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<document id= "17FE759BC2F640F00CACF6B428D19385" ID-CLB-Dataset= "35553" ID-DOI= "http://dx.doi.org/10.3897/zookeys.269.4252" ID-GBIF-Dataset= "9e397890-e472-47a5-a5db-1bfa65137034" ID-PMC= "PMC3592266" ID-Pensoft-Pub= "1313-2970-269-11" ID-PubMed= "23653523" ModsDocAuthor= "" ModsDocDate= "2013" ModsDocID= "1313-2970-269-11" ModsDocOrigin= "ZooKeys 269" ModsDocTitle= "Review of the Palaearctic Acomopterella Zaitzev (Diptera, Sciaroidea, Mycetophilidae)" checkinTime= "1451247671814" checkinUser= "pensoft" docAuthor= "Kallweit, Uwe" docDate= "2013" docId= "3AB8E195ED5178CECB60438E64AAE916" docLanguage= "en" docName= "ZooKeys 269: 11-32" docOrigin= "ZooKeys 269" docSource= "http://dx.doi.org/10.3897/zookeys.269.4252" docTitle= "Acomopterella Zaitzev 1989" docType= "treatment" docVersion= "5" lastPageNumber= "12" masterDocId= "FF9B955C197EFFF76C122727FF92FF8E" masterDocTitle= "Review of the Palaearctic Acomopterella Zaitzev (Diptera, Sciaroidea, Mycetophilidae)" masterLastPageNumber= "32" masterPageNumber= "11" pageNumber= "12" updateTime= "1732831155614" updateUser= "ExternalLinkService" >
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<mods:title id= "F9F6E552328BDDEDF2B4B2251E845A04" > Review of the Palaearctic Acomopterella Zaitzev (Diptera, Sciaroidea, Mycetophilidae)</mods:title>
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<mods:namePart id= "4F18F17C6C67652CAD777AE5A8D4C4A3" > Kallweit, Uwe</mods:namePart>
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<treatment id= "3AB8E195ED5178CECB60438E64AAE916" ID-GBIF-Taxon= "152040638" LSID= "urn:lsid:plazi:treatment:3AB8E195ED5178CECB60438E64AAE916" httpUri= "http://treatment.plazi.org/id/3AB8E195ED5178CECB60438E64AAE916" lastPageId= "2" lastPageNumber= "12" pageId= "1" pageNumber= "12" scope_family= "Mycetophilidae" scope_order= "Diptera" >
<subSubSection id= "43690763F9AAE893D605DCA1FA5A5FDC" pageId= "1" pageNumber= "12" type= "nomenclature" >
<paragraph id= "68ECB13528854A44D68BB6840976CD62" pageId= "1" pageNumber= "12" >
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Genus
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<taxonomicName id= "9BD7A49E4753206CFD8BAC6F991C7277" ID-CoL= "8ZY82" LSID= "http://species-id.net/wiki/Acomopterella" authority= "Zaitzev, 1989" authorityName= "Zaitzev" authorityYear= "1989" class= "Insecta" family= "Mycetophilidae" genus= "Acomopterella" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Acomopterella" order= "Diptera" pageId= "1" pageNumber= "12" phylum= "Arthropoda" rank= "genus" > Acomopterella Zaitzev, 1989</taxonomicName>
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</paragraph>
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<subSubSection id= "5841F86E746BF8459FEC5684F17561A9" pageId= "1" pageNumber= "12" type= "reference_group" >
<paragraph id= "497988608B479EC5EBC910A65B9CD75C" pageId= "1" pageNumber= "12" >
<taxonomicName id= "DDA8796F57AF8BC8EE4EE8C155643664" class= "Insecta" family= "Mycetophilidae" genus= "Acomopterella" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Acomopterella" order= "Diptera" pageId= "1" pageNumber= "12" phylum= "Arthropoda" rank= "genus" > Acomopterella</taxonomicName>
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Zaitzev, 1989: 134 (description);
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<normalizedToken id= "54CAD75C9E177A501D52725285B5DEA0" originalValue= "Ševčík" > Sevcik</normalizedToken>
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& Chandler: 2008: 63 (discussion).
</paragraph>
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<subSubSection id= "279813958728E14A26CD324FBDF866E3" pageId= "1" pageNumber= "12" type= "type species" >
<paragraph id= "A7F45AB58437A90496FD4CCC0F36078A" pageId= "1" pageNumber= "12" > Type species.</paragraph>
<paragraph id= "5958512718A5C4B709B7408FEC48D2CA" pageId= "1" pageNumber= "12" >
<taxonomicName id= "0F072E1559ACBB83CEF175ECC4AFEF99" class= "Insecta" family= "Mycetophilidae" genus= "Acomopterella" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Acomopterella arnaudi" order= "Diptera" pageId= "1" pageNumber= "12" phylum= "Arthropoda" rank= "species" species= "arnaudi" > Acomopterella arnaudi</taxonomicName>
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Zaitzev, 1989: 134 (by original designation) =
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<taxonomicName id= "3B6024A8431FD153A1E7C3B290C23AC7" class= "Insecta" family= "Mycetophilidae" genus= "Tetragoneura" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Tetragoneura fallax" order= "Diptera" pageId= "1" pageNumber= "12" phylum= "Arthropoda" rank= "species" species= "fallax" > Tetragoneura fallax</taxonomicName>
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Sherman, 1925: 20.
</paragraph>
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<subSubSection id= "72FF9914A6831565AE7751F7ECA2D5EC" pageId= "1" pageNumber= "12" type= "diagnosis" >
<paragraph id= "4195E067531A54D8EAB70E3D16A088E7" pageId= "1" pageNumber= "12" > Diagnosis.</paragraph>
<paragraph id= "2579E8419BE81E5A39B7ADE9961AC2F2" pageId= "1" pageNumber= "12" >
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(based on
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<normalizedToken id= "78FD71AFC70BE594A63F1BB22D824621" originalValue= "Zaitzev’ > Zaitzev's</normalizedToken>
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original genus description, with additions).
</paragraph>
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<paragraph id= "C6E26349A0A343E70C817C0C8FC8484B" pageId= "1" pageNumber= "12" > Body length 4.6-5.9 mm. Head. With 3 ocelli. Mid ocellus slightly in front of the lateral ocelli. Lateral ocelli scarcely to distinctly remote from eye margin. Eyes with short setulae. Face rather flat, bare or setose. Clypeus oval, slightly bulging, setose. Face and clypeus loosely connected. Palpi pentamerous. Antennae 16-segmented, 2-3 times length of the thorax. Length of mesial flagellomeres 3-5 times their own width. Thorax. Scutum with short, close lying acrostichal and long erect dorsocentral bristles. Scutellum with 2 very long medial bristles. Mediotergite, laterotergites and pleura bare. Wing. Transparent. Wing membrane with microtrichia only. C, the apical part of ta, R1, R5, M1, M2, M4 and CuA with macrotrichia. C extending to the tip of R5. R4 present; ta oblique, slightly shorter than stem of fork M1 + M2. Base of M4 on level of base of ta or slightly more proximal. Vein A well developed, its tip behind the level of base of ta or ending slightly before. Between A and the stem of fork M4 + CuA lies the weakly sclerotized vein CuP, traceable as fold line. Legs. Long. Fore coxae with numerous long bristles on its frontal side. Apical part of mid coxae with bristles on its lateral and frontal surface. Hind coxae with a row of very long bristles on the hind lateral surface. Mid tibiae with well developed tibial organ on its basal part. Tarsal claws with teeth. Empodia well developed. Abdomen. Segments 1-7 of normal size. Tergite 8 vestigial; Sternite 8 exceeding half length of sternite 7. Tergite 9 rectangular, with deep distal incision. Gonocoxites with apical processes.</paragraph>
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</subSubSection>
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<subSubSection id= "BFE1578EFEA4879976426F820C84D96C" lastPageId= "2" lastPageNumber= "13" pageId= "1" pageNumber= "12" type= "comments" >
<paragraph id= "8A052BA4CEBD608DE8AE4667B77960DF" pageId= "1" pageNumber= "12" > Comments.</paragraph>
<paragraph id= "66B98FAB150E04DE0D98A3DDD8D974A1" lastPageId= "2" lastPageNumber= "13" pageId= "1" pageNumber= "12" >
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The described genus belongs to the tribe
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<taxonomicName id= "5756DF1907C825B272BA6D2491F23817" lsidName= "" pageId= "1" pageNumber= "12" rank= "tribe" tribe= "Gnoristini" > Gnoristini</taxonomicName>
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in the subfamily
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<taxonomicName id= "79E3D861D1EC0A5FFCB5E30C9FC9EB9E" lsidName= "" pageId= "1" pageNumber= "12" rank= "subFamily" subFamily= "Sciophilinae" > Sciophilinae</taxonomicName>
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. It is similar to
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<taxonomicName id= "5F93C76CB6EFF79B564234B8291DB6A5" class= "Insecta" family= "Mycetophilidae" genus= "Dziedzickia" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Dziedzickia marginata" order= "Diptera" pageId= "1" pageNumber= "12" phylum= "Arthropoda" rank= "species" species= "marginata" > Dziedzickia marginata</taxonomicName>
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(
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<bibRefCitation id= "31EC7FE7BC640FCBD09988B15CE4796A" author= "Dziedzicki, H" journalOrPublisher= "Pamietnik Fizyiograficzny" pageId= "11" pageNumber= "22" pagination= "164 - 194" title= "Przyczynek do fauny owadow dwuskrzydlych. Rodzaje nowe: Hertwigia nov. gen., Eurycera nov. gen. i gatunki rodzajow: Boletina, Sciophila." volume= "5" year= "1885" > Dziedzicki 1885</bibRefCitation>
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), but differing
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<pageBreakToken id= "BBAAC4613E8D195E6A0DB955E63AB338" pageId= "2" pageNumber= "13" start= "start" > by</pageBreakToken>
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bare laterotergites and a much longer vein ta. It differs from the Holarctic genus
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<taxonomicName id= "AA53F74214E4D841EFD541967AC4CA48" class= "Insecta" family= "Mycetophilidae" genus= "Acomoptera" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Acomoptera" order= "Diptera" pageId= "2" pageNumber= "13" phylum= "Arthropoda" rank= "genus" > Acomoptera</taxonomicName>
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Vockeroth, 1980, in the position of Sc (in
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<taxonomicName id= "9D8132942B9CE7627016170708884230" class= "Insecta" family= "Mycetophilidae" genus= "Acomopterella" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Acomopterella" order= "Diptera" pageId= "2" pageNumber= "13" phylum= "Arthropoda" rank= "genus" > Acomopterella</taxonomicName>
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it joins R1, in
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<taxonomicName id= "D8DA239DB70C3F4B71116E4B4837ADE7" class= "Insecta" family= "Mycetophilidae" genus= "Acomoptera" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Acomoptera" order= "Diptera" pageId= "2" pageNumber= "13" phylum= "Arthropoda" rank= "genus" > Acomoptera</taxonomicName>
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C), in the shape of the cell between Rs and R4 (distinctly longer in
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<taxonomicName id= "ADC34754661BD6D3A295A6CAB8EF6A5E" class= "Insecta" family= "Mycetophilidae" genus= "Acomoptera" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Acomoptera" order= "Diptera" pageId= "2" pageNumber= "13" phylum= "Arthropoda" rank= "genus" > Acomoptera</taxonomicName>
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) and in the strikingly longer ta. The genus is also similar to
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<taxonomicName id= "F1C764A238960AD9FE99DABF4A6A6EA7" class= "Insecta" family= "Mycetophilidae" genus= "Austrosynapha" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Austrosynapha" order= "Diptera" pageId= "2" pageNumber= "13" phylum= "Arthropoda" rank= "genus" > Austrosynapha</taxonomicName>
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Tonnoir, 1929, known from South America, New Zealand and Tasmania. Typical in both genera is the absence of setae on the laterotergites and the presence of a relatively long oblique vein ta. In
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<taxonomicName id= "16A9617DDE7CBA208E0DD0A23AAABEBB" class= "Insecta" family= "Mycetophilidae" genus= "Austrosynapha" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "Austrosynapha" order= "Diptera" pageId= "2" pageNumber= "13" phylum= "Arthropoda" rank= "genus" > Austrosynapha</taxonomicName>
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vein R4 is absent.
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