBE5CA30CD9FDC9FD2F" authorityName="Williston" authorityYear="1896" box="[433,524,687,712]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="E791EA812855FFBE5CA30CD9FDC9FD2F" box="[433,524,687,712]" italics="true" pageId="17" pageNumber="18">Manota</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D55A36932855FFBE5C2A0CA1FDC0FAAE" blockId="17.[264,1323,687,1393]" pageId="17" pageNumber="18">
Male.
<emphasis id="E791EA812855FFBE5C910CA0FE00FD17" bold="true" box="[387,453,726,752]" pageId="17" pageNumber="18">Head</emphasis>
. Flagellomere 4,
<figureCitation id="4DDE2A162855FFBE5F810CA1FD35FD16" box="[659,752,727,753]" captionStart="FIGURE 5" captionStartId="16.[264,367,1792,1816]" captionTargetBox="[322,1296,318,1750]" captionTargetId="figure-73@16.[288,1296,311,1771]" captionTargetPageId="16" captionText="FIGURE 5. Antennal flagellomere 4, lateral view (A, D); hypopygium, ventral (B, E) and dorsal view (C, F). A, B, C. Manota ovata sp. n. (A, C paratype; B holotype). D, E, F. Manota angustata sp. n. (D, E holotype; F paratype). Scale for A and D 0.05; for B, C, E, and F 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058344" httpUri="https://zenodo.org/record/5058344/files/figure.png" pageId="17" pageNumber="18">Fig. 5D</figureCitation>
. Maxillary palpus with palpomere 3 bearing 4–5 apically expanded curved sensilla: palpomere 4 with parasegment.
<emphasis id="E791EA812855FFBE59090C88FBBBFCFF" bold="true" box="[1051,1150,766,792]" pageId="17" pageNumber="18">Thorax.</emphasis>
Anepisternum setose. Anterior basalare non­setose. Preepisternum 2 setose. Laterotergite setose. Episternum 3 setose. Front leg,
<figureCitation id="4DDE2A162855FFBE5F840D39FD30FC8E" box="[662,757,847,873]" captionStart="FIGURE 1" captionStartId="4.[264,368,1691,1715]" captionTargetBox="[300,1268,323,1675]" captionTargetId="figure-116@4.[285,1310,298,1688]" captionTargetPageId="4" captionText="FIGURE 1. Left palp, dorsal view (A); apical part of palpomere 3, lateral view (B, C, D); and apical part of tibia 1 with basal part of tarsomere 1, prolateral view (E). A, C. Manota heptacantha sp. n (paratypes). B. Manota pappi sp. n. (paratype). D. Manota spadix sp. n. (paratype). E. Manota angustata sp. n. (paratype). Scale 0.1 mm. 1 = palpomere 1, 2 = palpomere 2, 3 = palpomere 3, 4 = palpomere 4, 5 = palpomere 5, 6 = sensory pit, 7 = curved sensillum, 8 = bluntended sensillum, 9 = seta, 10 = parasegment." figureDoi="http://doi.org/10.5281/zenodo.5058332" httpUri="https://zenodo.org/record/5058332/files/figure.png" pageId="17" pageNumber="18">Fig. 1E</figureCitation>
.
<emphasis id="E791EA812855FFBE5E140D38FC8CFC8F" bold="true" box="[774,841,846,872]" pageId="17" pageNumber="18">Wing</emphasis>
. Length
<quantity id="121D9B762855FFBE5EAC0D39FB96FC8E" box="[958,1107,847,873]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="17" pageNumber="18" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.3–1.5 mm</quantity>
. Wing membrane dorsally with few setae at posterior margin. Sc distally of h non­setose. Setae indicating A1 not observed.
<emphasis id="E791EA812855FFBE5CA20DE8FD94FC5F" bold="true" box="[432,593,926,952]" pageId="17" pageNumber="18">Hypopygium</emphasis>
(
<figureCitation id="4DDE2A162855FFBE5F710DE9FD2EFC5E" box="[611,747,927,953]" captionStart="FIGURE 5" captionStartId="16.[264,367,1792,1816]" captionTargetBox="[322,1296,318,1750]" captionTargetId="figure-73@16.[288,1296,311,1771]" captionTargetPageId="16" captionText="FIGURE 5. Antennal flagellomere 4, lateral view (A, D); hypopygium, ventral (B, E) and dorsal view (C, F). A, B, C. Manota ovata sp. n. (A, C paratype; B holotype). D, E, F. Manota angustata sp. n. (D, E holotype; F paratype). Scale for A and D 0.05; for B, C, E, and F 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058344" httpUri="https://zenodo.org/record/5058344/files/figure.png" pageId="17" pageNumber="18">Figs. 5E, F</figureCitation>
). Sternite 9 large, laterally sharply delimited but posterior margin weakly indicated, setae subequal to ventral setae of gonocoxa. Dorsomesial margin of gonocoxa simple, without apicolateral triangular lobe but with small subapical mesial rounded lobe covered by mesial margin, with 1 long sigmoid megaseta and 1 shorter less sigmoid one in position III, both arising from long common basal body; in
<specimenCount id="C3E3FD1A2855FFBE5CA10A11FDF5FB66" box="[435,560,1127,1153]" pageId="17" pageNumber="18" type="generic" typeStatus="paratype">1 paratype</specimenCount>
, the difference between these 2 setae is not as great as in
<figureCitation id="4DDE2A162855FFBE59DF0A11FAE0FB66" box="[1229,1317,1127,1153]" captionStart="FIGURE 5" captionStartId="16.[264,367,1792,1816]" captionTargetBox="[322,1296,318,1750]" captionTargetId="figure-73@16.[288,1296,311,1771]" captionTargetPageId="16" captionText="FIGURE 5. Antennal flagellomere 4, lateral view (A, D); hypopygium, ventral (B, E) and dorsal view (C, F). A, B, C. Manota ovata sp. n. (A, C paratype; B holotype). D, E, F. Manota angustata sp. n. (D, E holotype; F paratype). Scale for A and D 0.05; for B, C, E, and F 0.10 mm." figureDoi="http://doi.org/10.5281/zenodo.5058344" httpUri="https://zenodo.org/record/5058344/files/figure.png" pageId="17" pageNumber="18">Fig. 5E</figureCitation>
, but the shorter one, too, is curved in the apical part. Gonostylus elongate, parallel­sided, with subapical process on the mesial side. Parastylar lobe well exposed, directed obliquely anteriad, with 1 apical setae. Tegmen rather short, triangular, with slight lateral shoulders. Setae on sternite 10 about
<quantity id="121D9B762855FFBE5F580B71FDBCFAC6" box="[586,633,1287,1313]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="17" pageNumber="18" unit="in" value="7.0">7 in</quantity>
number, in scattered row extending anteriorly to level of apical part of tegmen.
</paragraph>
<paragraph id="D55A36932855FFBE5C2A0B21FCDBFA96" blockId="17.[264,1323,687,1393]" box="[312,798,1367,1393]" pageId="17" pageNumber="18">Female and preimaginal stages unknown.</paragraph>
<paragraph id="D55A36932855FFBE5C1A0BDFFE4DFA25" blockId="17.[264,1322,1449,1555]" box="[264,392,1449,1474]" pageId="17" pageNumber="18">
<emphasis id="E791EA812855FFBE5C1A0BDFFE4DFA25" box="[264,392,1449,1474]" italics="true" pageId="17" pageNumber="18">Discussion</emphasis>
</paragraph>
<paragraph id="D55A36932855FFBE5C2A0BA7FCE2F9F4" blockId="17.[264,1322,1449,1555]" pageId="17" pageNumber="18">
<taxonomicName id="12E54D102855FFBE5C2A0BA7FDCAFA0D" authorityName="Hippa" authorityYear="2006" box="[312,527,1489,1514]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="angustata">
<emphasis id="E791EA812855FFBE5C2A0BA7FDCAFA0D" box="[312,527,1489,1514]" italics="true" pageId="17" pageNumber="18">Manota angustata</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="12E54D102855FFBE5FA00BA7FCD8FA0D" authorityName="Hippa" authorityYear="2006" box="[690,797,1489,1514]" class="Insecta" family="Mycetophilidae" genus="Manota" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ovata">
<emphasis id="E791EA812855FFBE5FA00BA7FCD8FA0D" box="[690,797,1489,1514]" italics="true" pageId="17" pageNumber="18">M. ovata</emphasis>
</taxonomicName>
, but is distinguished by its narrow parallelsided gonostylus with a mesial subapical lobe.
</paragraph>
<paragraph id="D55A36932855FFBE5C1A083AFE8FF982" blockId="17.[264,1323,1612,1798]" box="[264,330,1612,1637]" pageId="17" pageNumber="18">
<emphasis id="E791EA812855FFBE5C1A083AFE8FF982" box="[264,330,1612,1637]" italics="true" pageId="17" pageNumber="18">Types</emphasis>
</paragraph>
<paragraph id="D55A36932855FFBE5C2A0802FD24F8E1" blockId="17.[264,1323,1612,1798]" pageId="17" pageNumber="18">
<materialsCitation id="658D3CCE2855FFBE5C2A0802FE7AF939" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Selangor" typeStatus="holotype">
<emphasis id="E791EA812855FFBE5C2A0802FE66F96A" box="[312,419,1652,1677]" italics="true" pageId="17" pageNumber="18">
<typeStatus id="0A5E88312855FFBE5C2A0802FE66F96A" box="[312,419,1652,1677]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingCounty id="3C3B4E1F2855FFBE5CA60802FE36F969" box="[436,499,1652,1678]" pageId="17" pageNumber="18">Male</collectingCounty>
,
<collectingMunicipality id="353EACE92855FFBE5F100802FDF9F969" box="[514,572,1652,1678]" pageId="17" pageNumber="18">West</collectingMunicipality>
<collectingCountry id="ADF276032855FFBE5F570802FD76F969" box="[581,691,1652,1678]" name="Malaysia" pageId="17" pageNumber="18">Malaysia</collectingCountry>
,
<collectingRegion id="1721F8712855FFBE5FD30802FCEEF969" box="[705,811,1652,1678]" country="Malaysia" name="Selangor" pageId="17" pageNumber="18">Selangor</collectingRegion>
,
<location id="D03A60482855FFBE5E280802FC12F969" LSID="urn:lsid:plazi:treatment:5D4C87852855FFBE5C1A0C4FFD24F8E1:D03A60482855FFBE5E280802FC12F969" box="[826,983,1652,1678]" country="Malaysia" county="Male" municipality="West" name="Ulu Gombak" pageId="17" pageNumber="18" stateProvince="Selangor">Ulu Gombak</location>
,
<location id="D03A60482855FFBE5EF70802FE60F951" LSID="urn:lsid:plazi:treatment:5D4C87852855FFBE5C1A0C4FFD24F8E1:D03A60482855FFBE5EF70802FE60F951" country="Malaysia" county="Male" municipality="West" name="University of Malaya Field Study Centre" pageId="17" pageNumber="18" stateProvince="Selangor">University of Malaya Field Study Centre</location>
,
<quantity id="121D9B762855FFBE5CA008EAFE3EF951" box="[434,507,1692,1718]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="17" pageNumber="18" unit="ft" value="800.0">
<elevation id="5EC8D1A02855FFBE5CA008EAFE3EF951" box="[434,507,1692,1718]" metricMagnitude="2" metricUnit="m" metricValue="2.4384" pageId="17" pageNumber="18" unit="ft" value="800.0">800 ft</elevation>
</quantity>
, Malaise in jungle,
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<collectingDate id="B11FE9BB2855FFBE5FF608EAFBD4F951" box="[740,1041,1692,1718]" pageId="17" pageNumber="18" value="1997-02-24" valueMax="1997-03-21" valueMin="1997-02-24">24. Feb.–21. March 1997</collectingDate>
</date>
,
<collectorName id="781053452855FFBE590D08EAFB4BF951" box="[1055,1166,1692,1718]" pageId="17" pageNumber="18">H. Hippa</collectorName>
,
<collectorName id="781053452855FFBE598808EAFAEFF951" box="[1178,1322,1692,1718]" pageId="17" pageNumber="18">M. Jaschhof</collectorName>
and
<collectorName id="781053452855FFBE5C2808B2FE7AF939" box="[314,447,1732,1758]" pageId="17" pageNumber="18">B. Viklund</collectorName>
</materialsCitation>
.
<materialsCitation id="658D3CCE2855FFBE5CD908B2FC31F939" box="[459,1012,1732,1758]" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="17" pageNumber="18" specimenCount="4" specimenCount-male="4" stateProvince="Selangor" typeStatus="paratype">
<typeStatus id="0A5E88312855FFBE5CD908B2FD84F939" box="[459,577,1732,1758]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="C3E3FD1A2855FFBE5F5F08B2FD62F939" box="[589,679,1732,1758]" pageId="17" pageNumber="18" type="male">4 males</specimenCount>
with same data as holotype
</materialsCitation>
;
<materialsCitation id="658D3CCE2855FFBE591208B2FEABF8E1" collectingDate="1997-02-24" collectingDateMax="1997-03-21" collectingDateMin="1997-02-24" collectorName="H. Hippa &amp; M. Jaschhof &amp; B. Viklund" country="Malaysia" county="Male" elevation="244" location="Ulu Gombak" municipality="West" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Selangor" typeStatus="paratype">
<specimenCount id="C3E3FD1A2855FFBE591208B2FB95F939" box="[1024,1104,1732,1758]" pageId="17" pageNumber="18" type="male">1 male</specimenCount>
with same data as holotype
</materialsCitation>
except for 22. Feb.–21. March.
</paragraph>
</subSubSection>
</treatment>
</document>
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