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<document id="D9086725931074C5B759337C230F90A5" ID="10.11646/zootaxa.4236.1.2SLASH10251" ID-CLB-Dataset="36479" ID-DOI="10.11646/zootaxa.4236.1.2" ID-GBIF-Dataset="0f39b5c0-0af5-476f-8448-a02339cc1dfd" ID-ISSN="1175-5326" ID-Zenodo-Dep="322014" ID-ZooBank="23412386-CB17-49CA-9C47-BD71DD9C5372" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487752177037" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N. &amp; Yushkova, Olga V." docDate="2017" docId="4F3A5135FFB5DC0E4FE3FE806B28F908" docLanguage="en" docName="zootaxa.4236.1.2SLASH10251.pdf" docOrigin="Zootaxa 4236 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Placoblatta beybienkoi Anisyutkin, sp.nov." docType="treatment" docVersion="7" lastPageNumber="54" masterDocId="B303294DFFB8DC034F74FFCC6F0CFF9F" masterDocTitle="New data on cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from India and Sri Lanka, with descriptions of new species and the genital complex of Aptera fusca (Thunberg, 1784)" masterLastPageNumber="64" masterPageNumber="41" pageNumber="54" updateTime="1698437163562" updateUser="ExternalLinkService">
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<mods:title id="09FDA42CF74078EBC2E7A8A013C96153">New data on cockroaches of the subfamily Epilamprinae (Dictyoptera: Blaberidae) from India and Sri Lanka, with descriptions of new species and the genital complex of Aptera fusca (Thunberg, 1784)</mods:title>
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<mods:namePart id="A596058B86E7D7E55DE34C34301CA528">Anisyutkin, Leonid N.</mods:namePart>
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<mods:namePart id="3CD9BE49924E096EEADC6B4CFB2EF112">Yushkova, Olga V.</mods:namePart>
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<mods:typeOfResource id="BF8D117AFAEE1F81CE960169FCE57840">text</mods:typeOfResource>
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<mods:title id="D2D97A9BB238C95D2FB8FC5CCFFBC946">Zootaxa</mods:title>
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<mods:date id="49417DDDC2A9F46EABB6C41B1121530E">2017</mods:date>
<mods:detail id="824D21A1C8820652CB124FEA869B7338" type="volume">
<mods:number id="0DD5D5A9A735BDE2E5872E6203EBDD82">4236</mods:number>
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<mods:classification id="3920EFF75DBE90AC8C03BA44648AF106">journal article</mods:classification>
<mods:identifier id="F50880FD42E02D3B0B3E52C42ED92552" type="CLB-Dataset">36479</mods:identifier>
<mods:identifier id="D94429843D499BDE9487DE0953290B19" type="DOI">10.11646/zootaxa.4236.1.2</mods:identifier>
<mods:identifier id="0004783E650FBFE54CB4E06F816902B0" type="GBIF-Dataset">0f39b5c0-0af5-476f-8448-a02339cc1dfd</mods:identifier>
<mods:identifier id="E76BA7B9D5938876EA6DA739D73E0E14" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="5BFE66CFB9635E308B3D1821B8896C52" type="Zenodo-Dep">322014</mods:identifier>
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<treatment id="4F3A5135FFB5DC0E4FE3FE806B28F908" ID-DOI="http://doi.org/10.5281/zenodo.6015709" ID-GBIF-Taxon="127697118" ID-Zenodo-Dep="6015709" LSID="urn:lsid:plazi:treatment:4F3A5135FFB5DC0E4FE3FE806B28F908" httpUri="http://treatment.plazi.org/id/4F3A5135FFB5DC0E4FE3FE806B28F908" lastPageNumber="54" pageId="13" pageNumber="54">
<subSubSection id="8F89B3A8FFB5DC0E4FE3FE806EBDFE18" pageId="13" pageNumber="54" type="nomenclature">
<paragraph id="C72CE023FFB5DC0E4FE3FE806D93FEF9" blockId="13.[151,671,332,391]" box="[151,671,332,358]" pageId="13" pageNumber="54">
<heading id="9C64574FFFB5DC0E4FE3FE806D93FEF9" bold="true" box="[151,671,332,358]" fontSize="11" level="1" pageId="13" pageNumber="54" reason="1">
<emphasis id="F5E73C31FFB5DC0E4FE3FE806D93FEF9" bold="true" box="[151,671,332,358]" pageId="13" pageNumber="54">
<taxonomicName id="00939BA0FFB5DC0E4FE3FE806D30FEF9" authority="Anisyutkin" authorityName="Anisyutkin" box="[151,572,332,358]" class="Insecta" family="Blaberidae" genus="Placoblatta" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="beybienkoi" status="sp. nov.">
<emphasis id="F5E73C31FFB5DC0E4FE3FE806EA4FEF9" bold="true" box="[151,424,332,358]" italics="true" pageId="13" pageNumber="54">Placoblatta beybienkoi</emphasis>
Anisyutkin
</taxonomicName>
,
<taxonomicNameLabel id="EED4814AFFB5DC0E4D3DFE806D93FEF9" box="[585,671,332,358]" pageId="13" pageNumber="54" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C72CE023FFB5DC0E4FE3FEA26EBDFE18" blockId="13.[151,671,332,391]" box="[151,433,366,391]" pageId="13" pageNumber="54">
(
<figureCitation id="5FA8FCA6FFB5DC0E4FEBFEA26FE0FE18" box="[159,236,366,391]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="13" pageNumber="54">Figs. 4</figureCitation>
F, G, 7A–K, 11D)
</paragraph>
</subSubSection>
<subSubSection id="8F89B3A8FFB5DC0E4FE3FE7A6EDAFE6C" pageId="13" pageNumber="54" type="materials_examined">
<paragraph id="C72CE023FFB5DC0E4FE3FE7A6EDAFE6C" blockId="13.[151,1437,438,1687]" pageId="13" pageNumber="54">
<materialsCitation id="77FBEA7EFFB5DC0E4FE3FE7A6EDEFE6C" ID-GBIF-Occurrence="1434656674" collectingDate="1970-01-29" collectionCode="MHNG" collectorName="C. Besuchet &amp; Y. Lobl" country="Sri Lanka" location="Ceylan Pidurutalagala" pageId="13" pageNumber="54" specimenCount="1" stateProvince="Central">
<emphasis id="F5E73C31FFB5DC0E4FE3FE7A6E0AFE50" bold="true" box="[151,262,438,463]" pageId="13" pageNumber="54">Material.</emphasis>
Holotype—male,
<collectingCountry id="BF84A0B3FFB5DC0E4EA8FE7B6D7BFE50" box="[476,631,439,463]" name="Sri Lanka" pageId="13" pageNumber="54">SRI LANKA</collectingCountry>
,
<collectingRegion id="05572EC1FFB5DC0E4DF2FE7A6DD4FE50" box="[646,728,438,463]" country="Sri Lanka" name="Central" pageId="13" pageNumber="54">Central</collectingRegion>
prov., &quot;31
<location id="C24CB6F8FFB5DC0E4C2AFE7A6B6CFE50" LSID="urn:lsid:plazi:treatment:4F3A5135FFB5DC0E4FE3FE806B28F908:C24CB6F8FFB5DC0E4C2AFE7A6B6CFE50" box="[862,1120,438,463]" country="Sri Lanka" name="Ceylan Pidurutalagala" pageId="13" pageNumber="54" stateProvince="Central">Ceylan Pidurutalagala</location>
&quot;,
<collectingDate id="A3693F0BFFB5DC0E4B02FE7B6A35FE50" box="[1142,1337,439,463]" pageId="13" pageNumber="54" value="1970-01-29">
<date id="B32DC6E3FFB5DC0E4B02FE7B6A35FE50" box="[1142,1337,439,463]" pageId="13" pageNumber="54" value="1970-01-29">29 January 1970</date>
</collectingDate>
, coll.
<collectorName id="6A6685F5FFB5DC0E4AF7FE7B6E0DFE6C" pageId="13" pageNumber="54">C. Besuchet</collectorName>
,
<collectorName id="6A6685F5FFB5DC0E4E78FE176E53FE6C" box="[268,351,474,499]" pageId="13" pageNumber="54">Y. Löbl</collectorName>
(
<collectionCode id="A18278E6FFB5DC0E4E1AFE176EC6FE6C" box="[366,458,475,499]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="13" pageNumber="54">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8F89B3A8FFB5DC0E4FB3FE336E23FDA4" pageId="13" pageNumber="54" type="etymology">
<paragraph id="C72CE023FFB5DC0E4FB3FE336E23FDA4" blockId="13.[151,1437,438,1687]" pageId="13" pageNumber="54">
<emphasis id="F5E73C31FFB5DC0E4FB3FE336E42FD88" bold="true" box="[199,334,511,535]" pageId="13" pageNumber="54">Etymology.</emphasis>
This species is named in honour of the famous Russian entomologist, Prof. G.Ya. Bey-Bienko (
<date id="B32DC6E3FFB5DC0E4FEBFDEF6E2EFDA4" box="[159,290,547,571]" pageId="13" pageNumber="54" value="1903" valueMax="1971">1903–1971</date>
).
</paragraph>
</subSubSection>
<subSubSection id="8F89B3A8FFB5DC0E4FB3FD8A6B28F908" pageId="13" pageNumber="54" type="description">
<paragraph id="C72CE023FFB5DC0E4FB3FD8A6E23FA94" blockId="13.[151,1437,438,1687]" pageId="13" pageNumber="54">
<emphasis id="F5E73C31FFB5DC0E4FB3FD8A6E55FDC0" bold="true" box="[199,345,582,607]" pageId="13" pageNumber="54">Description.</emphasis>
Male (the
<typeStatus id="18285E81FFB5DC0E4EA6FD8A6D35FDC0" box="[466,569,582,607]" pageId="13" pageNumber="54" type="holotype">holotype</typeStatus>
). General colour yellowish brown, with scattered small brown spots (
<figureCitation id="5FA8FCA6FFB5DC0E4A35FD8A6A88FDC0" box="[1345,1412,582,607]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="13" pageNumber="54">Fig. 4</figureCitation>
G, F); facial part of head darker, dark brown; eyes black; mouthparts and, partly, legs dirty yellow. Surfaces lustrous; antennae with lustrous proximal 8–9 segments, the rest of segments dull; thoracic and abdominal tergites uneven, somewhat rugose, densely covered with small tubercles and longitudinal inflations along caudal margins of thoracal and abdominal tergites (
<figureCitation id="5FA8FCA6FFB5DC0E4D7CFD1A6D5AFD70" box="[520,598,726,751]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="13" pageNumber="54">Figs. 4</figureCitation>
F, 7B); facial part of head moderately punctuated. Head rounded at vertex, epicranial sutures present (
<figureCitation id="5FA8FCA6FFB5DC0E4EA0FD366D2BFC8C" box="[468,551,762,787]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="13" pageNumber="54">Figs. 4</figureCitation>
G, 7A); ocellar spots small, but distinct; a weak transverse hollow located between antennal sockets; distance between eyes 1.6 times eye length; distance between antennal sockets about twice of the scape length (about
<quantity id="006B4DC6FFB5DC0E4D74FC8E6D55FCC4" box="[512,601,834,859]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="54" unit="mm" value="0.6">0.6 mm</quantity>
); approximate length ratio of 3rd–5th segments of maxillary palps 1.1: 1.0: 1.5. Pronotum campaniform, transverse, anterior and lateral margins semicircular, posterior margin nearly straight (
<figureCitation id="5FA8FCA6FFB5DC0E4FEBFC466FFDFC3C" box="[159,241,906,931]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="13" pageNumber="54">Figs. 4</figureCitation>
F, 7B). Meso- and metanotum transverse (
<figureCitation id="5FA8FCA6FFB5DC0E4D96FC466C38FC3C" box="[738,820,906,931]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="13" pageNumber="54">Figs. 4</figureCitation>
F, 7B). Tegmina short, only reaching 2nd abdominal tergite, transformed into wing scales; venation obsolete. Anterior margin of fore femur armed as in the
<typeStatus id="18285E81FFB5DC0E4A39FC636A71FC58" box="[1357,1405,943,967]" pageId="13" pageNumber="54">type</typeStatus>
D
<emphasis id="F5E73C31FFB5DC0E4FE3FC186FD9FC74" box="[151,213,980,1003]" italics="true" pageId="13" pageNumber="54">sensu</emphasis>
<bibRefCitation id="A3029DD2FFB5DC0E4FA8FC1E6E69FC74" author="Roth" box="[220,357,978,1003]" pageId="13" pageNumber="54" refString="Roth, L. M. (2003) Systematics and Phylogeny of Cockroaches (Dictyoptera: Blattaria). Oriental Insects, 37, 1 - 186. http: // dx. doi. org / 10.1080 / 00305316.2003.10417344" type="journal article" year="2003">Roth (2003)</bibRefCitation>
or C
<emphasis id="F5E73C31FFB5DC0E4ED1FC186EEFFC74" box="[421,483,980,1003]" italics="true" pageId="13" pageNumber="54">sensu</emphasis>
<bibRefCitation id="A3029DD2FFB5DC0E4E9FFC1E6DCAFC74" author="Bey-Bienko" box="[491,710,978,1003]" pageId="13" pageNumber="54" refString="Bey-Bienko, G. Ya. (1950) Nasekomye Tarakanovye [Cockroach insects]. Fauna USSR, New Series 40. Leningrad, Moscow, 343 pp." type="book" year="1950">Bey-Bienko (1950)</bibRefCitation>
, all spines, including apical, substituted for uniform row of very short spinules. Fore tibiae moderately thickened distally. Structure of hind tarsi (
<figureCitation id="5FA8FCA6FFB5DC0E4B62FC3A6B55FB90" box="[1046,1113,1014,1039]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">Fig. 7</figureCitation>
C): metatarsus a little shorter than other segments combined, with large euplantula along lower margin, 2nd–4th segments with large euplantulae; tarsal spines absent; all euplantulae with spinules; claws symmetrical and simple; arolium about half of claw length. Abdominal tergites without visible glandular specializations; posterolateral angles of tergites obtuse (
<figureCitation id="5FA8FCA6FFB5DC0E4FEBFB4A6FE1FB00" box="[159,237,1158,1183]" captionStart="FIGURE 4" captionStartId="9.[151,249,1910,1932]" captionTargetBox="[156,1430,440,1893]" captionTargetId="figure@9.[151,1436,431,1905]" captionTargetPageId="9" captionText="FIGURE 4. A, D, F—general view from above; B—head and pronotum from above; C, G—general view from below; Eanterior part of body from below. A – C— Rhabdoblattella alexeevi sp. nov. (male, holotype); D, E— R. euptera sp. nov. (male, holotype); F, G— Placoblatta beybienkoi sp. nov. (male, holotype). In Figs. A, C, G, F abdominal apex removed. Out of scale." httpUri="https://zenodo.org/record/322018/files/figure.png" pageId="13" pageNumber="54">Figs. 4</figureCitation>
F, 7B) Anal plate (tergite X) wide and trapezoidal, caudal margin with weak median incision (
<figureCitation id="5FA8FCA6FFB5DC0E4A61FB4A6A54FB01" box="[1301,1368,1158,1183]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">Fig. 7</figureCitation>
D, E). Cerci shortened, flatten, with segments partly fused (
<figureCitation id="5FA8FCA6FFB5DC0E4C5DFB666C79FB5D" box="[809,885,1194,1219]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">Fig. 7</figureCitation>
D, E). Paraprocts of blaberid-type (
<figureCitation id="5FA8FCA6FFB5DC0E4A5BFB666A70FB5C" box="[1327,1404,1194,1219]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">Fig. 7</figureCitation>
E). Hypandrium nearly symmetrical (
<figureCitation id="5FA8FCA6FFB5DC0E4D63FB026D50FB78" box="[535,604,1230,1255]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">Fig. 7</figureCitation>
F), its caudal margin rounded, with a distinct median incision; styli short and fusiform.
</paragraph>
<paragraph id="C72CE023FFB5DC0E4FB3FADA6BA3FA7C" blockId="13.[151,1437,438,1687]" pageId="13" pageNumber="54">
Male genitalia (
<figureCitation id="5FA8FCA6FFB5DC0E4E0DFADA6EB0FAB0" box="[377,444,1302,1327]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">Fig. 7</figureCitation>
G–K, 11D). Right phallomere (R+N) with caudal part of sclerite R1T subrectangular in shape (
<figureCitation id="5FA8FCA6FFB5DC0E4F98FAF66E3FFACC" box="[236,307,1338,1363]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">Fig. 7</figureCitation>
G, H, 11D,
<emphasis id="F5E73C31FFB5DC0E4EB7FAF76D17FACC" box="[451,539,1339,1363]" italics="true" pageId="13" pageNumber="54">c.p.R1T</emphasis>
), densely covered with bristles; R2 weakly curved; R3 elongated, caudally bifurcated; R4 plate-like; R5 large, fused with sclerite R3. Sclerite L2D (L1) divided into basal and apical parts (
<figureCitation id="5FA8FCA6FFB5DC0E4FEBFA4E6FEBFA04" box="[159,231,1410,1435]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">
Fig.
<date id="B32DC6E3FFB5DC0E4FACFA4F6FEBFA04" box="[216,231,1411,1435]" pageId="13" pageNumber="54">7</date>
</figureCitation>
I); basal part rod-like, weakly widened cranially; apical part rounded, densely covered with recumbent bristles; &quot;dorsal outgrows&quot; flat (
<figureCitation id="5FA8FCA6FFB5DC0E4D71FA6A6D47FA21" box="[517,587,1446,1471]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">
Fig.
<date id="B32DC6E3FFB5DC0E4D48FA6B6D47FA21" box="[572,587,1447,1470]" pageId="13" pageNumber="54">7</date>
</figureCitation>
I,
<emphasis id="F5E73C31FFB5DC0E4D10FA6A6D81FA20" box="[612,653,1446,1471]" italics="true" pageId="13" pageNumber="54">d.o.</emphasis>
). Sclerite L3 (L2d) without basal subsclerite, &quot;folded structure&quot; and bristles (
<figureCitation id="5FA8FCA6FFB5DC0E4F83FA066E36FA7D" box="[247,314,1482,1507]" captionStart="FIGURE 7" captionStartId="14.[151,249,1853,1875]" captionTargetBox="[182,1389,203,1820]" captionTargetId="figure@14.[171,1399,193,1830]" captionTargetPageId="14" captionText="FIGURE 7. Placoblatta beybienkoi sp. nov. (male, holotype). A—facial part of head; B—thorax and 1 – 7 th abdominal tergites from above; C—right hind tarsus from outside; D, E—abdominal apex from above (D) and below, hypandrium and genitalia removed (E); F—hypandrium from below; G, H—right phallomere from below (G) and above (H); I—sclerite L 2 D from above; J, K—sclerite L 3. Dotted area shows membranous parts (not shown in C). Abbreviations: c. p. R 1 T, d. o., R 2, R 3, R 4, R 5, spi. — see text. Scale bar 1 mm: a = A; b = B; c = C; d = D; e = E; f = F; g = G, H; h = I; i = J, K." httpUri="https://zenodo.org/record/322021/files/figure.png" pageId="13" pageNumber="54">Fig. 7</figureCitation>
J, K); groove hge absent. Sclerite L4U (L3d) weakly sclerotized, subtriangular.
</paragraph>
<paragraph id="C72CE023FFB5DC0E4FB3FA226E9CF998" blockId="13.[151,1437,438,1687]" box="[199,400,1518,1543]" pageId="13" pageNumber="54">Female unknown.</paragraph>
<paragraph id="C72CE023FFB5DC0E4FB3F9DF6DE6F9D0" blockId="13.[151,1437,438,1687]" pageId="13" pageNumber="54">
<emphasis id="F5E73C31FFB5DC0E4FB3F9DF6E7AF9B4" bold="true" box="[199,374,1555,1579]" pageId="13" pageNumber="54">Measurements</emphasis>
(mm). Head length: 2.4, width 2.4, pronotum length 3.3, width 5.4, mesonotum length 1.0, tegmina length 3.0, width 1.9, metanotum length 1.1.
</paragraph>
<paragraph id="C72CE023FFB5DC0E4FB3F9966B28F908" blockId="13.[151,1437,438,1687]" pageId="13" pageNumber="54">
<emphasis id="F5E73C31FFB5DC0E4FB3F9966E6DF9EC" bold="true" box="[199,353,1626,1651]" pageId="13" pageNumber="54">Comparison.</emphasis>
The new species differs from the
<typeStatus id="18285E81FFB5DC0E4C75F9976C3DF9EC" box="[769,817,1627,1651]" pageId="13" pageNumber="54">type</typeStatus>
species (
<taxonomicName id="00939BA0FFB5DC0E4CD7F9906B1EF9EC" box="[931,1042,1627,1651]" class="Insecta" family="Blaberidae" genus="Placoblatta" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="54" phylum="Arthropoda" rank="species" species="rugosa">
<emphasis id="F5E73C31FFB5DC0E4CD7F9906CBDF9EC" box="[931,945,1628,1651]" italics="true" pageId="13" pageNumber="54">P</emphasis>
.
<emphasis id="F5E73C31FFB5DC0E4CB0F9906B1EF9EC" box="[964,1042,1628,1651]" italics="true" pageId="13" pageNumber="54">rugosa</emphasis>
</taxonomicName>
) in its smaller size, the unarmed anterior margin of fore femur, the absence of tarsal spines and the larger arolium.
</paragraph>
</subSubSection>
</treatment>
</document>
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