treatments-xml/data/1C/11/87/1C1187FCEB75ED65FF9EFDF3639B83CE.xml

<document id="5284F3AFA575AD650F3B9254C9E0BFFF" ID-CLB-Dataset="299696" ID-DOI="10.5281/zenodo.11512952" ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512952" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1717721037103" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Cueva-Dabkoski, Mollie" docDate="2023" docId="1C1187FCEB75ED65FF9EFDF3639B83CE" docLanguage="en" docName="ProcCalifAcadSci.67.21.493-537.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (21)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Cyclosomus vespertilio Cueva-Dabkoski &amp; Kavanaugh 2023" docType="treatment" docVersion="5" lastPageNumber="525" masterDocId="E028FF84EB68ED45FFE5FFAF6327867B" masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia." masterLastPageNumber="537" masterPageNumber="493" pageNumber="522" updateTime="1720721490288" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="608617F77660242B82F96348C1879766" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C43E3D9D9A109F04A473193659229E8C">
<mods:title id="6BD9198051BB46136962E2BFDFFAA90A">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
</mods:titleInfo>
<mods:name id="C44CAF5552D9CC25DE1594BDA6EE9DCD" type="personal">
<mods:role id="3C84A9E933D622CCB21A97E167719655">
<mods:roleTerm id="6FA5BCDF6575639E189B6D9BC5FF2E68">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="371D88A1D63BB0CE3110C0443A18867C">Kavanaugh, David H.</mods:namePart>
<mods:affiliation id="66AA01B2E0A2316A81D8BEBF1D00C163">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
</mods:name>
<mods:name id="5DB7AC97CFF2B7FBCBD597FCBFD393E8" type="personal">
<mods:role id="EA9C18051AE5144FA32E471B30C0D372">
<mods:roleTerm id="35A2DF0C6E8E8BA1C6BC850BC663CEEC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8AEA2564C725747624DAA0F49F9AA362">Cueva-Dabkoski, Mollie</mods:namePart>
<mods:affiliation id="E383F0271FA17DD9D98D34C9054FAC30">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FF56F586B449BBD5374D27B4030FA4B0">text</mods:typeOfResource>
<mods:relatedItem id="BABE7E546FB12D4B18CE345F9A710FB2" type="host">
<mods:titleInfo id="0BE2BC6EDCBB118D4FA4353BAC93FF40">
<mods:title id="BD35895FE3990E59009DCD3B1442656F">Proceedings of the California Academy of Sciences</mods:title>
</mods:titleInfo>
<mods:part id="8BA1D8C6AAE9D92024279FBF728FAD3A">
<mods:date id="C85B4A583161D0AB2E222E8DDED778B5">2023</mods:date>
<mods:detail id="FF4E2F8A97B2B69415EAA1240298080B" type="pubDate">
<mods:number id="6C9D093B9D888220948F2CC3288DFB14">2023-02-15</mods:number>
</mods:detail>
<mods:detail id="3CBFF9EB87D11872747390E36DDCD584" type="volume">
<mods:number id="AC18E95781C490233AC020129372EBF0">67</mods:number>
</mods:detail>
<mods:detail id="1C291E7F8D3288B248841968E4BB262A" type="issue">
<mods:number id="1DABE57E42360866CE5BAAF9485C51C2">21</mods:number>
</mods:detail>
<mods:extent id="403B6EAEF141458392E9D205D69E959C" unit="page">
<mods:start id="315C74FB3611B9FB999A71D040B3F5EF">493</mods:start>
<mods:end id="5607616CAD29E2F72C696657EAFABE23">537</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9307924348B617DAA89B0C53DBF41450">journal article</mods:classification>
<mods:identifier id="69C8242DB50FF69FB3BEB43C888DF06A" type="CLB-Dataset">299696</mods:identifier>
<mods:identifier id="E8741C735840FDC12B46BA3D15E52F54" type="DOI">10.5281/zenodo.11512952</mods:identifier>
<mods:identifier id="2D721B22BB350DF836A9894ADFCA8056" type="GBIF-Dataset">2b56e3b0-2274-4963-b243-9c991b07072c</mods:identifier>
<mods:identifier id="67AACC8E968CB88260CCA3D6F27C79CC" type="ISSN">0068-547X</mods:identifier>
<mods:identifier id="4E1C29BC501B70CE831BE8C4AD7D37FB" type="Zenodo-Dep">11512952</mods:identifier>
</mods:mods>
<treatment id="1C1187FCEB75ED65FF9EFDF3639B83CE" ID-DOI="http://doi.org/10.5281/zenodo.12726736" ID-GBIF-Taxon="235039993" ID-Zenodo-Dep="12726736" LSID="urn:lsid:plazi:treatment:1C1187FCEB75ED65FF9EFDF3639B83CE" httpUri="http://treatment.plazi.org/id/1C1187FCEB75ED65FF9EFDF3639B83CE" lastPageId="32" lastPageNumber="525" pageId="29" pageNumber="522">
<subSubSection id="DCA26561EB75ED58FF9EFDF3606F840F" box="[123,840,604,628]" pageId="29" pageNumber="522" type="nomenclature">
<paragraph id="940736EAEB75ED58FF9EFDF3606F840F" blockId="29.[123,1174,604,1725]" box="[123,840,604,628]" pageId="29" pageNumber="522">
<heading id="CF4F8186EB75ED58FF9EFDF3606F840F" box="[123,840,604,628]" fontSize="10" level="2" pageId="29" pageNumber="522" reason="4">
<emphasis id="A6CCEAF8EB75ED58FF9EFDF3606F840F" bold="true" box="[123,840,604,628]" pageId="29" pageNumber="522">
<taxonomicName id="53B84D69EB75ED58FF9EFDF361C1840F" authority="Cueva-Dabkoski &amp; Kavanaugh" authorityName="Cueva-Dabkoski &amp; Kavanaugh" authorityYear="2023" box="[123,742,604,628]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="vespertilio" status="sp. nov.">
<emphasis id="A6CCEAF8EB75ED58FF9EFDF36250840F" bold="true" box="[123,375,604,628]" italics="true" pageId="29" pageNumber="522">Cyclosomus vespertilio</emphasis>
Cueva-Dabkoski &amp; Kavanaugh
</taxonomicName>
, sp. nov.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB75ED58FF9EFDD362F384E9" box="[123,468,636,658]" pageId="29" pageNumber="522" type="description">
<paragraph id="940736EAEB75ED58FF9EFDD362F384E9" blockId="29.[123,1174,604,1725]" box="[123,468,636,658]" pageId="29" pageNumber="522">
<figureCitation id="0C832A6FEB75ED58FF9EFDD363C484E9" box="[123,227,636,658]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="29" pageNumber="522">Figures 13</figureCitation>
, 
<figureCitation id="0C832A6FEB75ED58FF0BFDD3623784E9" box="[238,272,636,658]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="29" pageNumber="522">14I</figureCitation>
, 
<figureCitation id="0C832A6FEB75ED58FEFFFDD3626284E9" box="[282,325,636,658]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="29" pageNumber="522">15D</figureCitation>
, 
<figureCitation id="0C832A6FEB75ED58FEB5FDD3625B84E9" box="[336,380,636,658]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="29" pageNumber="522">16H</figureCitation>
, 
<figureCitation id="0C832A6FEB75ED58FE63FDD3629684E9" box="[390,433,636,658]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="29" pageNumber="522">17H</figureCitation>
, 
<figureCitation id="0C832A6FEB75ED58FE59FDD362F384E9" box="[444,468,636,658]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="29" pageNumber="522">22</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB75ED58FF4EFD0D61F28360" pageId="29" pageNumber="522" type="materials_examined">
<paragraph id="940736EAEB75ED58FF4EFD0D604C8280" blockId="29.[123,1174,604,1725]" pageId="29" pageNumber="522">
<emphasis id="A6CCEAF8EB75ED58FF4EFD0D626D84C2" bold="true" box="[171,330,674,697]" pageId="29" pageNumber="522">Type material</emphasis>
. 
<typeStatus id="4B038848EB75ED58FEBCFD0E62CE84C2" box="[345,489,673,697]" pageId="29" pageNumber="522" type="holotype">HOLOTYPE</typeStatus>
, a male, in 
<collectionCode id="F2A9AE2FEB75ED58FD83FD0D619D84C2" box="[614,698,674,697]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="522" type="Museum">NMNH</collectionCode>
, labeled: “ 
<collectingCountry id="ECAF767AEB75ED58FCCEFD0E60A284C2" box="[811,901,673,697]" name="Nepal" pageId="29" pageNumber="522">NEPAL</collectingCountry>
: Royal Chitwan National Park, black light 
<quantity id="53409B0FEB75ED58FECAFD6E624084A2" box="[303,359,705,729]" metricMagnitude="2" metricUnit="m" metricValue="2.49936" pageId="29" pageNumber="522" unit="ft" value="820.0">820ft</quantity>
<date id="E006102AEB75ED58FE88FD6E62CD84A2" box="[365,490,705,729]" pageId="29" pageNumber="522" value="1977-10-31">31 Oct1977</date>
Gary F. Hevel”/ “ 
<typeStatus id="4B038848EB75ED58FD43FD6E601284A2" box="[678,821,705,729]" pageId="29" pageNumber="522" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="53B84D69EB75ED58FCDEFD6E671484A2" authority="Kavanaugh &amp; Cueva-Dabkoski, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski" authorityYear="2023" box="[827,1075,705,729]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="vespertilio" status="sp. nov.">
<emphasis id="A6CCEAF8EB75ED58FCDEFD6E671484A2" box="[827,1075,705,729]" italics="true" pageId="29" pageNumber="522">Cyclosomus vespertilio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDFF5783EB75ED58FBDCFD6D675684A2" box="[1081,1137,706,729]" pageId="29" pageNumber="522" rank="species">sp. n.</taxonomicNameLabel>
M. Cueva-Dabkoski and D.H. 
<bibRefCitation id="F0294B1BEB75ED58FE7EFD4E61758482" author="KAVANAUGH, D. H. &amp; M. CUEVA-DABKOSKI &amp; H. B. LIANG" box="[411,594,737,761]" pageId="29" pageNumber="522" pagination="451 - 491" refId="ref23619" refString="KAVANAUGH, D. H., M. CUEVA-DABKOSKI, AND H. B. LIANG. 2023. Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with descriptions of two new species. Proceedings of the California Academy of Sciences (Series 4) 67: 451 - 491." type="journal article" year="2023">Kavanaugh 2023</bibRefCitation>
” [red label]. 
<collectionCode id="F2A9AE2FEB75ED58FD39FD4E61C98482" box="[732,750,737,761]" pageId="29" pageNumber="522">A</collectionCode>
total of 
<specimenCount id="82BEFD63EB75ED58FCA3FD4E60F68481" box="[838,977,737,762]" count="44" pageId="29" pageNumber="522" type="generic" typeStatus="paratype">44 paratypes</specimenCount>
: 
<specimenCount id="82BEFD63EB75ED58FC3EFD4D671B8482" box="[987,1084,737,761]" count="1" pageId="29" pageNumber="522" type="male">one male</specimenCount>
and 
<specimenCount id="82BEFD63EB75ED58FB8AFD4D63E18562" count="1" pageId="29" pageNumber="522" type="female">one female</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FF09FCAE62078562" box="[236,288,769,793]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="522" type="Museum">CAS</collectionCode>
and 
<collectionCode id="F2A9AE2FEB75ED58FEB1FCAD628F8562" box="[340,424,770,793]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="522" type="Museum">NMNH</collectionCode>
, same label data as 
<typeStatus id="4B038848EB75ED58FD99FCAE61FA8562" box="[636,733,769,793]" pageId="29" pageNumber="522" type="holotype">holotype</typeStatus>
; 
<specimenCount id="82BEFD63EB75ED58FD0DFCAD606C8562" box="[744,843,769,793]" count="1" pageId="29" pageNumber="522" type="male">one male</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FC97FCAD60FD8562" box="[882,986,770,793]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
, labeled: “ 
<collectingCountry id="ECAF767AEB75ED58FBACFCAE67A48562" box="[1097,1155,769,793]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
”/ “Bowring, 63.47*”/ “ 
<collectionCode id="F2A9AE2FEB75ED58FEBBFC8E62E28542" box="[350,453,801,825]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
010795925”; 
<specimenCount id="82BEFD63EB75ED58FDB8FC8D61F18542" box="[605,726,801,825]" count="1" pageId="29" pageNumber="522" type="female">one female</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FD19FC8D60438542" box="[764,868,802,825]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
, labeled: “ 
<collectingCountry id="ECAF767AEB75ED58FC36FC8E672A8542" box="[979,1037,801,825]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
”/ “Bowring, 63.47*”/ “H.E.Andrewes Coll. B.M.1945-97.”/ “Ex coll. Brit. Mus.”/ “ 
<taxonomicName id="53B84D69EB75ED58FC6AFCEE621A8501" authority="Motch. Compared" authorityName="Motch. Compared" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="marginatus">Cyclosomus marginatus Motch. Compared</taxonomicName>
with type H.E. 
<collectionCode id="F2A9AE2FEB75ED58FE01FCCD62DC8501" box="[484,507,866,890]" pageId="29" pageNumber="522">A</collectionCode>
.”; 
<specimenCount id="82BEFD63EB75ED58FDF2FCCD615C8501" box="[535,635,866,890]" count="1" pageId="29" pageNumber="522" type="male">one male</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FD46FCCD602C8502" box="[675,779,866,889]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
, labeled: “Indian Orient”/ “Fry Coll. 1905.100.”/ “ 
<collectionCode id="F2A9AE2FEB75ED58FEECFC2D625785E1" box="[265,368,898,922]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
010796097”; 
<specimenCount id="82BEFD63EB75ED58FDECFC2C614B85E1" box="[521,620,898,922]" count="1" pageId="29" pageNumber="522" type="male">one male</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FD76FC2C61C085E1" box="[659,743,899,922]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="522" type="Museum">NMNH</collectionCode>
, labeled: “ 
<collectingCountry id="ECAF767AEB75ED58FCB3FC2D608485E1" box="[854,931,898,922]" name="India" pageId="29" pageNumber="522">INDIA</collectingCountry>
New 
<collectingRegion id="567CF808EB75ED58FC06FC2D670785E1" box="[995,1056,898,922]" country="India" name="Delhi" pageId="29" pageNumber="522">Delhi</collectingRegion>
”/ “ 
<date id="E006102AEB75ED58FBA7FC2D639785C1" pageId="29" pageNumber="522" value="1967-07-14">14-VII-1967</date>
KEGibson light trap”; 
<specimenCount id="82BEFD63EB75ED58FE48FC0C610085C1" box="[429,551,930,954]" count="1" pageId="29" pageNumber="522" type="female">one female</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FDAAFC0C619085C1" box="[591,695,931,954]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
, labeled: “ 
<collectingCountry id="ECAF767AEB75ED58FCCDFC0D604685C1" box="[808,865,930,954]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
77.15 K.” [handwritten]; 
<specimenCount id="82BEFD63EB75ED58FB9EFC0D639E85A1" count="12" pageId="29" pageNumber="522" type="male">12 males</specimenCount>
and 
<specimenCount id="82BEFD63EB75ED58FF12FC6D625385A1" box="[247,372,962,986]" count="22" pageId="29" pageNumber="522" type="female">22 females</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FE41FC6C612E85A1" box="[420,521,963,986]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
and 
<collectionCode id="F2A9AE2FEB75ED58FDADFC6D615985A1" box="[584,638,962,986]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="29" pageNumber="522" type="Museum">CAS</collectionCode>
, labeled: “ 
<collectingCountry id="ECAF767AEB75ED58FD1CFC6D601285A1" box="[761,821,962,986]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
. Nevinson Coll. 1918-14”; 
<specimenCount id="82BEFD63EB75ED58FB8AFC6C63E18581" count="1" pageId="29" pageNumber="522" type="female">one female</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FF09FC4C62738581" box="[236,340,995,1018]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
, labeled: “ 
<collectingCountry id="ECAF767AEB75ED58FE26FC4D62D88581" box="[451,511,994,1018]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
. Nevinson Coll. 1918-14”/ “ 
<taxonomicName id="53B84D69EB75ED58FCC8FC4D67A38581" authority="Motch." authorityName="Motch." box="[813,1156,994,1018]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="marginatus">Cyclosomus marginatus Motch.</taxonomicName>
”/ “NNHMUK 010796368”; 
<specimenCount id="82BEFD63EB75ED58FE7CFBAC62DA8261" box="[409,509,1026,1050]" count="1" pageId="29" pageNumber="522" type="male">one male</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FDC3FBAC61A98261" box="[550,654,1027,1050]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
, labelled: “ 
<collectingCountry id="ECAF767AEB75ED58FCE2FBAD60648261" box="[775,835,1026,1050]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
. Nevinson Coll. 1918-14”/ “Ex coll. Brit. Mus.”/ “H.E.Andrewes Coll. B.M.1945-97.”/ “ 
<taxonomicName id="53B84D69EB75ED58FD0EFB8D67398241" authority="Wied." authorityName="Wied." box="[747,1054,1058,1082]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="suturalis">Cyclosomus suturalis Wied.</taxonomicName>
(see back) Compared with type H.E. 
<collectionCode id="F2A9AE2FEB75ED58FE43FBED629A8221" box="[422,445,1090,1114]" pageId="29" pageNumber="522">A</collectionCode>
.” [back labeled: “The median band in type is very narrow”]/ “NNHMUK 010796362”; 
<specimenCount id="82BEFD63EB75ED58FE73FBCC61298200" box="[406,526,1123,1147]" count="1" pageId="29" pageNumber="522" type="female">one female</specimenCount>
, in 
<collectionCode id="F2A9AE2FEB75ED58FDD6FBCC61BF8201" box="[563,664,1123,1146]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
labeled: “ 
<collectingCountry id="ECAF767AEB75ED58FCE7FBCC601C8200" box="[770,827,1123,1147]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
<collectingRegion id="567CF808EB75ED58FCA4FBCC605F8200" box="[833,888,1123,1147]" country="India" name="Delhi" pageId="29" pageNumber="522">Dilhi</collectingRegion>
[sic]”/ “39155” [= Moradabad, based on 
<collectionCode id="F2A9AE2FEB75ED58FECBFB2B62B482E0" box="[302,403,1156,1179]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="29" pageNumber="522" type="Museum">NHMUK</collectionCode>
accession records (M. Barclay, personal communication)]/ Fry Coll. 1906.100.”/ 
<taxonomicName id="53B84D69EB75ED58FEE9FB0C614C82C0" authority="Mots." authorityName="Mots." box="[268,619,1187,1211]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="marginatus">Cyclosomus marginatus Mots.</taxonomicName>
<collectingCountry id="ECAF767AEB75ED58FD9EFB0C619482C0" box="[635,691,1187,1211]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
<taxonomicName id="53B84D69EB75ED58FD21FB0C607B82C0" box="[708,860,1187,1211]" class="Insecta" family="Carabidae" genus="Omophron" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="variety" variety="major">Or. v. major</taxonomicName>
” [handwritten]/ “NMHUK 0100795973”. Each 
<typeStatus id="4B038848EB75ED58FEB2FB6B629482A7" box="[343,435,1220,1244]" pageId="29" pageNumber="522" type="paratype">paratype</typeStatus>
also bears the following label: “ 
<typeStatus id="4B038848EB75ED58FCEBFB6C60B282A0" box="[782,917,1219,1243]" pageId="29" pageNumber="522" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="53B84D69EB75ED58FC78FB6C67B182A0" authority="Kavanaugh &amp; Cueva-Dabkoski, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski" authorityYear="2023" box="[925,1174,1219,1243]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="vespertilio" status="sp. nov.">
<emphasis id="A6CCEAF8EB75ED58FC78FB6C67B182A0" box="[925,1174,1219,1243]" italics="true" pageId="29" pageNumber="522">Cyclosomus vespertilio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDFF5783EB75ED58FF9EFB4B63928280" box="[123,181,1252,1275]" pageId="29" pageNumber="522" rank="species">sp. n.</taxonomicNameLabel>
M. Cueva-Dabkoski and D.H. 
<bibRefCitation id="F0294B1BEB75ED58FDE0FB4C619A8280" author="KAVANAUGH, D. H. &amp; M. CUEVA-DABKOSKI &amp; H. B. LIANG" box="[517,701,1251,1275]" pageId="29" pageNumber="522" pagination="451 - 491" refId="ref23619" refString="KAVANAUGH, D. H., M. CUEVA-DABKOSKI, AND H. B. LIANG. 2023. Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with descriptions of two new species. Proceedings of the California Academy of Sciences (Series 4) 67: 451 - 491." type="journal article" year="2023">Kavanaugh 2023</bibRefCitation>
” [yellow label].
</paragraph>
<paragraph id="940736EAEB75ED58FF4EFAAB61F28360" blockId="29.[123,1174,604,1725]" box="[171,725,1283,1307]" pageId="29" pageNumber="522">
<emphasis id="A6CCEAF8EB75ED58FF4EFAAB621A8360" bold="true" box="[171,317,1284,1307]" pageId="29" pageNumber="522">
<typeStatus id="4B038848EB75ED58FF4EFAAB63C48360" box="[171,227,1284,1307]" pageId="29" pageNumber="522">Type</typeStatus>
locality
</emphasis>
. 
<collectingCountry id="ECAF767AEB75ED58FEAEFAAC62AA8360" box="[331,397,1283,1307]" name="Nepal" pageId="29" pageNumber="522">Nepal</collectingCountry>
, 
<location id="91676031EB75ED58FE7DFAAC61F28360" LSID="urn:lsid:plazi:treatment:1C1187FCEB75ED65FF9EFDF3639B83CE:91676031EB75ED58FE7DFAAC61F28360" box="[408,725,1283,1307]" country="Nepal" name="Royal Chitwan National Park" pageId="29" pageNumber="522">Royal Chitwan National Park</location>
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB75ED58FF4EFA8B60058307" pageId="29" pageNumber="522" type="etymology">
<paragraph id="940736EAEB75ED58FF4EFA8B60058307" blockId="29.[123,1174,604,1725]" pageId="29" pageNumber="522">
<emphasis id="A6CCEAF8EB75ED58FF4EFA8B620C8340" bold="true" box="[171,299,1316,1339]" pageId="29" pageNumber="522">Etymology.</emphasis>
The species epithet, 
<taxonomicName id="53B84D69EB75ED58FDFFFA8C61AC8340" authorityName="Kavanaugh &amp; Cueva-Dabkoski" authorityYear="2023" box="[538,651,1315,1339]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="vespertilio">
<emphasis id="A6CCEAF8EB75ED58FDFFFA8C61AC8340" box="[538,651,1315,1339]" italics="true" pageId="29" pageNumber="522">vespertilio</emphasis>
</taxonomicName>
, is a Latin word for bat, here used as a noun in apposition. It refers to the shape of the middle transverse dark band of the joined elytra, which resembles a black bat with its wings spread in most specimens.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB75ED5AFF4EFA2A625284ED" lastPageId="31" lastPageNumber="524" pageId="29" pageNumber="522" type="diagnosis">
<paragraph id="940736EAEB75ED5AFF4EFA2A61D787CF" blockId="29.[123,1174,604,1725]" lastBlockId="31.[123,1174,219,1725]" lastPageId="31" lastPageNumber="524" pageId="29" pageNumber="522">
<emphasis id="A6CCEAF8EB75ED58FF4EFA2A623883E7" bold="true" box="[171,287,1413,1436]" pageId="29" pageNumber="522">Diagnosis.</emphasis>
Adults of 
<taxonomicName id="53B84D69EB75ED58FE7BFA2A611483E7" authorityName="Kavanaugh &amp; Cueva-Dabkoski" authorityYear="2023" box="[414,563,1412,1436]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="vespertilio">
<emphasis id="A6CCEAF8EB75ED58FE7BFA2A611483E7" box="[414,563,1412,1436]" italics="true" pageId="29" pageNumber="522">C. vespertilio</emphasis>
</taxonomicName>
can be distinguished from those of other 
<taxonomicName id="53B84D69EB75ED58FBF1FA2B63EF83C7" pageId="29" pageNumber="522">
<emphasis id="A6CCEAF8EB75ED58FBF1FA2B67B283E7" box="[1044,1173,1412,1436]" italics="true" pageId="29" pageNumber="522">Cyclosomus</emphasis>
species
</taxonomicName>
in Asia by the following combination of character states: Body size medium for genus, BL males = 
<quantity id="53409B0FEB75ED58FF33FA6B624883A7" box="[214,367,1476,1500]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.4" metricValueMin="6.6" pageId="29" pageNumber="522" unit="mm" value="7.0" valueMax="7.4" valueMin="6.6">6.6 to 7.4 mm</quantity>
, females = 
<quantity id="53409B0FEB75ED58FE0FFA6B61A483A7" box="[490,643,1476,1500]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.6" metricValueMin="6.5" pageId="29" pageNumber="522" unit="mm" value="7.05" valueMax="7.6" valueMin="6.5">6.5 to 7.6 mm</quantity>
; body form (
<figureCitation id="0C832A6FEB75ED58FCE8FA6B608883A7" box="[781,943,1476,1500]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="29" pageNumber="522">Fig. 13A, C, D</figureCitation>
) roundly ovoid (ratio BL/EW = 1.36 to 1.41; ratio EL/EW = 0.88 to 0.93), with elytra widest at or near mid-length; dorsal surface with distinctly contrasting pale and dark areas; pronotum (
<figureCitation id="0C832A6FEB75ED58FC9AF9AB60FF8067" box="[895,984,1540,1564]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="29" pageNumber="522">Fig. 14I</figureCitation>
) relatively broad (ratio PWM/PL = 2.79 to 2.93) and more broadened basally (ratio PWM/PWA = 1.63 to 1.75), with disc rufous to piceous, lateral pale bands broad and more or less well-defined, anterior angles broad, lateral margins evenly arcuate, not or faintly sinuate near anterior angles; free apex of prosternal intercoxal process short (
<figureCitation id="0C832A6FEB75ED58FE12F92A617D80E6" box="[503,602,1669,1693]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="29" pageNumber="522">Fig. 15D</figureCitation>
); elytra pale yellowish-brown with basal, middle, and longitudinal dark bands present and dark reddish brown to black, preapical dark spot absent, middle transverse dark band varied, moderately thick in most specimens (
<figureCitation id="0C832A6FEB77ED5AFC97FF7460F28688" box="[882,981,219,243]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13A</figureCitation>
), markedly thickened (
<figureCitation id="0C832A6FEB77ED5AFF58FF5462398768" box="[189,286,251,275]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13C</figureCitation>
) or narrowed (
<figureCitation id="0C832A6FEB77ED5AFE26FF5461028768" box="[451,549,251,275]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13D</figureCitation>
) (much as in 
<taxonomicName id="53B84D69EB77ED5AFD5AFF53607E8768" box="[703,857,252,275]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="A6CCEAF8EB77ED5AFD5AFF53607E8768" box="[703,857,252,275]" italics="true" pageId="31" pageNumber="524">C. marginatus</emphasis>
</taxonomicName>
), in a few specimens; elytral striae moderately impressed, elytral intervals flat or nearly so; elytral epipleura with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; median lobe of male genitalia with shaft distinctly thicker and more arcuate, ventral curvature distinctly curved throughout in lateral aspect (
<figureCitation id="0C832A6FEB77ED5AFE86FED362EF87EF" box="[355,456,380,404]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="31" pageNumber="524">Fig. 16H</figureCitation>
), apical lamella smoothly rounded and slightly elongate in dorsal aspect (
<figureCitation id="0C832A6FEB77ED5AFF28FE33621787CF" box="[205,304,412,436]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="31" pageNumber="524">Fig. 17H</figureCitation>
); specimen from northern 
<collectingCountry id="ECAF767AEB77ED5AFDABFE3361A287CF" box="[590,645,412,436]" name="India" pageId="31" pageNumber="524">India</collectingCountry>
or 
<collectingCountry id="ECAF767AEB77ED5AFD4CFE3361CB87CF" box="[681,748,412,436]" name="Nepal" pageId="31" pageNumber="524">Nepal</collectingCountry>
.
</paragraph>
<caption id="C0C76662EB76ED5BFF4EF98A60F28029" ID-DOI="http://doi.org/10.5281/zenodo.11512984" ID-Zenodo-Dep="11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="30" pageNumber="523" startId="30.[171,183,1573,1592]" targetBox="[134,1162,233,1556]" targetPageId="30" targetType="figure">
<paragraph id="940736EAEB76ED5BFF4EF98A67B38043" blockId="30.[123,1172,1572,1618]" box="[171,1172,1572,1592]" pageId="30" pageNumber="523">
<smallCapsWord id="92E1A036EB76ED5BFF4EF98A63CF804C" baselines="1587,1587" box="[171,232,1573,1592]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="30" pageNumber="523">FIGURE</smallCapsWord>
13. 
<taxonomicName id="53B84D69EB76ED5BFEF4F98B61D8804C" authority="Cueva-Dabkoski and Kavanaugh" authorityName="Cueva-Dabkoski and Kavanaugh" authorityYear="2023" box="[273,767,1572,1591]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="523" phylum="Arthropoda" rank="species" species="vespertilio" status="sp. nov.">
<emphasis id="A6CCEAF8EB76ED5BFEF4F98B62FE804C" box="[273,473,1572,1591]" italics="true" pageId="30" pageNumber="523">Cyclosomus vespertilio</emphasis>
Cueva-Dabkoski and Kavanaugh
</taxonomicName>
<taxonomicNameLabel id="BDFF5783EB76ED5BFCE3F98A606C8043" box="[774,843,1573,1592]" pageId="30" pageNumber="523" rank="species">sp. nov.</taxonomicNameLabel>
A. Holotype male, dorsal habitus; B.
</paragraph>
<paragraph id="940736EAEB76ED5BFF9EF99160F28029" blockId="30.[123,1172,1572,1618]" box="[123,981,1598,1618]" pageId="30" pageNumber="523">Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm.</paragraph>
</caption>
<paragraph id="940736EAEB77ED5AFF4EFE13625284ED" blockId="31.[123,1174,219,1725]" pageId="31" pageNumber="524">
Specimens of 
<taxonomicName id="53B84D69EB77ED5AFEA0FE1262F087AF" authorityName="Cueva-Dabkoski and Kavanaugh" authorityYear="2023" box="[325,471,444,468]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="vespertilio">
<emphasis id="A6CCEAF8EB77ED5AFEA0FE1262F087AF" box="[325,471,444,468]" italics="true" pageId="31" pageNumber="524">C. vespertilio</emphasis>
</taxonomicName>
are most similar to those of 
<taxonomicName id="53B84D69EB77ED5AFCF6FE12605887AF" box="[787,895,445,468]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="inustus">
<emphasis id="A6CCEAF8EB77ED5AFCF6FE12605887AF" box="[787,895,445,468]" italics="true" pageId="31" pageNumber="524">C. inustus</emphasis>
</taxonomicName>
and 
<taxonomicName id="53B84D69EB77ED5AFC53FE12677787AF" box="[950,1104,445,468]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="A6CCEAF8EB77ED5AFC53FE12677787AF" box="[950,1104,445,468]" italics="true" pageId="31" pageNumber="524">C. marginatus</emphasis>
</taxonomicName>
. Most specimens of 
<taxonomicName id="53B84D69EB77ED5AFEE8FE7262BA878F" authorityName="Cueva-Dabkoski and Kavanaugh" authorityYear="2023" box="[269,413,476,500]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="vespertilio">
<emphasis id="A6CCEAF8EB77ED5AFEE8FE7262BA878F" box="[269,413,476,500]" italics="true" pageId="31" pageNumber="524">C. vespertilio</emphasis>
</taxonomicName>
have the middle dark band of the elytral color pattern darker and thick-er than those of the other two species, with its form in most specimens as in 
<figureCitation id="0C832A6FEB77ED5AFC52FE526732846E" box="[951,1045,509,533]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13A</figureCitation>
or intermediate between that form and the extreme as shown in 
<figureCitation id="0C832A6FEB77ED5AFD5FFDB2603D844E" box="[698,794,541,565]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13C</figureCitation>
. The extreme minimally developed form shown in 
<figureCitation id="0C832A6FEB77ED5AFEFBFD92625A842E" box="[286,381,573,597]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13D</figureCitation>
was found in only 
<specimenCount id="82BEFD63EB77ED5AFDA9FD9161CB842E" box="[588,748,573,597]" count="2" pageId="31" pageNumber="524" type="generic">two specimens</specimenCount>
, and these specimens resemble those of 
<taxonomicName id="53B84D69EB77ED5AFF9EFDF1623F840E" box="[123,280,606,629]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="A6CCEAF8EB77ED5AFF9EFDF1623F840E" box="[123,280,606,629]" italics="true" pageId="31" pageNumber="524">C. marginatus</emphasis>
</taxonomicName>
. Refer to the key and Diagnosis sections for 
<taxonomicName id="53B84D69EB77ED5AFCC7FDF160B5840E" box="[802,914,606,629]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="inustus">
<emphasis id="A6CCEAF8EB77ED5AFCC7FDF160B5840E" box="[802,914,606,629]" italics="true" pageId="31" pageNumber="524">C. inustus</emphasis>
</taxonomicName>
and 
<taxonomicName id="53B84D69EB77ED5AFC2BFDF1674C840E" box="[974,1131,606,629]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="A6CCEAF8EB77ED5AFC2BFDF1674C840E" box="[974,1131,606,629]" italics="true" pageId="31" pageNumber="524">C. marginatus</emphasis>
</taxonomicName>
for distinguishing features.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB77ED65FF4EFD30639B83CE" lastPageId="32" lastPageNumber="525" pageId="31" pageNumber="524" type="description">
<paragraph id="940736EAEB77ED5AFF4EFD306155848D" blockId="31.[123,1174,219,1725]" pageId="31" pageNumber="524">
<emphasis id="A6CCEAF8EB77ED5AFF4EFD30621184CD" bold="true" box="[171,310,671,694]" pageId="31" pageNumber="524">Description.</emphasis>
Size medium for genus, BL males = 
<quantity id="53409B0FEB77ED5AFD31FD31605584CD" box="[724,882,670,694]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.4" metricValueMin="6.6" pageId="31" pageNumber="524" unit="mm" value="7.0" valueMax="7.4" valueMin="6.6">6.6 to 7.4 mm</quantity>
, females = 
<quantity id="53409B0FEB77ED5AFC17FD3167B684CD" box="[1010,1169,670,694]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.6" metricValueMin="6.5" pageId="31" pageNumber="524" unit="mm" value="7.05" valueMax="7.6" valueMin="6.5">6.5 to 7.6 mm</quantity>
; body form (
<figureCitation id="0C832A6FEB77ED5AFF18FD11628484AD" box="[253,419,702,726]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13A, C, D</figureCitation>
) roundly ovoid (ratio BL/EW = 1.36 to 1.41; ratio EL/EW = 0.88 to 0.93), with elytra widest at or near mid-length.
</paragraph>
<paragraph id="940736EAEB77ED5AFF4EFD51674A8263" blockId="31.[123,1174,219,1725]" pageId="31" pageNumber="524">
Color. Head rufous to black, clypeus and labrum slightly lighter in dark specimens, venter rufous; antennae, mandibles, maxillae, and maxillary and labial palpi pale yellow-tan to rufous. Pronotum with disc rufous to piceous, lateral pale bands yellow-tan, broad and more or less well-defined. Elytra mainly yellow-brown, with piceous to black markings; elytral base, scutellum, and longitudinal dark band on elytral interval 1 dark reddish brown to black, dark basal band slender, piceous to black, extended laterally to interval 5; middle transverse dark band piceous or black, varied in form, moderately thick in most specimens (
<figureCitation id="0C832A6FEB77ED5AFD6FFC6F61CB85A3" box="[650,748,960,984]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13A</figureCitation>
), markedly thickened (
<figureCitation id="0C832A6FEB77ED5AFC04FC6F671885A3" box="[993,1087,960,984]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13C</figureCitation>
) or narrowed (
<figureCitation id="0C832A6FEB77ED5AFF29FC4F620B8583" box="[204,300,992,1016]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="31" pageNumber="524">Fig. 13D</figureCitation>
) (much as in 
<taxonomicName id="53B84D69EB77ED5AFE59FC4E61738583" box="[444,596,993,1016]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="524" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="A6CCEAF8EB77ED5AFE59FC4E61738583" box="[444,596,993,1016]" italics="true" pageId="31" pageNumber="524">C. marginatus</emphasis>
</taxonomicName>
), in a few specimens; preapical dark spot absent. Venter rufous, proepipleura and elytral epipleura pale yellow-tan. Legs pale, yellow-tan to rufous.
</paragraph>
<paragraph id="940736EAEB77ED5AFF4EFB8F600282E2" blockId="31.[123,1174,219,1725]" pageId="31" pageNumber="524">Reflection, luster, and microsculpture. Dorsum and venter without metallic reflection. Head and pronotum dull, with microsculpture moderately impressed and comprised of isodiametric meshes; elytra slightly shiny with isodiametric meshes slightly less deeply impressed; venter with isodiametric to slightly transverse meshes shallowly impressed,</paragraph>
<paragraph id="940736EAEB77ED5AFF4EFB0E626A8361" blockId="31.[123,1174,219,1725]" pageId="31" pageNumber="524">Head. Eyes large, hemispheric; antennae slightly short, extended only to basal one-fifth of elytra; clypeus bisetose; labrum with apical margin deeply emarginate, three pairs of setae present; mentum asetose, with a broad, apically emarginate medial tooth; submentum anteriorly with a single pair of setae.</paragraph>
<paragraph id="940736EAEB77ED5AFF4EFA8D624E8007" blockId="31.[123,1174,219,1725]" pageId="31" pageNumber="524">
Prothorax. Pronotum (
<figureCitation id="0C832A6FEB77ED5AFE79FA8D62D38341" box="[412,500,1314,1338]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="31" pageNumber="524">Fig. 14I</figureCitation>
) trapezoidal, almost as wide as elytra at humeral angles, relatively broad (ratio PWM/PL = 2.79 to 2.93) and more broadened basally (ratio PWM/PWA = 1.63 to 1.75), anterior angles broad, rectangular or slightly acute, narrowly rounded apically; anterior margin smoothly concave between anterior angles; lateral margins gently arcuate, not or faintly sinuate near anterior angles; basal margin slightly bisinuate, faintly lobate medially; anterior and basal margination thin but distinct laterally, obsolete medially; lateral margination very slender and finely impressed; midlateral setae present, inserted just medial to lateral margination and at about one-third the distance between the anterior and basal angles along the lateral margin; basolateral setae present, inserted on lateral edge of pronotum and just anterior to posterior angles. Prosternal intercoxal process moderately long, lanceolate, with complete margination, free apex of process short (as in 
<figureCitation id="0C832A6FEB77ED5AFF1FF9CB627A8007" box="[250,349,1636,1660]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="31" pageNumber="524">Fig. 15D</figureCitation>
).
</paragraph>
<paragraph id="940736EAEB77ED65FF4EF92B67688728" blockId="31.[123,1174,219,1725]" lastBlockId="32.[123,1174,219,1461]" lastPageId="32" lastPageNumber="525" pageId="31" pageNumber="524">Elytra. Broadly ovate, slightly shorter than wide (ratio EL/EW = 0.88 to 0.93), broadly rounded apically, humeri slightly obtuse, angulate; elytral striae moderately impressed throughout; elytral intervals flat or nearly so, smooth; parascutellar setiferous pore present at base of interval 1 near junction of striae 1 and 2; two discal setiferous pores present on interval 3 adjacent to stria 2, one inserted just posterior to elytral mid-length and the other inserted near apical one-third; apical seta present, inserted near apex of interval 3; umbilicate series comprised of 11 to 13 setae.</paragraph>
<paragraph id="940736EAEB48ED65FF4EFEF361038788" blockId="32.[123,1174,219,1461]" pageId="32" pageNumber="525">
Legs. Males and females with front tarsi similar in shape and width (as in 
<figureCitation id="0C832A6FEB48ED65FC22FEF4673C8708" box="[967,1051,347,371]" captionStart="FIGURE" captionStartId="2.[171,183,897,916]" captionTargetBox="[120,1176,216,878]" captionTargetId="figure-193@2.[120,1176,216,878]" captionTargetPageId="2" captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11512958" httpUri="https://zenodo.org/record/11512958/files/figure.png" pageId="32" pageNumber="525">Fig. 2C</figureCitation>
), but males with tarsomeres 1 to 3 with two rows of adhesive squamosetae ventrally (as in 
<figureCitation id="0C832A6FEB48ED65FC3CFED4670A87E8" box="[985,1069,379,403]" captionStart="FIGURE" captionStartId="2.[171,183,897,916]" captionTargetBox="[120,1176,216,878]" captionTargetId="figure-193@2.[120,1176,216,878]" captionTargetPageId="2" captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11512958" httpUri="https://zenodo.org/record/11512958/files/figure.png" pageId="32" pageNumber="525">Fig. 2B</figureCitation>
), females without such setae; middle tarsi with tarsomeres 1 to 3 distinctly broader in males (as in 
<figureCitation id="0C832A6FEB48ED65FBDFFE3467AA87C8" box="[1082,1165,411,435]" captionStart="FIGURE" captionStartId="2.[171,183,897,916]" captionTargetBox="[120,1176,216,878]" captionTargetId="figure-193@2.[120,1176,216,878]" captionTargetPageId="2" captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11512958" httpUri="https://zenodo.org/record/11512958/files/figure.png" pageId="32" pageNumber="525">Fig. 2D</figureCitation>
) than in females and with two rows of adhesive squamosetae ventrally (as in 
<figureCitation id="0C832A6FEB48ED65FC55FE14672487A8" box="[944,1027,443,467]" captionStart="FIGURE" captionStartId="2.[171,183,897,916]" captionTargetBox="[120,1176,216,878]" captionTargetId="figure-193@2.[120,1176,216,878]" captionTargetPageId="2" captionText="FIGURE 2. Parts of legs, Cyclosomus acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China). A. Left hind tibial medial spur and tarsomere 1, dorsal aspect; B,C. Male front tarsus, B. ventral aspect, C. dorsal aspect; D,E. Male middle tarsus, D. dorsal aspect, E. ventral aspect. Scale lines = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.11512958" httpUri="https://zenodo.org/record/11512958/files/figure.png" pageId="32" pageNumber="525">Fig. 2E</figureCitation>
), absent from females. Tarsal claws smooth, edentate.
</paragraph>
<paragraph id="940736EAEB48ED65FF4EFE5463D48408" blockId="32.[123,1174,219,1461]" pageId="32" pageNumber="525">
Male genitalia. Median lobe with shaft distinctly thickened and markedly arcuate, ventral curvature distinctly curved throughout, shaft tapered gently to apex in lateral aspect (
<figureCitation id="0C832A6FEB48ED65FBC4FDB467AE8448" box="[1057,1161,539,563]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="32" pageNumber="525">Fig. 16H</figureCitation>
), apical lamella smoothly rounded and slightly elongate, slightly narrowed basally in dorsal aspect (
<figureCitation id="0C832A6FEB48ED65FF66FDF463C18408" box="[131,230,603,627]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="32" pageNumber="525">Fig. 17H</figureCitation>
).
</paragraph>
<paragraph id="940736EAEB48ED65FF4EFDD36177856F" blockId="32.[123,1174,219,1461]" pageId="32" pageNumber="525">
<emphasis id="A6CCEAF8EB48ED65FF4EFDD362B084E8" bold="true" box="[171,407,636,659]" pageId="32" pageNumber="525">Habitat distribution.</emphasis>
Unknown, but presumed to be restricted to the sandy shores of medium to large rivers at low elevation, like members of most other species of the genus. The elevation of New 
<collectingRegion id="567CF808EB48ED65FF52FD1463D584A8" box="[183,242,699,723]" country="India" name="Delhi" pageId="32" pageNumber="525">Delhi</collectingRegion>
is about 
<quantity id="53409B0FEB48ED65FEB8FD1462BA84A8" box="[349,413,699,723]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="525" unit="m" value="200.0">200m</quantity>
, of Morababad about 
<quantity id="53409B0FEB48ED65FD76FD1461FB84A8" box="[659,732,699,723]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="32" pageNumber="525" unit="m" value="190.0">190 m</quantity>
, and that of suitable habitat in Chitwan National Park about 
<quantity id="53409B0FEB48ED65FEBDFD7362B98488" box="[344,414,732,756]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="32" pageNumber="525" unit="m" value="550.0">550 m</quantity>
, so members of this species occupy at least that altitudinal range at the southern base of the Himalayan Mountains.
</paragraph>
<paragraph id="940736EAEB48ED65FF4EFCB361EE820F" blockId="32.[123,1174,219,1461]" pageId="32" pageNumber="525">
<emphasis id="A6CCEAF8EB48ED65FF4EFCB362FE8548" bold="true" box="[171,473,796,819]" pageId="32" pageNumber="525">Geographical distribution.</emphasis>
<figureCitation id="0C832A6FEB48ED65FE3AFCB3610A854F" box="[479,557,796,820]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="32" pageNumber="525">Fig. 22</figureCitation>
. 
<collectingCounty id="7D664E66EB48ED65FDD3FCB36142854F" box="[566,613,796,820]" pageId="32" pageNumber="525">This</collectingCounty>
species is apparently restricted to northern 
<collectingCountry id="ECAF767AEB48ED65FBD7FCB3674E854F" box="[1074,1129,796,820]" name="India" pageId="32" pageNumber="525">India</collectingCountry>
and 
<collectingCountry id="ECAF767AEB48ED65FF9EFC93639A852F" box="[123,189,828,852]" name="Nepal" pageId="32" pageNumber="525">Nepal</collectingCountry>
, along tributaries of the 
<collectingMunicipality id="7463AC90EB48ED65FE20FC936170852F" box="[453,599,828,852]" pageId="32" pageNumber="525">Ganges River</collectingMunicipality>
draining the southern slope of the 
<location id="91676031EB48ED65FC35FC936396850F" LSID="urn:lsid:plazi:treatment:1C1187FCEB75ED65FF9EFDF3639B83CE:91676031EB48ED65FC35FC936396850F" country="India" county="This" municipality="Ganges River" name="Himalayan Mountains" pageId="32" pageNumber="525" stateProvince="Delhi">Himalayan Mountains</location>
. We have found only three specific records, two of which represent the known extremes of the range of the species. 
<location id="91676031EB48ED65FE65FCD3628E85EF" LSID="urn:lsid:plazi:treatment:1C1187FCEB75ED65FF9EFDF3639B83CE:91676031EB48ED65FE65FCD3628E85EF" box="[384,425,892,916]" country="India" county="This" municipality="Ganges River" name="The" pageId="32" pageNumber="525" stateProvince="Delhi">The</location>
western locality is 
<location id="91676031EB48ED65FD92FCD2618E85EF" LSID="urn:lsid:plazi:treatment:1C1187FCEB75ED65FF9EFDF3639B83CE:91676031EB48ED65FD92FCD2618E85EF" box="[631,681,893,916]" country="India" county="This" municipality="Ganges River" name="New" pageId="32" pageNumber="525" stateProvince="Delhi">New</location>
<collectingRegion id="567CF808EB48ED65FD4AFCD361CD85EF" box="[687,746,892,916]" country="India" name="Delhi" pageId="32" pageNumber="525">Delhi</collectingRegion>
(
<collectingRegion id="567CF808EB48ED65FD1DFCD3605485EF" box="[760,883,892,916]" country="India" name="Delhi" pageId="32" pageNumber="525">Delhi State</collectingRegion>
, 
<collectingCountry id="ECAF767AEB48ED65FC9BFCD3609285EF" box="[894,949,892,916]" name="India" pageId="32" pageNumber="525">India</collectingCountry>
[in 
<collectionCode id="F2A9AE2FEB48ED65FC3BFCD3671385EF" box="[990,1076,892,916]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="32" pageNumber="525" type="Museum">NMNH</collectionCode>
]) and the eastern site is in 
<location id="91676031EB48ED65FECEFC33613985CF" LSID="urn:lsid:plazi:treatment:1C1187FCEB75ED65FF9EFDF3639B83CE:91676031EB48ED65FECEFC33613985CF" box="[299,542,924,948]" country="India" county="This" municipality="Ganges River" name="Chitwan National Park" pageId="32" pageNumber="525" stateProvince="Delhi">Chitwan National Park</location>
(Narayani Zone, 
<collectingCountry id="ECAF767AEB48ED65FD3CFC33603E85CF" box="[729,793,924,948]" name="Nepal" pageId="32" pageNumber="525">Nepal</collectingCountry>
[in 
<collectionCode id="F2A9AE2FEB48ED65FCA6FC33605E85CF" box="[835,889,924,948]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="32" pageNumber="525" type="Museum">CAS</collectionCode>
, 
<collectionCode id="F2A9AE2FEB48ED65FC61FC3360FA85CF" box="[900,989,924,948]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="32" pageNumber="525" type="Museum">NMNH</collectionCode>
]). However, most of the specimens we examined were labeled simply “ 
<collectingCountry id="ECAF767AEB48ED65FD53FC1361D685AF" box="[694,753,956,980]" name="India" pageId="32" pageNumber="525">India</collectingCountry>
” and were from the “Nevinson Collection” in 
<collectionCode id="F2A9AE2FEB48ED65FF32FC726218858F" box="[215,319,989,1012]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="32" pageNumber="525" type="Museum">NHMUK</collectionCode>
. According to Max Barclay (
<collectionCode id="F2A9AE2FEB48ED65FD65FC7361CE858F" box="[640,745,988,1012]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="32" pageNumber="525" type="Museum">NHMUK</collectionCode>
, personal communication), most of the Nevinson material labeled “ 
<collectingCountry id="ECAF767AEB48ED65FE46FC5362C7826F" box="[419,480,1020,1044]" name="India" pageId="32" pageNumber="525">India</collectingCountry>
,” and for which no further supporting specificity has been found, has been from northern 
<collectingCountry id="ECAF767AEB48ED65FE9CFBB36295824F" box="[377,434,1052,1076]" name="India" pageId="32" pageNumber="525">India</collectingCountry>
, including Punjab, 
<collectingRegion id="567CF808EB48ED65FD65FBB361EA824F" box="[640,717,1052,1076]" country="India" name="Sikkim" pageId="32" pageNumber="525">Sikkim</collectingRegion>
and 
<collectingRegion id="567CF808EB48ED65FCE4FBB260AF824F" box="[769,904,1052,1076]" country="India" name="West Bengal" pageId="32" pageNumber="525">West Bengal</collectingRegion>
(“Darjeeling”). It is likely, therefore, that the range of this species extends somewhat further east and west along the southern edge of the Himalaya than has yet been confirmed.
</paragraph>
<paragraph id="940736EAEB48ED65FF4EFBD2615382AF" blockId="32.[123,1174,219,1461]" pageId="32" pageNumber="525">
<emphasis id="A6CCEAF8EB48ED65FF4EFBD2629A82EF" bold="true" box="[171,445,1149,1172]" pageId="32" pageNumber="525">Geographical variation.</emphasis>
Although there is some individual variation (
<figureCitation id="0C832A6FEB48ED65FC5FFBD3674982EF" box="[954,1134,1148,1172]" captionStart="FIGURE" captionStartId="30.[171,183,1573,1592]" captionTargetBox="[134,1162,233,1556]" captionTargetId="figure-9@30.[120,1174,216,1560]" captionTargetPageId="30" captionText="FIGURE 13. Cyclosomus vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. A. Holotype male, dorsal habitus; B. Labels associated with holotype; C. Dark male (India); D. Pale male (India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512984" httpUri="https://zenodo.org/record/11512984/files/figure.png" pageId="32" pageNumber="525">Figs. 13A, C, D</figureCitation>
) in development of the dark color pattern of the elytra in this species, we could not discern any particular geographic component to that variation.
</paragraph>
<paragraph id="940736EAEB48ED65FF4EFB72639B83CE" blockId="32.[123,1174,219,1461]" pageId="32" pageNumber="525">
<emphasis id="A6CCEAF8EB48ED65FF4EFB726065828F" bold="true" box="[171,834,1245,1269]" pageId="32" pageNumber="525">
Geographical relationships with other 
<taxonomicName id="53B84D69EB48ED65FD84FB726065828F" box="[609,834,1245,1269]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="525" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A6CCEAF8EB48ED65FD84FB7261C2828E" bold="true" box="[609,741,1245,1269]" italics="true" pageId="32" pageNumber="525">Cyclosomus</emphasis>
species.
</taxonomicName>
</emphasis>
The known range of 
<taxonomicName id="53B84D69EB48ED65FBC2FB726381836F" authorityName="Cueva-Dabkoski and Kavanaugh" authorityYear="2023" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="525" phylum="Arthropoda" rank="species" species="vespertilio">
<emphasis id="A6CCEAF8EB48ED65FBC2FB726381836F" italics="true" pageId="32" pageNumber="525">C. vespertilio</emphasis>
</taxonomicName>
is within the broad range of 
<taxonomicName id="53B84D69EB48ED65FE36FB526172836F" authorityName=", Chaudoir" authorityYear="1876" box="[467,597,1276,1300]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="525" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB48ED65FE36FB526172836F" box="[467,597,1276,1300]" italics="true" pageId="32" pageNumber="525">C. flexuosus</emphasis>
</taxonomicName>
at its northern limit, so these two species may be sympatric or even syntopic in at least some areas. However, they have not yet been recorded together from any locality. It is also possible that 
<taxonomicName id="53B84D69EB48ED65FDA2FA9261FD832E" authorityName="Cueva-Dabkoski and Kavanaugh" authorityYear="2023" box="[583,730,1341,1365]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="525" phylum="Arthropoda" rank="species" species="vespertilio">
<emphasis id="A6CCEAF8EB48ED65FDA2FA9261FD832E" box="[583,730,1341,1365]" italics="true" pageId="32" pageNumber="525">C. vespertilio</emphasis>
</taxonomicName>
is sympatric with 
<taxonomicName id="53B84D69EB48ED65FC49FA92676F832E" box="[940,1096,1341,1365]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="525" phylum="Arthropoda" rank="species" species="marginatus">
<emphasis id="A6CCEAF8EB48ED65FC49FA92676F832E" box="[940,1096,1341,1365]" italics="true" pageId="32" pageNumber="525">C. marginatus</emphasis>
</taxonomicName>
and/or 
<taxonomicName id="53B84D69EB48ED65FF9EFAF263DF830E" box="[123,248,1373,1397]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="525" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A6CCEAF8EB48ED65FF9EFAF263DF830E" box="[123,248,1373,1397]" italics="true" pageId="32" pageNumber="525">C. suturalis</emphasis>
</taxonomicName>
in the eastern or southeastern part of its range, depending on just where in “Bengal” (i.e, the area now including 
<collectingRegion id="567CF808EB48ED65FE4FFAD2611383EE" box="[426,564,1405,1429]" country="India" name="West Bengal" pageId="32" pageNumber="525">West Bengal</collectingRegion>
, 
<collectingCountry id="ECAF767AEB48ED65FDA5FAD2615083EE" box="[576,631,1405,1429]" name="India" pageId="32" pageNumber="525">India</collectingCountry>
and 
<collectingCountry id="ECAF767AEB48ED65FD48FAD2600E83EE" box="[685,809,1405,1429]" name="Bangladesh" pageId="32" pageNumber="525">Bangladesh</collectingCountry>
[see above]) the last two actually occur.
</paragraph>
</subSubSection>
</treatment>
</document>