treatments-xml/data/03/D3/87/03D3878DFF9A58191EE26771DE31F942.xml

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<document id="7B2E7F7678B02C0449CE397D3C6BCA27" ID-CLB-Dataset="27716" ID-DOI="10.5852/ejt.2019.510" ID-GBIF-Dataset="08fe4f6b-5422-45af-bc2e-29ad35fed42f" ID-Zenodo-Dep="2616378" ID-ZooBank="BF58E108-2C6C-4A80-BD7D-47FEF5805B06" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1553845906407" checkinUser="plazi" docAuthor="Fujimoto, Shinta &amp; Hansen, Jesper Guldberg" docDate="2019" docId="03D3878DFF9A58191EE26771DE31F942" docLanguage="en" docName="ejt-510_fujimoto_hansen.pdf.imf" docOrigin="European Journal of Taxonomy 510" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Angursa Pollock 1979" docType="key" docVersion="8" lastPageNumber="13" masterDocId="FFEAFFF5FF9658151E5F6411DB46FFE9" masterDocTitle="Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan" masterLastPageNumber="19" masterPageNumber="1" pageNumber="13" updateTime="1698715651624" updateUser="ExternalLinkService">
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<mods:title id="071F611AF7369D6C527FAC1842B7E333">Revision of Angursa (Arthrotardigrada: Styraconyxidae) with the description of a new species from Japan</mods:title>
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<treatment id="03D3878DFF9A58191EE26771DE31F942" ID-DOI="http://doi.org/10.5281/zenodo.5941842" ID-GBIF-Taxon="215923441" ID-Zenodo-Dep="5941842" LSID="urn:lsid:plazi:treatment:03D3878DFF9A58191EE26771DE31F942" httpUri="http://treatment.plazi.org/id/03D3878DFF9A58191EE26771DE31F942" lastPageNumber="13" pageId="12" pageNumber="13">
<subSubSection id="C3606510FF9A58191EE26771D9C3FC93" box="[189,645,864,890]" pageId="12" pageNumber="13" type="nomenclature">
<paragraph id="8BC5369BFF9A58191EE26771D9C3FC93" blockId="12.[189,645,864,890]" box="[189,645,864,890]" pageId="12" pageNumber="13">
<heading id="D08D81F7FF9A58191EE26771D9C3FC93" bold="true" box="[189,645,864,890]" fontSize="11" level="3" pageId="12" pageNumber="13" reason="3">
<emphasis id="B90EEA89FF9A58191EE26771D9C3FC93" bold="true" box="[189,645,864,890]" pageId="12" pageNumber="13">
Identification key to
<taxonomicName id="4C7A4D18FF9A58191FE06771D961FC93" ID-CoL="7443d693-6738-42b4-9e43-1b41918cea81" authorityName="Pollock" authorityYear="1979" box="[447,551,864,890]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="genus">
<emphasis id="B90EEA89FF9A58191FE06771D961FC93" bold="true" box="[447,551,864,890]" italics="true" pageId="12" pageNumber="13">Angursa</emphasis>
</taxonomicName>
species
</emphasis>
</heading>
</paragraph>
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<key id="FFEB61AAFF9A58191EE26781DE31F942" pageId="12" pageNumber="13">
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<keyLead id="308B95AEFF9A58191EE26781DE30FC26" pageId="12" pageNumber="13">
<paragraph id="8BC5369BFF9A58191EE26781DE30FC43" blockId="12.[189,1399,912,1707]" box="[189,1398,912,939]" pageId="12" pageNumber="13">
1. Club-shaped primary clavae longer than lateral cirri ........................................ 2 (
<taxonomicName id="4C7A4D18FF9A58191AC26781DE67FC42" ID-CoL="5c7b78a3-04a7-4982-8887-6b8ad60882ec" authorityName="Pollock" authorityYear="1979" box="[1181,1313,912,939]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="bicuspis">
<emphasis id="B90EEA89FF9A58191AC26781DE67FC42" bold="true" box="[1181,1313,912,939]" italics="true" pageId="12" pageNumber="13">A. bicuspis</emphasis>
</taxonomicName>
group)
</paragraph>
<paragraph id="8BC5369BFF9A58191E9E67A4DE30FC26" blockId="12.[189,1399,912,1707]" box="[193,1398,948,975]" pageId="12" pageNumber="13">
Bulbous primary clavae shorter than lateral cirri ...........................................6 (
<taxonomicName id="4C7A4D18FF9A58191ADE67A4DE66FC26" ID-CoL="86743074-accd-4d51-93f9-8ffac7ea8271" authorityName="Renaud-Mornant" authorityYear="1981" box="[1153,1312,948,975]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="lanceolata">
<emphasis id="B90EEA89FF9A58191ADE67A4DE66FC26" bold="true" box="[1153,1312,948,975]" italics="true" pageId="12" pageNumber="13">A. lanceolata</emphasis>
</taxonomicName>
group)
</paragraph>
</keyLead>
</keyStep>
<keyStep id="308E2E3EFF9A58191EE267E9DE31FBDF" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191EE267E9DE31FBDF" pageId="12" pageNumber="13">
<paragraph id="8BC5369BFF9A58191EE267E9DE31FBFB" blockId="12.[189,1399,912,1707]" box="[189,1399,1015,1042]" pageId="12" pageNumber="13">
2. Well developed anal papillae .....................................................................
<taxonomicName id="4C7A4D18FF9A58191A1767E9DE31FBFB" ID-CoL="5c7b78a3-04a7-4982-8887-6b8ad60882ec" authority="Pollock, 1979" authorityName="Pollock" authorityYear="1979" box="[1096,1399,1015,1042]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="bicuspis">
<emphasis id="B90EEA89FF9A58191A1767E9DF8AFBFB" bold="true" box="[1096,1228,1015,1042]" italics="true" pageId="12" pageNumber="13">A. bicuspis</emphasis>
<bibRefCitation id="EFEB4B6AFF9A58191A8C67E6DE31FBFB" author="Pollock L. W." box="[1235,1399,1015,1042]" pageId="12" pageNumber="13" pagination="558 - 562" refId="ref9514" refString="Pollock L. W. 1979. Angursa bicuspis n. g., n. sp., a new marine arthrotardigrade from the western North Atlantic. Transactions of the American Microscopical Society 98: 558 - 562. https: // doi. org / 10.2307 / 3225907" type="journal article" year="1979">Pollock, 1979</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BC5369BFF9A58191E9E600CDE31FBDF" blockId="12.[189,1399,912,1707]" box="[193,1399,1052,1079]" pageId="12" pageNumber="13"> Anal papillae absent........................................................................................................................... 3</paragraph>
</keyLead>
</keyStep>
<keyStep id="308E2E3EFF9A58191EE2604EDE31FB77" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191EE2604EDE31FB77" pageId="12" pageNumber="13">
<paragraph id="8BC5369BFF9A58191EE2604EDE31FB93" blockId="12.[189,1399,912,1707]" box="[189,1399,1119,1146]" pageId="12" pageNumber="13">
3. Legs II and III sensory organs present; spiral seminal receptacle ducts........
<taxonomicName id="4C7A4D18FF9A58191A05604EDE31FB93" ID-CoL="75d2002b-4a9a-44fe-bba3-6a282a92a626" authority="Noda, 1985" authorityName="Noda" authorityYear="1985" box="[1114,1399,1119,1146]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="clavifera">
<emphasis id="B90EEA89FF9A58191A05604EDFA0FB93" bold="true" box="[1114,1254,1119,1146]" italics="true" pageId="12" pageNumber="13">A. clavifera</emphasis>
<bibRefCitation id="EFEB4B6AFF9A58191AB1604EDE31FB93" author="Noda H." box="[1262,1399,1119,1146]" pageId="12" pageNumber="13" pagination="269 - 276" refId="ref9298" refString="Noda H. 1985. Description of a new subspecies of Angursa bicuspis Pollock (Heterotardigrada, Halechiniscidae) from Tanabe Bay, Japan. Publications of the Seto Marine Biological Laboratory 30: 269 - 276. Available from https: // hdl. handle. net / 2433 / 176110 [accessed 4 Feb. 2019]." type="journal article" year="1985">Noda, 1985</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BC5369BFF9A58191E9E6095DE31FB77" blockId="12.[189,1399,912,1707]" box="[193,1399,1156,1182]" pageId="12" pageNumber="13"> Leg II and III sensory organs absent.................................................................................................. 4</paragraph>
</keyLead>
</keyStep>
<keyStep id="308E2E3EFF9A58191EE260D6DE31FAC2" pageId="12" pageNumber="13">
<paragraph id="8BC5369BFF9A58191EE260D6DE31FAEF" blockId="12.[189,1399,912,1707]" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191EE260D6DE31FAEF" pageId="12" pageNumber="13">
4. Three-lobed tertiary clavae; D-shaped secondary clavae; seminal receptacles present ...................... .....................................................................................................................
<taxonomicName id="4C7A4D18FF9A58191A0E60FDDE53FAEF" ID-CoL="9bec59d6-dfd9-4ff8-b1e4-3c5bf21491ef" authority="Fujimoto &amp; Hansen, 2019" authorityName="Fujimoto &amp; Hansen" authorityYear="2019" box="[1105,1301,1260,1286]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="seisuimaruae" status="sp. nov.">
<emphasis id="B90EEA89FF9A58191A0E60FDDE53FAEF" bold="true" box="[1105,1301,1260,1286]" italics="true" pageId="12" pageNumber="13">A. seisuimaruae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57F2FF9A58191B4360FDDE31FAEF" box="[1308,1399,1260,1286]" pageId="12" pageNumber="13" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC5369BFF9A58191E9E6100DE31FAC2" blockId="12.[189,1399,912,1707]" box="[193,1399,1296,1323]" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191E9E6100DE31FAC2" box="[193,1399,1296,1323]" pageId="12" pageNumber="13"> Tertiary clavae with no indentation or only slightly indented around external cirri.......................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E2E3EFF9A58191EE26142DE31FA5E" pageId="12" pageNumber="13">
<paragraph id="8BC5369BFF9A58191EE26142DE31FA7B" blockId="12.[189,1399,912,1707]" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191EE26142DE31FA7B" pageId="12" pageNumber="13">
5. Secondary clavae situated close together (ca 3 μm); short (10 μm) cirrus E ...................................... ....................................................................................................
<taxonomicName id="4C7A4D18FF9A58191D8F6169DE31FA7B" ID-CoL="85d4a8a4-7dd8-4e00-9e0c-3faf8b02d2ad" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" box="[976,1399,1400,1427]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="abyssalis">
<emphasis id="B90EEA89FF9A58191D8F6169DF18FA7A" bold="true" box="[976,1118,1400,1427]" italics="true" pageId="12" pageNumber="13">A. abyssalis</emphasis>
<bibRefCitation id="EFEB4B6AFF9A58191A3A6169DE31FA7B" author="Renaud-Mornant J." box="[1125,1399,1400,1426]" pageId="12" pageNumber="13" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant, 1981</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC5369BFF9A58191E9E618CDE31FA5E" blockId="12.[189,1399,912,1707]" box="[193,1399,1436,1463]" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191E9E618CDE31FA5E" box="[193,1399,1436,1463]" pageId="12" pageNumber="13">
Secondary clavae situated apart (ca 9 μm); long (19 μm) cirrus E...............
<taxonomicName id="4C7A4D18FF9A58191A0D618CDE31FA5E" ID-CoL="443f3a26-cc50-439f-8800-4be5d848728a" authority="Bussau, 1992" authorityName="Bussau" authorityYear="1992" box="[1106,1399,1436,1463]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="capsula">
<emphasis id="B90EEA89FF9A58191A0D618CDF97FA5E" bold="true" box="[1106,1233,1436,1463]" italics="true" pageId="12" pageNumber="13">A. capsula</emphasis>
<bibRefCitation id="EFEB4B6AFF9A58191A87618CDE31FA5E" author="Bussau C." box="[1240,1399,1437,1463]" pageId="12" pageNumber="13" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau, 1992</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E2E3EFF9A58191EE261CEDE31F9AD" pageId="12" pageNumber="13">
<paragraph id="8BC5369BFF9A58191EE261CEDE31F9F7" blockId="12.[189,1399,912,1707]" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191EE261CEDE31F9F7" pageId="12" pageNumber="13">
6. Leg IV sensory organs as papillae enveloped in hemispherical sheaths; cirri E with annulated scapi ............................................................................................
<taxonomicName id="4C7A4D18FF9A58191D8D6215DE31F9F7" ID-CoL="0cfad9a9-2791-4cab-9641-8f247c8ae880" authority="Villora-Moreno, 1998" authorityName="Villora-Moreno" authorityYear="1998" box="[978,1399,1540,1566]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="antarctica">
<emphasis id="B90EEA89FF9A58191D8D6215DF28F9F7" bold="true" box="[978,1134,1540,1566]" italics="true" pageId="12" pageNumber="13">A. antarctica</emphasis>
<bibRefCitation id="EFEB4B6AFF9A58191A2A6215DE31F9F7" author="Villora-Moreno S." box="[1141,1399,1540,1566]" pageId="12" pageNumber="13" pagination="336 - 341" refId="ref9804" refString="Villora-Moreno S. 1998. Deep-sea Tardigrada from South Shetland Islands (Antarctica) with description of Angursa antarctica sp. nov. (Arthrotardigrada, Halechiniscidae). Polar Biology 19: 336 - 341. https: // doi. org / 10.1007 / s 003000050255" type="journal article" year="1998">Villora-Moreno, 1998</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC5369BFF9A58191E9E623BDE31F9AD" blockId="12.[189,1399,912,1707]" box="[193,1399,1577,1604]" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191E9E623BDE31F9AD" box="[193,1399,1577,1604]" pageId="12" pageNumber="13"> Leg IV sensory organs as elongate papillae....................................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="308E2E3EFF9A58191EE2627DDE31F942" pageId="12" pageNumber="13">
<keyLead id="308B95AEFF9A58191EE2627DDE31F942" pageId="12" pageNumber="13">
<paragraph id="8BC5369BFF9A58191EE2627DDE31F96F" blockId="12.[189,1399,912,1707]" box="[189,1399,1644,1671]" pageId="12" pageNumber="13">
7. Cirri E lance-shaped ................................................................
<taxonomicName id="4C7A4D18FF9A58191DE1627DDE31F96F" ID-CoL="86743074-accd-4d51-93f9-8ffac7ea8271" authority="Renaud-Mornant, 1981" authorityName="Renaud-Mornant" authorityYear="1981" box="[958,1399,1644,1671]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="lanceolata">
<emphasis id="B90EEA89FF9A58191DE1627DDF1BF96E" bold="true" box="[958,1117,1644,1671]" italics="true" pageId="12" pageNumber="13">A. lanceolata</emphasis>
<bibRefCitation id="EFEB4B6AFF9A58191A3A627DDE31F96F" author="Renaud-Mornant J." box="[1125,1399,1644,1670]" pageId="12" pageNumber="13" pagination="161 - 164" refId="ref9565" refString="Renaud-Mornant J. 1981. Deux nouveaux Angursa Pollock, 1979, du domaine abyssal (Tardigrada, Arthrotardigrada). Tethys 10: 161 - 164." type="journal article" year="1981">Renaud-Mornant, 1981</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BC5369BFF9A58191E9E6280DE31F942" blockId="12.[189,1399,912,1707]" box="[193,1399,1680,1707]" pageId="12" pageNumber="13">
Cirri E tongue-shaped .....................................................................................
<taxonomicName id="4C7A4D18FF9A58191A3E6280DE31F942" ID-CoL="be864703-f48d-4cbc-9e6a-20c878d4e8f5" authority="Bussau, 1992" authorityName="Bussau" authorityYear="1992" box="[1121,1399,1680,1707]" class="Heterotardigrada" family="Styraconyxidae" genus="Angursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Arthrotardigrada" pageId="12" pageNumber="13" phylum="Tardigrada" rank="species" species="lingua">
<emphasis id="B90EEA89FF9A58191A3E6280DF97F942" bold="true" box="[1121,1233,1680,1707]" italics="true" pageId="12" pageNumber="13">A. lingua</emphasis>
<bibRefCitation id="EFEB4B6AFF9A58191A876280DE31F942" author="Bussau C." box="[1240,1399,1681,1707]" pageId="12" pageNumber="13" pagination="79 - 91" refId="ref8261" refString="Bussau C. 1992. New deep-sea Tardigrada (Arthrotardigrada, Halechiniscidae) from manganese nodule area of the eastern South Pacific. Zoologica Scripta 21: 79 - 91. https: // doi. org / 10.1111 / j. 1463 - 6409.1992. tb 00311. x" type="journal article" year="1992">Bussau, 1992</bibRefCitation>
</taxonomicName>
</paragraph>
</keyLead>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>