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156 lines
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<document id="2EB7A4A193949C9371845854AC3EFBAC" ID-DOI="10.11646/zootaxa.2435.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5315673" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629671386604" checkinUser="felipe" docAuthor="Oláh, János & Johanson, Kjell Arne" docDate="2010" docId="0393CE26FFD3FFD17CFB8B13FBB7F831" docLanguage="en" docName="zt02435p063.pdf" docOrigin="Zootaxa 2435 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2435.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Nyctiophylax (Paranyctiophylax) dhauli Oláh & Johanson 2010, new species" docType="treatment" docVersion="5" lastPageNumber="46" masterDocId="FFAAB65EFFFEFFFC7C6C8801FFE1FFE5" masterDocTitle="Generic review of Polycentropodidae with description of 32 new species and 19 new species records from the Oriental, Australian and Afrotropical Biogeographical Regions 2435" masterLastPageNumber="63" masterPageNumber="1" pageNumber="46" updateTime="1699093793393" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="85EB37447475AC377E1F37C1AA0811ED">
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<mods:title id="BE133B614DBFBB18CA21EC76A4850916">Generic review of Polycentropodidae with description of 32 new species and 19 new species records from the Oriental, Australian and Afrotropical Biogeographical Regions 2435</mods:title>
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<mods:name id="AA3B35820F342F2A485461DD3998B490" type="personal">
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<mods:namePart id="04EBFA12DB35DBD5412B9E7C0CD6051A">Oláh, János</mods:namePart>
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</mods:name>
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<mods:namePart id="62EBAEB2A5CC7CBB8649F8219CA5DF3B">Johanson, Kjell Arne</mods:namePart>
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<mods:typeOfResource id="C85BBDADF887D86A5E2F3D774D77D9A1">text</mods:typeOfResource>
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<mods:title id="04EFAE33532251EC0D5947F17BEF22D1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D9FF9F508D9E64857465F38278015EBE">
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<mods:date id="D28ECB9F1F0278B7F731EF22FE266105">2010</mods:date>
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<mods:number id="C389A4E1A636E1DF76BB21A474AF719B">2010-04-23</mods:number>
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</mods:detail>
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<mods:detail id="7AED29C694BCEFE7A0346C8358CD6353" type="volume">
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<mods:number id="F4C6C9B0B87E3607682B1F4A58FB8197">2435</mods:number>
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</mods:detail>
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<mods:detail id="007291E979223024227E8F93E8072C76" type="issue">
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<mods:number id="30EE8948445B007CF747641DB4623156">1</mods:number>
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<mods:url id="989C4FD50CDA2A7B21A7B95D5CF067F7">https://biotaxa.org/Zootaxa/article/view/zootaxa.2435.1.1</mods:url>
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</mods:location>
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<mods:classification id="5597380DB2220A37E4B1E9439DAB676E">journal article</mods:classification>
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<mods:identifier id="706A21755D813B14B9B30AC298CEFC02" type="DOI">10.11646/zootaxa.2435.1.1</mods:identifier>
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<mods:identifier id="A3D694986837B57F06E5D902ABA169CB" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="037724E3222EB5E1BB4C062ABCA50B30" type="Zenodo-Dep">5315673</mods:identifier>
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<treatment id="0393CE26FFD3FFD17CFB8B13FBB7F831" ID-DOI="http://doi.org/10.5281/zenodo.5324102" ID-Zenodo-Dep="5324102" LSID="urn:lsid:plazi:treatment:0393CE26FFD3FFD17CFB8B13FBB7F831" httpUri="http://treatment.plazi.org/id/0393CE26FFD3FFD17CFB8B13FBB7F831" lastPageNumber="46" pageId="45" pageNumber="46">
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<subSubSection id="C3202CBBFFD3FFD17CFB8B13FCFBFCC9" box="[151,794,786,812]" pageId="45" pageNumber="46" type="nomenclature">
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<paragraph id="8B857F30FFD3FFD17CFB8B13FCFBFCC9" blockId="45.[151,794,786,847]" box="[151,794,786,812]" pageId="45" pageNumber="46">
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<heading id="D0CDC85CFFD3FFD17CFB8B13FCFBFCC9" bold="true" box="[151,794,786,812]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
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<emphasis id="B94EA322FFD3FFD17CFB8B13FCFBFCC9" bold="true" box="[151,794,786,812]" pageId="45" pageNumber="46">
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<taxonomicName id="4C3A04B3FFD3FFD17CFB8B13FD9DFCC9" authority="Oláh & Johanson, 2010" authorityName="Oláh & Johanson" authorityYear="2010" box="[151,636,786,812]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="dhauli" status="sp. nov." subGenus="Paranyctiophylax">
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<emphasis id="B94EA322FFD3FFD17CFB8B13FED6FCC9" bold="true" box="[151,311,786,812]" italics="true" pageId="45" pageNumber="46">Nyctiophylax</emphasis>
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(
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<emphasis id="B94EA322FFD3FFD17D2B8B13FDFDFCC9" bold="true" box="[327,540,786,812]" italics="true" pageId="45" pageNumber="46">Paranyctiophylax</emphasis>
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)
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<emphasis id="B94EA322FFD3FFD17E428B13FD9DFCC9" bold="true" box="[558,636,786,812]" italics="true" pageId="45" pageNumber="46">dhauli</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A27D1E59FFD3FFD17EE78B13FCFBFCC9" box="[651,794,786,812]" pageId="45" pageNumber="46" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3202CBBFFD3FFD17CFB8B34FEDEFCAA" box="[151,319,821,847]" pageId="45" pageNumber="46" type="description">
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<paragraph id="8B857F30FFD3FFD17CFB8B34FEDEFCAA" blockId="45.[151,794,786,847]" box="[151,319,821,847]" pageId="45" pageNumber="46">
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<figureCitation id="130163B5FFD3FFD17CFB8B34FEDEFCAA" box="[151,319,821,847]" captionStart="FIGURES 80–82" captionStartId="46.[151,270,1408,1432]" captionTargetBox="[159,1424,202,1369]" captionTargetId="figure-198@46.[151,1436,165,1387]" captionTargetPageId="46" captionText="FIGURES 80–82. Nyctiophylax (Paranyctiophylax) dhauli, new species, holotype. 80—genitalia, left lateral; 81—left gonopod, ventral; 82—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361556" httpUri="https://zenodo.org/record/5361556/files/figure.png" pageId="45" pageNumber="46">Figures 80–82</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3202CBBFFD3FFD17CFB8B7EFC35FB21" pageId="45" pageNumber="46" type="diagnosis">
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<paragraph id="8B857F30FFD3FFD17CFB8B7EFC35FB21" blockId="45.[151,1436,895,2004]" pageId="45" pageNumber="46">
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<emphasis id="B94EA322FFD3FFD17CFB8B7EFEFDFC7C" bold="true" box="[151,284,895,921]" pageId="45" pageNumber="46">Diagnosis:</emphasis>
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The gonopod shape of this species is intermediate between those of species with short, basally curved gonopods and species with a long ventral lobe of each gonopod. The gonopods in this species are accompanied by slender dorsal and ventral paraproctal processes. There are several species from
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<collectingCountry id="F32D3FA0FFD3FFD179678BCBFA9CFC01" box="[1291,1405,970,996]" name="Sri Lanka" pageId="45" pageNumber="46">Sri Lanka</collectingCountry>
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to
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<collectingCountry id="F32D3FA0FFD3FFD17CFB8BF1FEEBFBEF" box="[151,266,1008,1034]" name="Indonesia" pageId="45" pageNumber="46">Indonesia</collectingCountry>
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with similar genitalia. This species strongly resembles
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<taxonomicName id="4C3A04B3FFD3FFD17FC98BF1FB5EFBEF" authority="Malicky" authorityName="Malicky" authorityYear="1997" box="[933,1215,1008,1034]" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="amykos" subGenus="Paranyctiophylax">
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<emphasis id="B94EA322FFD3FFD17FC98BF1FBB5FBEC" box="[933,1108,1008,1033]" italics="true" pageId="45" pageNumber="46">N. (P.) amykos</emphasis>
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Malicky
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</taxonomicName>
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from
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<collectingCountry id="F32D3FA0FFD3FFD179658BF1FAB5FBEF" box="[1289,1364,1008,1034]" name="Nepal" pageId="45" pageNumber="46">Nepal</collectingCountry>
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. In
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<taxonomicName id="4C3A04B3FFD3FFD179ED8BF1FEF2FBCB" authorityName="Oláh & Johanson" authorityYear="2010" class="Insecta" family="Polycentropodidae" genus="Nyctiophylax" kingdom="Animalia" order="Trichoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="dhauli" subGenus="Paranyctiophylax">
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<emphasis id="B94EA322FFD3FFD179ED8BF1FEF2FBCB" italics="true" pageId="45" pageNumber="46">N. (P.) dhauli</emphasis>
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</taxonomicName>
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, the sternite IX is vertically tall, not horizontally elongate; the apicoventral mesal lobe of sternite IX is sharply triangular, not blunt and rounded in lateral view; the cerci are elongate, foliaceous, not narrow at basal 1/3rd; the mesoventral branch of each gonopod is longer; the dorsal branch of each gonopod is more slender with a concave dorsum, not stout and convex; the dorsal paraproctal processes are slightly longer than the cerci; and the ventral paraproctal processes are dorsally geniculate.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3202CBBFFD3FFD17CA88CCEFA7DF94F" pageId="45" pageNumber="46" type="description">
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<paragraph id="8B857F30FFD3FFD17CA88CCEFD71FAEA" blockId="45.[151,1436,895,2004]" pageId="45" pageNumber="46">
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<emphasis id="B94EA322FFD3FFD17CA88CCEFEBEFB0C" bold="true" box="[196,351,1231,1257]" pageId="45" pageNumber="46">Description:</emphasis>
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Male. Body uniformly pale brown. Maxillary palp formula (I,II)-IV-III-V. Forewing length 4.0 mm. Forewing A1, A2, and A3 looped.
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</paragraph>
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<paragraph id="8B857F30FFD3FFD17CA88D1BFA7DF94F" blockId="45.[151,1436,895,2004]" pageId="45" pageNumber="46">
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Male genitalia. Abdominal segment IX reduced to robust sternite, tall, nearly triangular (
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<figureCitation id="130163B5FFD3FFD178B78D1BFAD2FAD1" box="[1243,1331,1306,1332]" captionStart="FIGURES 80–82" captionStartId="46.[151,270,1408,1432]" captionTargetBox="[159,1424,202,1369]" captionTargetId="figure-198@46.[151,1436,165,1387]" captionTargetPageId="46" captionText="FIGURES 80–82. Nyctiophylax (Paranyctiophylax) dhauli, new species, holotype. 80—genitalia, left lateral; 81—left gonopod, ventral; 82—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361556" httpUri="https://zenodo.org/record/5361556/files/figure.png" pageId="45" pageNumber="46">Fig. 80</figureCitation>
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); tergum IX membranous, with no narrow sclerotized stripe discernible. Segment X membranous, covering dorsal part of phallic apparatus. Cerci setose, narrow, elongate, foliaceous in lateral view; slightly shorter than dorsal paraproctal processes. Paraproctal complexes well-developed, smooth; each composed of paraproctal base (anterior body formed by meeting point of dorsal and ventral paraproctal processes) heavily sclerotized, narrow; dorsal paraproctal process slender, weakly sclerotized, straight in lateral view; and ventral paraproctal process with dorsum geniculate, apex with several small setae. Gonopods with short, sharply triangular mesoventral branch (
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<figureCitation id="130163B5FFD3FFD17DE28E21FE07F9DF" box="[398,486,1568,1594]" captionStart="FIGURES 80–82" captionStartId="46.[151,270,1408,1432]" captionTargetBox="[159,1424,202,1369]" captionTargetId="figure-198@46.[151,1436,165,1387]" captionTargetPageId="46" captionText="FIGURES 80–82. Nyctiophylax (Paranyctiophylax) dhauli, new species, holotype. 80—genitalia, left lateral; 81—left gonopod, ventral; 82—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361556" httpUri="https://zenodo.org/record/5361556/files/figure.png" pageId="45" pageNumber="46">Fig. 80</figureCitation>
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); dorsal branch long, slender, with concave dorsum (
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<figureCitation id="130163B5FFD3FFD1783F8E21FB4AF9DF" box="[1107,1195,1568,1594]" captionStart="FIGURES 80–82" captionStartId="46.[151,270,1408,1432]" captionTargetBox="[159,1424,202,1369]" captionTargetId="figure-198@46.[151,1436,165,1387]" captionTargetPageId="46" captionText="FIGURES 80–82. Nyctiophylax (Paranyctiophylax) dhauli, new species, holotype. 80—genitalia, left lateral; 81—left gonopod, ventral; 82—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361556" httpUri="https://zenodo.org/record/5361556/files/figure.png" pageId="45" pageNumber="46">Fig. 80</figureCitation>
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); dorsal branch with blunt apex. Phallic apparatus (
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<figureCitation id="130163B5FFD3FFD17E768E44FD97F9BA" box="[538,630,1605,1631]" captionStart="FIGURES 80–82" captionStartId="46.[151,270,1408,1432]" captionTargetBox="[159,1424,202,1369]" captionTargetId="figure-198@46.[151,1436,165,1387]" captionTargetPageId="46" captionText="FIGURES 80–82. Nyctiophylax (Paranyctiophylax) dhauli, new species, holotype. 80—genitalia, left lateral; 81—left gonopod, ventral; 82—phallic apparatus, left lateral." figureDoi="http://doi.org/10.5281/zenodo.5361556" httpUri="https://zenodo.org/record/5361556/files/figure.png" pageId="45" pageNumber="46">Fig. 82</figureCitation>
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) located dorsally in genitalia, guided by paraproctal processes, membranous segment X, and membranous subphallic bridge. Phallotheca thick, horizontal; endophallus with pair of slightly sclerotized sclerites and ventrad-curving spines; apical part ending in row of elongate spicules.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3202CBBFFD3FFD17CA88EB4FF0AF911" pageId="45" pageNumber="46" type="materials_examined">
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<paragraph id="8B857F30FFD3FFD17CA88EB4FF0AF911" blockId="45.[151,1436,895,2004]" pageId="45" pageNumber="46">
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<materialsCitation id="3B52756DFFD3FFD17CA88EB4FF0AF911" collectingDate="1987-02-20" collectingDateMax="1987-02-28" collectingDateMin="1987-02-20" collectionCode="OPC" collectorName="J. Olah" country="India" location="Dhauli" municipality="Bhubaneswar" pageId="45" pageNumber="46" specimenCount="1" stateProvince="Odisha" typeStatus="holotype">
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<emphasis id="B94EA322FFD3FFD17CA88EB4FE0CF92A" bold="true" box="[196,493,1717,1743]" pageId="45" pageNumber="46">
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<typeStatus id="5481C192FFD3FFD17CA88EB4FED7F92A" box="[196,310,1717,1743]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
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male:
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<collectingCountry id="F32D3FA0FFD3FFD17DE18EB4FE09F92A" box="[397,488,1717,1743]" name="India" pageId="45" pageNumber="46">INDIA</collectingCountry>
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:
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</emphasis>
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<collectingRegion id="49FEB1D2FFD3FFD17D9A8EB4FD66F92A" box="[502,647,1717,1743]" country="India" name="Odisha" pageId="45" pageNumber="46">Orissa State</collectingRegion>
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,
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<collectingMunicipality id="6BE1E54AFFD3FFD17EFF8EB4FCD4F92A" box="[659,821,1717,1743]" pageId="45" pageNumber="46">Bhubaneswar</collectingMunicipality>
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,
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<location id="8EE529EBFFD3FFD17F2E8EB4FC74F92A" LSID="urn:lsid:plazi:treatment:0393CE26FFD3FFD17CFB8B13FBB7F831:8EE529EBFFD3FFD17F2E8EB4FC74F92A" box="[834,917,1717,1743]" country="India" municipality="Bhubaneswar" name="Dhauli" pageId="45" pageNumber="46" stateProvince="Odisha">Dhauli</location>
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, marshy area,
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<date id="FF8459F0FFD3FFD1782E8EB4FB09F92A" box="[1090,1256,1717,1743]" pageId="45" pageNumber="46" value="1987-02-20" valueMax="1987-02-28" valueMin="1987-02-20">
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<collectingDate id="EFC0A018FFD3FFD1782E8EB4FB09F92A" box="[1090,1256,1717,1743]" pageId="45" pageNumber="46" value="1987-02-20" valueMax="1987-02-28" valueMin="1987-02-20">20–28.ii.1987</collectingDate>
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</date>
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, light [
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<collectorName id="26CF1AE6FFD3FFD179518EB4FA75F92A" box="[1341,1428,1717,1743]" pageId="45" pageNumber="46">J. Oláh</collectorName>
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] (
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<collectionCode id="ED2BE7F5FFD3FFD17CF38EDBFF3FF911" box="[159,222,1754,1780]" pageId="45" pageNumber="46">OPC</collectionCode>
|
|||
|
|
).
|
|||
|
|
</materialsCitation>
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3202CBBFFD3FFD17CAA8EFEFE55F8FF" box="[198,436,1791,1818]" pageId="45" pageNumber="46" type="distribution">
|
|||
|
|
<paragraph id="8B857F30FFD3FFD17CAA8EFEFE55F8FF" blockId="45.[151,1436,895,2004]" box="[198,436,1791,1818]" pageId="45" pageNumber="46">
|
|||
|
|
<emphasis id="B94EA322FFD3FFD17CAA8EFEFE88F8FC" bold="true" box="[198,361,1791,1817]" pageId="45" pageNumber="46">Distribution:</emphasis>
|
|||
|
|
<collectingCountry id="F32D3FA0FFD3FFD17D1C8F01FE51F8FF" box="[368,432,1792,1818]" name="India" pageId="45" pageNumber="46">India</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3202CBBFFD3FFD17CA88F24FBB7F831" pageId="45" pageNumber="46" type="etymology">
|
|||
|
|
<paragraph id="8B857F30FFD3FFD17CA88F24FBB7F831" blockId="45.[151,1436,895,2004]" pageId="45" pageNumber="46">
|
|||
|
|
<emphasis id="B94EA322FFD3FFD17CA88F24FEB5F8DA" bold="true" box="[196,340,1829,1855]" pageId="45" pageNumber="46">Etymology:</emphasis>
|
|||
|
|
<emphasis id="B94EA322FFD3FFD17D308F24FE4CF8DB" box="[348,429,1829,1854]" italics="true" pageId="45" pageNumber="46">Dhauli</emphasis>
|
|||
|
|
, named after the Dhauli Hills at Daya River in
|
|||
|
|
<collectingRegion id="49FEB1D2FFD3FFD17FBB8F24FB87F8DA" box="[983,1126,1829,1855]" country="India" name="Odisha" pageId="45" pageNumber="46">Orissa State</collectingRegion>
|
|||
|
|
, near Bhubaneswar,
|
|||
|
|
<collectingCountry id="F32D3FA0FFD3FFD179348F24FA76F8DA" box="[1368,1431,1829,1855]" name="India" pageId="45" pageNumber="46">India</collectingCountry>
|
|||
|
|
, where this animal was collected in a marshy area of the famous battlefield of the bloody war between King Ashoka and the King of Kalinga. The water of River Daya turned red and Ashoka, witnessing the mass deaths, thereafter pursued a policy of non-violence. After converting to Buddhism, he was responsible for the proliferation of Buddhist ideals across the whole of
|
|||
|
|
<collectingCountry id="F32D3FA0FFD3FFD17E9B8FBBFC86F831" box="[759,871,1978,2004]" name="China" pageId="45" pageNumber="46">East Asia</collectingCountry>
|
|||
|
|
and Southeast Asia.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|