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499 lines
67 KiB
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<document id="2501D146A41462BE93BE75A4A1FB40B2" ID-CLB-Dataset="299696" ID-DOI="10.5281/zenodo.11512952" ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512952" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1717721037103" checkinUser="felipe" docAuthor="Kavanaugh, David H. & Cueva-Dabkoski, Mollie" docDate="2023" docId="1C1187FCEB72ED58FF9EFB736739844F" docLanguage="en" docName="ProcCalifAcadSci.67.21.493-537.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (21)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Cyclosomus suturalis" docType="treatment" docVersion="5" lastPageNumber="522" masterDocId="E028FF84EB68ED45FFE5FFAF6327867B" masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia." masterLastPageNumber="537" masterPageNumber="493" pageNumber="519" updateTime="1720721490288" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="1FF6D10CFF9B65934A58200EF2C7D5F4">
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<mods:title id="1E2FC6B015E000B386BE03239E2C35B5">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
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</mods:titleInfo>
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<mods:name id="8B2CA181FA8DFF0BCA02334EAC7E96FB" type="personal">
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<mods:roleTerm id="1DE65817EA4195FDB4A34DCEAD7A6FA2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FD95980708EEEEF49634D9EEE5B78A93">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="8AC0015A4FC30E16F373692027E00FF8">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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</mods:name>
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<mods:name id="2E0CAA24223C7457066D933E3DC71407" type="personal">
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<mods:roleTerm id="DFD157D826B5A8D9C1A865FB3E467D1B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DCCDE428B23F77293BE9E09D68A5D1E0">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:affiliation id="3498C8F46A6E4789CF73E3D405428B34">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="B0F95216E72BC217BB209CDBAC411493">text</mods:typeOfResource>
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<mods:titleInfo id="F0A47EE1E1FFAD18924FB3320CBE663B">
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<mods:title id="EE032F1C5FA055CB66F246DE80729887">Proceedings of the California Academy of Sciences</mods:title>
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</mods:titleInfo>
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<mods:part id="6C73AEA70444117CB1C8677E7A36A8FA">
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<mods:date id="96ADCA18823730452536B61947C207B5">2023</mods:date>
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<mods:detail id="F2AE0EE7A3DC6E6B03206C26DB8E1A2C" type="pubDate">
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<mods:number id="C3341B5C6482D574CF71E677BF0D47CD">2023-02-15</mods:number>
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</mods:detail>
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<mods:detail id="B3E030296F783B0A08D1C77986B00D6A" type="volume">
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<mods:number id="5A61C809483B17517D8B282D0F318946">67</mods:number>
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</mods:detail>
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<mods:detail id="484671E111F63E7BD7DE0B9072F6A7AC" type="issue">
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<mods:number id="82446A4EC7ACE3DB6A244C6CF5955667">21</mods:number>
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<mods:start id="59486B9850DE93CE7BC829EABD3D3A81">493</mods:start>
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<mods:end id="00940E85691F1795FB8B6A444160A7CF">537</mods:end>
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<mods:classification id="50950780144999C1B0A9926CEE9CEBCA">journal article</mods:classification>
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<mods:identifier id="5C04F1ED34674A084F31D22B11F218BC" type="CLB-Dataset">299696</mods:identifier>
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<mods:identifier id="24C430822BE572A439F50256CB0EE8E4" type="DOI">10.5281/zenodo.11512952</mods:identifier>
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<mods:identifier id="BA668E70D342D61F24287EC84CC64E08" type="GBIF-Dataset">2b56e3b0-2274-4963-b243-9c991b07072c</mods:identifier>
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<mods:identifier id="B1EF4F7CFFC3F8AA53BE933FE911BAFE" type="ISSN">0068-547X</mods:identifier>
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<mods:identifier id="770E961D29D4CF49AC26816977B2A592" type="Zenodo-Dep">11512952</mods:identifier>
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<treatment id="1C1187FCEB72ED58FF9EFB736739844F" ID-DOI="http://doi.org/10.5281/zenodo.12726734" ID-GBIF-Taxon="235039981" ID-Zenodo-Dep="12726734" LSID="urn:lsid:plazi:treatment:1C1187FCEB72ED58FF9EFB736739844F" httpUri="http://treatment.plazi.org/id/1C1187FCEB72ED58FF9EFB736739844F" lastPageId="29" lastPageNumber="522" pageId="26" pageNumber="519">
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<subSubSection id="DCA26561EB72ED5FFF9EFB73612F828F" box="[123,520,1244,1268]" pageId="26" pageNumber="519" type="nomenclature">
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<paragraph id="940736EAEB72ED5FFF9EFB73612F828F" blockId="26.[123,1174,1244,1726]" box="[123,520,1244,1268]" pageId="26" pageNumber="519">
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<heading id="CF4F8186EB72ED5FFF9EFB73612F828F" box="[123,520,1244,1268]" fontSize="10" level="2" pageId="26" pageNumber="519" reason="4">
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<taxonomicName id="53B84D69EB72ED5FFF9EFB73612F828F" ID-CoL="32X5F" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[123,520,1244,1268]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="suturalis">
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<emphasis id="A6CCEAF8EB72ED5FFF9EFB73612F828F" bold="true" box="[123,520,1244,1268]" pageId="26" pageNumber="519">
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<emphasis id="A6CCEAF8EB72ED5FFF9EFB736241828F" bold="true" box="[123,358,1244,1268]" italics="true" pageId="26" pageNumber="519">Cyclosomus suturalis</emphasis>
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(Wiedemann)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB72ED5FFF9EFB5362FB8369" box="[123,476,1276,1298]" pageId="26" pageNumber="519" type="description">
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<paragraph id="940736EAEB72ED5FFF9EFB5362FB8369" blockId="26.[123,1174,1244,1726]" box="[123,476,1276,1298]" pageId="26" pageNumber="519">
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<figureCitation id="0C832A6FEB72ED5FFF9EFB5363C48369" box="[123,227,1276,1298]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="26" pageNumber="519">Figures 12</figureCitation>
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,
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<figureCitation id="0C832A6FEB72ED5FFF0BFB53623E8369" box="[238,281,1276,1298]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="26" pageNumber="519">14H</figureCitation>
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,
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<figureCitation id="0C832A6FEB72ED5FFEC1FB5362688369" box="[292,335,1276,1298]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="26" pageNumber="519">16G</figureCitation>
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,
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<figureCitation id="0C832A6FEB72ED5FFEBFFB5362A28369" box="[346,389,1276,1298]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="26" pageNumber="519">17G</figureCitation>
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,
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<figureCitation id="0C832A6FEB72ED5FFE75FB53629E8369" box="[400,441,1276,1298]" captionStart="FIGURE" captionStartId="37.[171,183,1577,1596]" captionTargetBox="[120,1176,216,1561]" captionTargetId="figure-14@37.[120,1176,216,1561]" captionTargetPageId="37" captionText="FIGURE 18. Photographs of habitats for Cyclosomus species. A. C. acutangulus Kavanaugh & Cueva-Dabkoski (Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China; photograph by David H. Kavanaugh); B. C. suturalis (Wiedemann) (Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China; photograph by Hong-Bin Liang)." figureDoi="http://doi.org/10.5281/zenodo.11512996" httpUri="https://zenodo.org/record/11512996/files/figure.png" pageId="26" pageNumber="519">18B</figureCitation>
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,
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<figureCitation id="0C832A6FEB72ED5FFE21FB5362FB8369" box="[452,476,1276,1298]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="26" pageNumber="519">22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB72ED5FFF9EFA8F62CC802C" pageId="26" pageNumber="519" type="reference_group">
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<paragraph id="940736EAEB72ED5FFF9EFA8F600A807B" blockId="26.[123,1174,1244,1726]" pageId="26" pageNumber="519">
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<taxonomicName id="53B84D69EB72ED5FFF9EFA8F6120834D" authority="Wiedemann, 1819: 169" authorityName="Wiedemann. A" authorityPageNumber="169" authorityYear="1819" box="[123,519,1312,1334]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="suturalis">
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<emphasis id="A6CCEAF8EB72ED5FFF9EFA8F6202834D" box="[123,293,1312,1334]" italics="true" pageId="26" pageNumber="519">Scolytus suturalis</emphasis>
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<bibRefCitation id="F0294B1BEB72ED5FFECEFA8F6120834D" author="WIEDEMANN, C. R. W." box="[299,519,1312,1334]" pageId="26" pageNumber="519" pagination="157 - 183" refId="ref24121" refString="WIEDEMANN, C. R. W. 1819. Neue Kafer aus Bengalen und Java beschrieben vom herausgeben. Zoologische Magazin 1: 157 - 183." type="journal article" year="1819">Wiedemann, 1819:169</bibRefCitation>
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</taxonomicName>
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.
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<typeStatus id="4B038848EB72ED5FFDF4FA8F61BB834D" box="[529,668,1312,1334]" pageId="26" pageNumber="519" type="lectotype">LECTOTYPE</typeStatus>
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, here designated, a male, in
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<collectionCode id="F2A9AE2FEB72ED5FFC48FA8F60D2834D" box="[941,1013,1312,1334]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="519">ZMUC</collectionCode>
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, labeled: “Type” [red label]/ “Bengala Westermann flexuosus F. suturalis Wied.” [handwritten]/ “
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<collectionCode id="F2A9AE2FEB72ED5FFC37FA92673E8328" box="[978,1049,1341,1363]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="519">ZMUC</collectionCode>
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00022690”/ “
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<typeStatus id="4B038848EB72ED5FFF56FAF56219830B" box="[179,318,1370,1392]" pageId="26" pageNumber="519" type="lectotype">LECTOTYPE</typeStatus>
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<taxonomicName id="53B84D69EB72ED5FFEA0FAF561B8830B" authority="Wiedemann 1819" authorityName="Wiedemann. A" authorityYear="1819" box="[325,671,1370,1392]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="suturalis">
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<emphasis id="A6CCEAF8EB72ED5FFEA0FAF562C8830B" box="[325,495,1370,1392]" italics="true" pageId="26" pageNumber="519">Scolytus suturalis</emphasis>
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<bibRefCitation id="F0294B1BEB72ED5FFE13FAF561B8830B" author="WIEDEMANN, C. R. W." box="[502,671,1370,1392]" pageId="26" pageNumber="519" pagination="157 - 183" refId="ref24121" refString="WIEDEMANN, C. R. W. 1819. Neue Kafer aus Bengalen und Java beschrieben vom herausgeben. Zoologische Magazin 1: 157 - 183." type="journal article" year="1819">Wiedemann 1819</bibRefCitation>
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</taxonomicName>
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design. by D.H. Kavanaugh & M. Cueva-Dabkoski 2023” [red label].
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<typeStatus id="4B038848EB72ED5FFEB9FAD862CF83F6" box="[348,488,1399,1421]" pageId="26" pageNumber="519" type="paralectotype">Paralectotypes</typeStatus>
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examined:
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<specimenCount id="82BEFD63EB72ED5FFDBCFAD861BB83F6" box="[601,668,1399,1421]" count="1" pageId="26" pageNumber="519" type="male">1 male</specimenCount>
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, also in
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<collectionCode id="F2A9AE2FEB72ED5FFD09FAD8601383F6" box="[748,820,1399,1421]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="26" pageNumber="519">ZMUC</collectionCode>
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, labeled: “Mus. Westerm.”/ “TYPE” [red label]/ “
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<taxonomicName id="53B84D69EB72ED5FFECDFA3B62BA83D1" authorityName=", Kavanaugh" authorityYear="2015" box="[296,413,1428,1450]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexuosus">S. flexuosus</taxonomicName>
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var.
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<taxonomicName id="53B84D69EB72ED5FFE2BFA3B61E383D1" authority="Wied. Bengal" authorityName="Wied. Bengal" box="[462,708,1428,1450]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="suturalis">S. suturalis Wied. Bengal</taxonomicName>
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<date id="E006102AEB72ED5FFD2FFA3B600983D1" box="[714,814,1428,1450]" pageId="26" pageNumber="519" value="1809-05">May 1809</date>
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” [handwritten]/
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<typeStatus id="4B038848EB72ED5FFC2AFA3B67B183D1" box="[975,1174,1428,1450]" pageId="26" pageNumber="519" type="paralectotype">PARALECTOTYPE</typeStatus>
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<taxonomicName id="53B84D69EB72ED5FFF4EFA1E612083BC" ID-CoL="4VTPQ" authority="Wiedemann 1819" authorityName="Wiedemann. A" authorityYear="1819" box="[171,519,1457,1479]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFF4EFA1E627183BC" box="[171,342,1457,1479]" italics="true" pageId="26" pageNumber="519">Scolytus suturalis</emphasis>
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFEB8FA1E612083BC" author="WIEDEMANN, C. R. W." box="[349,519,1457,1479]" pageId="26" pageNumber="519" pagination="157 - 183" refId="ref24121" refString="WIEDEMANN, C. R. W. 1819. Neue Kafer aus Bengalen und Java beschrieben vom herausgeben. Zoologische Magazin 1: 157 - 183." type="journal article" year="1819">Wiedemann 1819</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
design. by D.H. Kavanaugh & M. Cueva-Dabkoski 2023” [yellow label]/ “
|
|||
|
|
<taxonomicName id="53B84D69EB72ED5FFF1DFA6261478398" ID-CoL="32X59" authority="Motschulsky" authorityName="Motschulsky" box="[248,608,1485,1507]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFF1DFA6262F98398" box="[248,478,1485,1507]" italics="true" pageId="26" pageNumber="519">Cyclosomus marginatus</emphasis>
|
|||
|
|
Motschulsky
|
|||
|
|
</taxonomicName>
|
|||
|
|
det. D.H.
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFD21FA62604C8398" author="KAVANAUGH, D. H. & M. CUEVA-DABKOSKI & H. B. LIANG" box="[708,875,1485,1507]" pageId="26" pageNumber="519" pagination="451 - 491" refId="ref23619" refString="KAVANAUGH, D. H., M. CUEVA-DABKOSKI, AND H. B. LIANG. 2023. Inventory of the carabid beetle fauna of the Gaoligong Mountains, western Yunnan Province, China: species of the tribe Cyclosomini Laporte, 1934 (Coleoptera: Carabidae), with descriptions of two new species. Proceedings of the California Academy of Sciences (Series 4) 67: 451 - 491." type="journal article" year="2023">Kavanaugh 2023</bibRefCitation>
|
|||
|
|
”.
|
|||
|
|
<typeStatus id="4B038848EB72ED5FFC98FA61608A8398" box="[893,941,1486,1507]" pageId="26" pageNumber="519">Type</typeStatus>
|
|||
|
|
locality: “Bengal” [area which now includes
|
|||
|
|
<collectingRegion id="567CF808EB72ED5FFE97FA4462CB807B" box="[370,492,1514,1536]" country="India" name="West Bengal" pageId="26" pageNumber="519">West Bengal</collectingRegion>
|
|||
|
|
(eastern
|
|||
|
|
<collectingCountry id="ECAF767AEB72ED5FFDA0FA45615F807B" box="[581,632,1514,1536]" name="India" pageId="26" pageNumber="519">India</collectingCountry>
|
|||
|
|
) and
|
|||
|
|
<collectingCountry id="ECAF767AEB72ED5FFD4BFA456003807B" box="[686,804,1514,1536]" name="Bangladesh" pageId="26" pageNumber="519">Bangladesh</collectingCountry>
|
|||
|
|
].
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB72ED5FFF9EF9A861368066" blockId="26.[123,1174,1244,1726]" box="[123,529,1543,1565]" pageId="26" pageNumber="519">
|
|||
|
|
<taxonomicName id="53B84D69EB72ED5FFF9EF9A8612B8066" authority=", Chaudoir 1876: 32" authorityName=", Chaudoir" authorityPageNumber="32" authorityYear="1876" box="[123,524,1543,1565]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFF9EF9A862698066" box="[123,334,1543,1565]" italics="true" pageId="26" pageNumber="519">Cyclosomus flexuosus</emphasis>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFEBEF9A8612B8066" author="CHAUDOIR, M." box="[347,524,1543,1565]" pageId="26" pageNumber="519" pagination="1 - 84" refId="ref23225" refString="CHAUDOIR, M. 1876. Etude monographique des Masoreides, des Tetragonoderides et du genre Nemotarsus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 51: 1 - 84." type="journal article" year="1876">Chaudoir 1876:32</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB72ED5FFF9EF98B62CC802C" blockId="26.[123,1174,1244,1726]" pageId="26" pageNumber="519">
|
|||
|
|
<taxonomicName id="53B84D69EB72ED5FFF9EF98B62D08041" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[123,503,1572,1594]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFF9EF98B627C8041" box="[123,347,1572,1594]" italics="true" pageId="26" pageNumber="519">Cyclosomus suturalis</emphasis>
|
|||
|
|
(Wiedemann)
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFDEFF98B61F98041" author="CHAUDOIR, M." box="[522,734,1572,1594]" pageId="26" pageNumber="519" pagination="1 - 84" refId="ref23225" refString="CHAUDOIR, M. 1876. Etude monographique des Masoreides, des Tetragonoderides et du genre Nemotarsus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 51: 1 - 84." type="journal article" year="1876">Chaudoir (1876:32)</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFD15F98B60E88041" author="ANDREWES, H. E." box="[752,975,1572,1594]" pageId="26" pageNumber="519" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:166</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFC06F98B67628041" author="ANDREWES, H. E." box="[995,1093,1572,1594]" pageId="26" pageNumber="519" pagination="459 - 468" refId="ref23019" refString="ANDREWES, H. E. 1924. XXII. On the Oriental Carabidae of the " Reise Novara. " Transactions of the Entomological Society of London 1923: 459 - 468." type="journal article" year="1924">1924:464</bibRefCitation>
|
|||
|
|
);
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFB84F98B67B18041" author="CSIKI, E." box="[1121,1174,1572,1594]" pageId="26" pageNumber="519" pagination="1279 - 1598" refId="ref23258" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki</bibRefCitation>
|
|||
|
|
(1932:1295);
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFECAF9ED62C0802C" author="LORENZ, W." box="[303,487,1601,1623]" pageId="26" pageNumber="519" refId="ref23840" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz (2005:452)</bibRefCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB72ED5EFF4EF9C9615184EE" lastPageId="27" lastPageNumber="520" pageId="26" pageNumber="519" type="discussion">
|
|||
|
|
<paragraph id="940736EAEB72ED5EFF4EF9C9615184EE" blockId="26.[123,1174,1244,1726]" lastBlockId="27.[123,1174,219,1725]" lastPageId="27" lastPageNumber="520" pageId="26" pageNumber="519">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFF4EF9C961128006" bold="true" box="[171,565,1638,1661]" pageId="26" pageNumber="519">Notes on types and nomenclature.</emphasis>
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFDDBF9CA60348006" author="CHAUDOIR, M." box="[574,787,1637,1661]" pageId="26" pageNumber="519" pagination="1 - 84" refId="ref23225" refString="CHAUDOIR, M. 1876. Etude monographique des Masoreides, des Tetragonoderides et du genre Nemotarsus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 51: 1 - 84." type="journal article" year="1876">Chaudoir (1876:32)</bibRefCitation>
|
|||
|
|
considered
|
|||
|
|
<taxonomicName id="53B84D69EB72ED5FFC7CF9C9673F8006" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[921,1048,1637,1661]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFC7CF9C9673F8006" box="[921,1048,1637,1661]" italics="true" pageId="26" pageNumber="519">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as a junior synonym of
|
|||
|
|
<taxonomicName id="53B84D69EB72ED5FFF1BF92962A780E6" authorityName=", Chaudoir" authorityYear="1876" box="[254,384,1669,1693]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFF1BF92962A780E6" box="[254,384,1669,1693]" italics="true" pageId="26" pageNumber="519">C. flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, but
|
|||
|
|
<bibRefCitation id="F0294B1BEB72ED5FFE56F92A61BC80E6" author="ANDREWES, H. E." box="[435,667,1669,1693]" pageId="26" pageNumber="519" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:166)</bibRefCitation>
|
|||
|
|
confirmed its status as a distinct species. In that same paper, Andrewes recognized
|
|||
|
|
<taxonomicName id="53B84D69EB72ED5FFE15F90961AE80C6" box="[496,649,1702,1725]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFE15F90961AE80C6" box="[496,649,1702,1725]" italics="true" pageId="26" pageNumber="519">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as a junior synonym of
|
|||
|
|
<taxonomicName id="53B84D69EB72ED5FFC75F909672A80C6" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[912,1037,1701,1725]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB72ED5FFC75F909672A80C6" box="[912,1037,1701,1725]" italics="true" pageId="26" pageNumber="519">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. This difference of opinion between these two superb taxonomists can possibly be explained by the fact that the type series of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFEA6FF5362E58768" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[323,450,251,275]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFEA6FF5362E58768" box="[323,450,251,275]" italics="true" pageId="27" pageNumber="520">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
includes
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFDCAFF5361F58768" box="[559,722,251,275]" count="2" pageId="27" pageNumber="520" type="generic">two specimens</specimenCount>
|
|||
|
|
, which are not, in our view, conspecific. Thus, they may have examined different specimens in making their assessments. The male specimen that we have selected as
|
|||
|
|
<typeStatus id="4B038848EB73ED5EFE59FE9461398728" box="[444,542,315,339]" pageId="27" pageNumber="520" type="lectotype">lectotype</typeStatus>
|
|||
|
|
actually resembles some small male specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFBA3FE93639F870F" authorityName=", Chaudoir" authorityYear="1876" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFBA3FE93639F870F" italics="true" pageId="27" pageNumber="520">C. flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in size and elytral color pattern. The female specimen we designate as
|
|||
|
|
<typeStatus id="4B038848EB73ED5EFC4BFEF36719870F" box="[942,1086,348,372]" pageId="27" pageNumber="520" type="paralectotype">paralectotype</typeStatus>
|
|||
|
|
appears to be a specimen of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFEB5FED262CF87EF" box="[336,488,381,404]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFEB5FED262CF87EF" box="[336,488,381,404]" italics="true" pageId="27" pageNumber="520">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, with the size and elytral color pattern matching the
|
|||
|
|
<typeStatus id="4B038848EB73ED5EFBF2FED3675E87EF" box="[1047,1145,380,404]" pageId="27" pageNumber="520" type="lectotype">lectotype</typeStatus>
|
|||
|
|
of that species. If Chaudoir had studied the male specimen and Andrewes the female specimen, their respective opinions are quite understandable. Because of variation in elytral color pattern seen in each of these species, Wiedemann’s original description is insufficient to distinguish between specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFF04FE526242846E" authorityName=", Chaudoir" authorityYear="1876" box="[225,357,509,533]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFF04FE526242846E" box="[225,357,509,533]" italics="true" pageId="27" pageNumber="520">C. flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFE90FE526129846E" box="[373,526,509,533]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFE90FE526129846E" box="[373,526,509,533]" italics="true" pageId="27" pageNumber="520">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, and
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFDA9FE5261ED846E" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[588,714,509,533]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFDA9FE5261ED846E" box="[588,714,509,533]" italics="true" pageId="27" pageNumber="520">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. By selecting the male specimen as
|
|||
|
|
<typeStatus id="4B038848EB73ED5EFBBCFE52638E844D" pageId="27" pageNumber="520" type="lectotype">lectotype</typeStatus>
|
|||
|
|
of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFF29FDB1626E844E" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[204,329,541,565]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFF29FDB1626E844E" box="[204,329,541,565]" italics="true" pageId="27" pageNumber="520">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, we are able to recognize Motschulsky’s
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFCE1FDB160BB844E" box="[772,924,542,565]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFCE1FDB160BB844E" box="[772,924,542,565]" italics="true" pageId="27" pageNumber="520">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
as the valid name for a distinct species (see above). If we had chosen the female specimen as
|
|||
|
|
<typeStatus id="4B038848EB73ED5EFC8BFD9260F4842E" box="[878,979,573,597]" pageId="27" pageNumber="520" type="lectotype">lectotype</typeStatus>
|
|||
|
|
, then
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFBF6FD9163BD840E" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFBF6FD9163BD840E" italics="true" pageId="27" pageNumber="520">C. marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
would have remained a junior synonym of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFD9BFDF161DA840E" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[638,765,605,629]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFD9BFDF161DA840E" box="[638,765,605,629]" italics="true" pageId="27" pageNumber="520">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and the male specimen would represented a new species currently without a name.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB73ED5EFF4EFD3061C082C2" pageId="27" pageNumber="520" type="diagnosis">
|
|||
|
|
<paragraph id="940736EAEB73ED5EFF4EFD3060478222" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="520">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFF4EFD30623884CD" bold="true" box="[171,287,671,694]" pageId="27" pageNumber="520">Diagnosis.</emphasis>
|
|||
|
|
Adults of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFE6BFD31612E84CD" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[398,521,670,694]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFE6BFD31612E84CD" box="[398,521,670,694]" italics="true" pageId="27" pageNumber="520">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
can be distinguished from those of other
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFC24FD3167B284CD" box="[961,1173,670,694]" pageId="27" pageNumber="520">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFC24FD31676584CD" box="[961,1090,670,694]" italics="true" pageId="27" pageNumber="520">Cyclosomus</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
in Asia by the following combination of character states: Body size medium to large for genus, BL males =
|
|||
|
|
<quantity id="53409B0FEB73ED5EFF33FD71624A848D" box="[214,365,734,758]" metricMagnitude="-3" metricUnit="m" metricValue="8.649999999999999" metricValueMax="9.1" metricValueMin="8.2" pageId="27" pageNumber="520" unit="mm" value="8.649999999999999" valueMax="9.1" valueMin="8.2">8.2 to 9.1 mm</quantity>
|
|||
|
|
, female =
|
|||
|
|
<quantity id="53409B0FEB73ED5EFE3BFD716114848D" box="[478,563,734,758]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="27" pageNumber="520" unit="mm" value="9.2">9.2 mm</quantity>
|
|||
|
|
; body form (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFD59FD71601A848D" box="[700,829,734,758]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="27" pageNumber="520">Fig. 12A, C</figureCitation>
|
|||
|
|
) slightly elongate ovoid; pronotum (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFF56FD516232856D" box="[179,277,766,790]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="27" pageNumber="520">Fig. 14H</figureCitation>
|
|||
|
|
) relatively broader (ratio PWM/PL = 2.44 to 2.50), disc rufopiceous to black, lateral pale bands well-defined, slightly narrowed at least in apical half and broadened basally, apical angles slightly narrowed and less broadly rounded, lateral margins smoothly arcuate, not sinuate near anterior angles; elytra with pattern of dark maculae less varied, with middle band of average (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFF66FCD063C385EC" box="[131,228,895,919]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="27" pageNumber="520">Fig. 12A</figureCitation>
|
|||
|
|
) to distinctly greater than average (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFD84FCD061E685EC" box="[609,705,895,919]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="27" pageNumber="520">Fig. 12C</figureCitation>
|
|||
|
|
) thickness for the genus and extended from midline to interval
|
|||
|
|
<quantity id="53409B0FEB73ED5EFEACFC30625485CC" box="[329,371,927,951]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="27" pageNumber="520" unit="in" value="6.0">6 in</quantity>
|
|||
|
|
most specimens, onto interval 7 and part of interval
|
|||
|
|
<quantity id="53409B0FEB73ED5EFC4BFC3060FF85CC" box="[942,984,927,951]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="27" pageNumber="520" unit="in" value="8.0">8 in</quantity>
|
|||
|
|
some specimens, preapical dark spot present, striae deeply impressed and intervals flat or slightly convex; elytral epipleura with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; male median lobe with apical lamella extended in line with ventral curvature of shaft in lateral aspect (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFE8AFB8F62F48243" box="[367,467,1056,1080]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="27" pageNumber="520">Fig. 16G</figureCitation>
|
|||
|
|
), apical lamella short and bluntly triangular in dorsal aspect (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFB8FFB8F638B8222" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="27" pageNumber="520">Fig. 17G</figureCitation>
|
|||
|
|
); specimen from eastern
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFE5FFBEE62D68222" box="[442,497,1089,1113]" name="India" pageId="27" pageNumber="520">India</collectingCountry>
|
|||
|
|
to westernmost Yunnan (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFCE7FBEE60748222" box="[770,851,1089,1113]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="27" pageNumber="520">Fig. 22</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB73ED5EFF4EFBCE61C082C2" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="520">
|
|||
|
|
Specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFEA6FBCD62988202" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[323,447,1121,1145]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFEA6FBCD62988202" box="[323,447,1121,1145]" italics="true" pageId="27" pageNumber="520">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
are most similar to those of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFD11FBCD60B28202" box="[756,917,1121,1145]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="acutangulus">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFD11FBCD60B28202" box="[756,917,1121,1145]" italics="true" pageId="27" pageNumber="520">C. acutangulus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and atypical specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFF7DFB2D622382E2" box="[152,260,1154,1177]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="inustus">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFF7DFB2D622382E2" box="[152,260,1154,1177]" italics="true" pageId="27" pageNumber="520">C. inustus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
that have the elytral preapical dark spot present. Refer to the key and Diagnosis sections for each of those species for distinguishing features.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB73ED5EFF4EFB6D63C68361" pageId="27" pageNumber="520" type="biology_ecology">
|
|||
|
|
<paragraph id="940736EAEB73ED5EFF4EFB6D63C68361" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="520">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFF4EFB6D62BD82A2" bold="true" box="[171,410,1218,1241]" pageId="27" pageNumber="520">Habitat distribution.</emphasis>
|
|||
|
|
Specimens from
|
|||
|
|
<collectingRegion id="567CF808EB73ED5EFDBAFB6D619782A2" box="[607,688,1218,1241]" country="China" name="Yunnan" pageId="27" pageNumber="520">Yunnan</collectingRegion>
|
|||
|
|
were collected at night on the upper sandy banks of the Longchuan River (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFE37FB4D61168281" box="[466,561,1250,1274]" captionStart="FIGURE" captionStartId="37.[171,183,1577,1596]" captionTargetBox="[120,1176,216,1561]" captionTargetId="figure-14@37.[120,1176,216,1561]" captionTargetPageId="37" captionText="FIGURE 18. Photographs of habitats for Cyclosomus species. A. C. acutangulus Kavanaugh & Cueva-Dabkoski (Longchuan Jiang just below bridge at Menglian village, Wuhe Township, Tengchong County, Yunnan, China; photograph by David H. Kavanaugh); B. C. suturalis (Wiedemann) (Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China; photograph by Hong-Bin Liang)." figureDoi="http://doi.org/10.5281/zenodo.11512996" httpUri="https://zenodo.org/record/11512996/files/figure.png" pageId="27" pageNumber="520">Fig. 18B</figureCitation>
|
|||
|
|
) at an elevation of
|
|||
|
|
<quantity id="53409B0FEB73ED5EFCE0FB4D606D8282" box="[773,842,1250,1274]" metricMagnitude="2" metricUnit="m" metricValue="7.34" pageId="27" pageNumber="520" unit="m" value="734.0">734 m</quantity>
|
|||
|
|
(H.B Liang, personal communication).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB73ED5EFF4EFA8C621C8380" pageId="27" pageNumber="520" type="materials_examined">
|
|||
|
|
<paragraph id="940736EAEB73ED5EFF4EFA8C621C8380" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="520">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFF4EFA8C62FE8341" bold="true" box="[171,473,1315,1338]" pageId="27" pageNumber="520">Geographical distribution.</emphasis>
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFE05FA8D61098341" box="[480,558,1314,1338]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="27" pageNumber="520">Fig. 22</figureCitation>
|
|||
|
|
. We examined a total of
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFCD0FA8C60CF8341" box="[821,1000,1314,1338]" count="7" pageId="27" pageNumber="520" type="generic">seven specimens</specimenCount>
|
|||
|
|
(
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFC10FA8D674F8341" box="[1013,1128,1314,1338]" count="4" pageId="27" pageNumber="520" type="male">four males</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFF9EFAED62348321" box="[123,275,1346,1370]" count="3" pageId="27" pageNumber="520" type="female">three females</specimenCount>
|
|||
|
|
) from the following localities:
|
|||
|
|
<materialsCitation id="24D03CB7EB73ED5EFD64FAEC620D83E0" ID-GBIF-Occurrence="4905152356" collectingDate="2020-05-22" collectionCode="IOZ" collectorName="H. B. Liang & Y. Xu" country="China" county="Mangshi" elevation="734" location="Longchuan Jiang" municipality="Zhefang Township" pageId="27" pageNumber="520" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Yunnan">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFD64FAEC61FD8321" bold="true" box="[641,730,1347,1370]" pageId="27" pageNumber="520">
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFD64FAEC61FD8321" box="[641,730,1347,1370]" name="China" pageId="27" pageNumber="520">CHINA</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="567CF808EB73ED5EFD0BFAEC60638321" box="[750,836,1347,1370]" country="China" name="Yunnan" pageId="27" pageNumber="520">Yunnan</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<collectingCounty id="7D664E66EB73ED5EFCB0FAED60928321" box="[853,949,1346,1370]" pageId="27" pageNumber="520">Mangshi</collectingCounty>
|
|||
|
|
,
|
|||
|
|
<collectingMunicipality id="7463AC90EB73ED5EFC22FAED67B18321" box="[967,1174,1346,1370]" pageId="27" pageNumber="520">Zhefang Township</collectingMunicipality>
|
|||
|
|
(
|
|||
|
|
<location id="91676031EB73ED5EFF67FACD621D8301" LSID="urn:lsid:plazi:treatment:1C1187FCEB72ED58FF9EFB736739844F:91676031EB73ED5EFF67FACD621D8301" box="[130,314,1378,1402]" country="China" county="Mangshi" municipality="Zhefang Township" name="Longchuan Jiang" pageId="27" pageNumber="520" stateProvince="Yunnan">Longchuan Jiang</location>
|
|||
|
|
at
|
|||
|
|
<location id="91676031EB73ED5EFEBCFACD612B8301" LSID="urn:lsid:plazi:treatment:1C1187FCEB72ED58FF9EFB736739844F:91676031EB73ED5EFEBCFACD612B8301" box="[345,524,1378,1402]" country="China" county="Mangshi" municipality="Zhefang Township" name="Nongkan village" pageId="27" pageNumber="520" stateProvince="Yunnan">Nongkan village</location>
|
|||
|
|
, 24.16534°/98.10266°,
|
|||
|
|
<quantity id="53409B0FEB73ED5EFCEDFACD606A8301" box="[776,845,1378,1402]" metricMagnitude="2" metricUnit="m" metricValue="7.34" pageId="27" pageNumber="520" unit="m" value="734.0">
|
|||
|
|
<elevation id="1F95D1D9EB73ED5EFCEDFACD606A8301" box="[776,845,1378,1402]" metricMagnitude="2" metricUnit="m" metricValue="7.34" pageId="27" pageNumber="520" unit="m" value="734.0">734 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="E006102AEB73ED5EFCBDFACD60CF8301" box="[856,1000,1378,1403]" pageId="27" pageNumber="520" value="2020-05-22">
|
|||
|
|
<collectingDate id="F042E9C2EB73ED5EFCBDFACD60CF8301" box="[856,1000,1378,1403]" pageId="27" pageNumber="520" value="2020-05-22">22 May 2020</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="394D533CEB73ED5EFC17FACC674F8301" box="[1010,1128,1378,1402]" pageId="27" pageNumber="520">H.B. Liang</collectorName>
|
|||
|
|
and
|
|||
|
|
<collectorName id="394D533CEB73ED5EFF9EFA2B639D83E0" box="[123,186,1412,1435]" pageId="27" pageNumber="520">Y. Xu</collectorName>
|
|||
|
|
collectors
|
|||
|
|
</materialsCitation>
|
|||
|
|
<materialsCitation id="24D03CB7EB73ED5EFED4FA2B619483E0" ID-GBIF-Occurrence="4905152353" box="[305,691,1411,1435]" collectingDate="2020-05-22" collectionCode="IOZ" collectorName="H. B. Liang & Y. Xu" country="China" county="Mangshi" elevation="734" location="Longchuan Jiang" municipality="Zhefang Township" pageId="27" pageNumber="520" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Yunnan">
|
|||
|
|
[
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFEDDFA2B628F83E0" box="[312,424,1411,1435]" count="2" pageId="27" pageNumber="520" type="male">two males</specimenCount>
|
|||
|
|
and
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFE3AFA2B614283E0" box="[479,613,1411,1435]" count="2" pageId="27" pageNumber="520" type="female">two females</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB73ED5EFD94FA2C618383E0" box="[625,676,1411,1435]" pageId="27" pageNumber="520">IOZ</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB73ED5EFD5BFA2B620283C0" ID-GBIF-Occurrence="4905152315" collectionCode="ZMUC" country="Bangladesh" location="Bengala" pageId="27" pageNumber="520" specimenCount="1" specimenCount-male="1">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFD5BFA2B60F083E0" bold="true" box="[702,983,1412,1435]" pageId="27" pageNumber="520">
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFD5BFA2B605083E0" box="[702,887,1412,1435]" name="Bangladesh" pageId="27" pageNumber="520">BANGLADESH</collectingCountry>
|
|||
|
|
/
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFC6DFA2B60F083E0" box="[904,983,1412,1435]" name="India" pageId="27" pageNumber="520">INDIA</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
: “
|
|||
|
|
<location id="91676031EB73ED5EFC15FA2C676C83E0" LSID="urn:lsid:plazi:treatment:1C1187FCEB72ED58FF9EFB736739844F:91676031EB73ED5EFC15FA2C676C83E0" box="[1008,1099,1411,1435]" country="Bangladesh" name="Bengala" pageId="27" pageNumber="520">Bengala</location>
|
|||
|
|
” ([
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFB8EFA2C639583C0" count="1" pageId="27" pageNumber="520" type="male">one male</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB73ED5EFF27FA0C623183C0" box="[194,278,1443,1467]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="27" pageNumber="520">ZMUC</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB73ED5EFED1FA0B67B583C0" ID-GBIF-Occurrence="4905152336" box="[308,1170,1443,1467]" collectionCode="MNFB" country="Bangladesh" location="BANGLADESH" pageId="27" pageNumber="520" specimenCount="1" specimenCount-female="1">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFED1FA0B61D383C0" bold="true" box="[308,756,1444,1467]" pageId="27" pageNumber="520">
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFED1FA0B62CA83C0" box="[308,493,1444,1467]" name="Bangladesh" pageId="27" pageNumber="520">BANGLADESH</collectingCountry>
|
|||
|
|
/
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFDE7FA0B616883C0" box="[514,591,1444,1467]" name="India" pageId="27" pageNumber="520">INDIA</collectingCountry>
|
|||
|
|
/
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFD81FA0B61D383C0" box="[612,756,1444,1467]" name="Myanmar" pageId="27" pageNumber="520">MYANMAR</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
: “India Or.” ([
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFC7AFA0C670683C0" box="[927,1057,1443,1467]" count="1" pageId="27" pageNumber="520" type="female">one female</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB73ED5EFBD4FA0C67A383C0" box="[1073,1156,1443,1467]" pageId="27" pageNumber="520">MNFB</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
<materialsCitation id="24D03CB7EB73ED5EFF9EFA6B62108380" ID-GBIF-Occurrence="4905152354" collectingDate="1918-10-14" collectionCode="NHMUK" collectorName="H. Stevens" country="India" elevation="240" location="Teesta River valley" municipality="Melli" pageId="27" pageNumber="520" specimenCount="1" specimenCount-male="1" stateProvince="Sikkim">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFF9EFA6B63ED83A0" bold="true" box="[123,202,1476,1499]" pageId="27" pageNumber="520">
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFF9EFA6B63ED83A0" box="[123,202,1476,1499]" name="India" pageId="27" pageNumber="520">INDIA</collectingCountry>
|
|||
|
|
</emphasis>
|
|||
|
|
:
|
|||
|
|
<collectingRegion id="567CF808EB73ED5EFF3FFA6C620C83A0" box="[218,299,1475,1499]" country="India" name="Sikkim" pageId="27" pageNumber="520">Sikkim</collectingRegion>
|
|||
|
|
:
|
|||
|
|
<collectingMunicipality id="7463AC90EB73ED5EFED2FA6C625683A0" box="[311,369,1475,1499]" pageId="27" pageNumber="520">Melli</collectingMunicipality>
|
|||
|
|
(
|
|||
|
|
<location id="91676031EB73ED5EFE64FA6C617183A0" LSID="urn:lsid:plazi:treatment:1C1187FCEB72ED58FF9EFB736739844F:91676031EB73ED5EFE64FA6C617183A0" box="[385,598,1475,1499]" country="India" municipality="Melli" name="Teesta River valley" pageId="27" pageNumber="520" stateProvince="Sikkim">Teesta River valley</location>
|
|||
|
|
,
|
|||
|
|
<quantity id="53409B0FEB73ED5EFD86FA6C618C83A0" box="[611,683,1475,1499]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="27" pageNumber="520" unit="m" value="240.0">
|
|||
|
|
<elevation id="1F95D1D9EB73ED5EFD86FA6C618C83A0" box="[611,683,1475,1499]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="27" pageNumber="520" unit="m" value="240.0">240 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<date id="E006102AEB73ED5EFD5CFA6C605483A0" box="[697,883,1475,1499]" pageId="27" pageNumber="520" value="1918-10-14">
|
|||
|
|
<collectingDate id="F042E9C2EB73ED5EFD5CFA6C605483A0" box="[697,883,1475,1499]" pageId="27" pageNumber="520" value="1918-10-14">14 October 1918</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
,
|
|||
|
|
<collectorName id="394D533CEB73ED5EFC64FA6B60D083A0" box="[897,1015,1475,1499]" pageId="27" pageNumber="520">H. Stevens</collectorName>
|
|||
|
|
collector [
|
|||
|
|
<specimenCount id="82BEFD63EB73ED5EFB89FA6B63958380" count="1" pageId="27" pageNumber="520" type="male">one male</specimenCount>
|
|||
|
|
;
|
|||
|
|
<collectionCode id="F2A9AE2FEB73ED5EFF58FA4C620E8380" box="[189,297,1507,1531]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="27" pageNumber="520" type="Museum">NHMUK</collectionCode>
|
|||
|
|
])
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="DCA26561EB73ED58FF4EF9AB6739844F" lastPageId="29" lastPageNumber="522" pageId="27" pageNumber="520" type="description">
|
|||
|
|
<paragraph id="940736EAEB73ED5EFF4EF9AB607080E7" blockId="27.[123,1174,219,1725]" pageId="27" pageNumber="520">
|
|||
|
|
This species ranges from the
|
|||
|
|
<typeStatus id="4B038848EB73ED5EFE0BF9AB613B8067" box="[494,540,1540,1564]" pageId="27" pageNumber="520">type</typeStatus>
|
|||
|
|
area, “Bengala” or “Bengal” (no specific locality), which includes present-day eastern
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFE52F98B62C98047" box="[439,494,1572,1596]" name="India" pageId="27" pageNumber="520">India</collectingCountry>
|
|||
|
|
and/or
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFDA0F98B61E38047" box="[581,708,1572,1596]" name="Bangladesh" pageId="27" pageNumber="520">Bangladesh</collectingCountry>
|
|||
|
|
, at least to westernmost Yunnan Province,
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFF9EF9EB63998027" box="[123,190,1604,1628]" name="China" pageId="27" pageNumber="520">China</collectingCountry>
|
|||
|
|
.
|
|||
|
|
<collectingMunicipality id="7463AC90EB73ED5EFF2CF9EB63D58027" box="[201,242,1604,1628]" pageId="27" pageNumber="520">The</collectingMunicipality>
|
|||
|
|
westernmost confirmed limit of the species is in
|
|||
|
|
<collectingRegion id="567CF808EB73ED5EFCF5F9EB607A8027" box="[784,861,1604,1628]" country="India" name="Sikkim" pageId="27" pageNumber="520">Sikkim</collectingRegion>
|
|||
|
|
and the easternmost limit in western Yunnan.
|
|||
|
|
<location id="91676031EB73ED5EFECAF9CB62858007" LSID="urn:lsid:plazi:treatment:1C1187FCEB72ED58FF9EFB736739844F:91676031EB73ED5EFECAF9CB62858007" box="[303,418,1636,1660]" country="India" municipality="The" name="Andrewes'" pageId="27" pageNumber="520" stateProvince="Sikkim">Andrewes’</location>
|
|||
|
|
records from northern
|
|||
|
|
<collectingCountry id="ECAF767AEB73ED5EFD72F9CB61E98007" box="[663,718,1636,1660]" name="India" pageId="27" pageNumber="520">India</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="F0294B1BEB73ED5EFD3EF9CB60A28007" author="ANDREWES, H. E." box="[731,901,1636,1660]" pageId="27" pageNumber="520" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes 1921</bibRefCitation>
|
|||
|
|
) may have been based on misidentified specimens of
|
|||
|
|
<taxonomicName id="53B84D69EB73ED5EFE46F92A611480E7" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[419,563,1668,1692]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="520" phylum="Arthropoda" rank="species" species="vespertilio">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFE46F92A611480E7" box="[419,563,1668,1692]" italics="true" pageId="27" pageNumber="520">C. vespertilio</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, which is described below.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="940736EAEB73ED58FF4EF90A602D8748" blockId="27.[123,1174,219,1725]" lastBlockId="29.[123,1174,219,565]" lastPageId="29" lastPageNumber="522" pageId="27" pageNumber="520">
|
|||
|
|
<emphasis id="A6CCEAF8EB73ED5EFF4EF90A629B80C7" bold="true" box="[171,444,1701,1724]" pageId="27" pageNumber="520">Geographical variation.</emphasis>
|
|||
|
|
We found some individual variation (
|
|||
|
|
<figureCitation id="0C832A6FEB73ED5EFCBCF90A60F880C6" box="[857,991,1701,1725]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="27" pageNumber="520">Figs.12A, C</figureCitation>
|
|||
|
|
) in development of the dark color pattern of the elytra within the sample of this species from Mangshi,
|
|||
|
|
<collectingRegion id="567CF808EB75ED58FBD8FF7467B68689" box="[1085,1169,219,242]" country="China" name="Yunnan" pageId="29" pageNumber="522">Yunnan</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry id="ECAF767AEB75ED58FF9EFF5463998768" box="[123,190,251,275]" name="China" pageId="29" pageNumber="522">China</collectingCountry>
|
|||
|
|
. Specimens from all other localities were similar to one or more of the specimens from that sample, so we recognized no particular geographic variation.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="C0C76662EB74ED59FF4EFA5B62808041" ID-DOI="http://doi.org/10.5281/zenodo.11512982" ID-Zenodo-Dep="11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="28" pageNumber="521" startId="28.[171,183,1524,1543]" targetBox="[120,1173,216,1504]" targetPageId="28" targetType="figure">
|
|||
|
|
<paragraph id="940736EAEB74ED59FF4EFA5B62808041" blockId="28.[123,1174,1523,1594]" pageId="28" pageNumber="521">
|
|||
|
|
<smallCapsWord id="92E1A036EB74ED59FF4EFA5B63CF807D" baselines="1538,1538" box="[171,232,1524,1543]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="28" pageNumber="521">FIGURE</smallCapsWord>
|
|||
|
|
12.
|
|||
|
|
<taxonomicName id="53B84D69EB74ED59FEEBFA5C6163807D" authority="(Wiedemann)" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[270,580,1523,1542]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="521" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB74ED59FEEBFA5C62E1807D" box="[270,454,1523,1542]" italics="true" pageId="28" pageNumber="521">Cyclosomus suturalis</emphasis>
|
|||
|
|
(Wiedemann)
|
|||
|
|
</taxonomicName>
|
|||
|
|
. A.
|
|||
|
|
<taxonomicName id="53B84D69EB74ED59FD83FA5C604B807D" authority="Wiedemann" authorityName="Wiedemann. A" authorityYear="1819" box="[614,876,1523,1542]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="521" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB74ED59FD83FA5C61D9807D" box="[614,766,1523,1542]" italics="true" pageId="28" pageNumber="521">Scolytus suturalis</emphasis>
|
|||
|
|
Wiedemann
|
|||
|
|
</taxonomicName>
|
|||
|
|
, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="940736EAEB75ED58FF4EFE936739844F" blockId="29.[123,1174,219,565]" pageId="29" pageNumber="522">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FF4EFE93617F8728" bold="true" box="[171,600,316,339]" pageId="29" pageNumber="522">Geographical relationships with other</emphasis>
|
|||
|
|
<taxonomicName id="53B84D69EB75ED58FDBAFE9360188728" box="[607,831,316,340]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="undetermined">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FDBAFE93601F8728" bold="true" box="[607,824,316,340]" pageId="29" pageNumber="522">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FDBAFE9361C4872F" bold="true" box="[607,739,316,340]" italics="true" pageId="29" pageNumber="522">Cyclosomus</emphasis>
|
|||
|
|
species
|
|||
|
|
</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicName>
|
|||
|
|
Because of the uncertainty with regard to where the exactly the
|
|||
|
|
<typeStatus id="4B038848EB75ED58FE28FEF362DC870F" box="[461,507,348,372]" pageId="29" pageNumber="522">type</typeStatus>
|
|||
|
|
specimen was collected in “Bengala”, the northwestern limits to the range of this species remain unclear.
|
|||
|
|
<taxonomicName id="53B84D69EB75ED58FDB2FED4607E87E8" box="[599,857,379,403]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="marginatus">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FDB2FED4607E87E8" box="[599,857,379,403]" italics="true" pageId="29" pageNumber="522">Cyclosomus marginatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is also known only from the similarly vague “
|
|||
|
|
<collectingCountry id="ECAF767AEB75ED58FED7FE34624C87C8" box="[306,363,411,435]" name="India" pageId="29" pageNumber="522">India</collectingCountry>
|
|||
|
|
Or.,” again without specific locality, so it is possible that these species co-occur or at least overlap in their geographical ranges. Whether or not the range of
|
|||
|
|
<taxonomicName id="53B84D69EB75ED58FC3EFE13677087AF" authorityName="Wiedemann. A" authorityYear="1819" box="[987,1111,444,468]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FC3EFE13677087AF" box="[987,1111,444,468]" italics="true" pageId="29" pageNumber="522">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
overlaps those of
|
|||
|
|
<taxonomicName id="53B84D69EB75ED58FEECFE7362AB878F" authorityName=", Chaudoir" authorityYear="1876" box="[265,396,476,500]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="flexuosus">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FEECFE7362AB878F" box="[265,396,476,500]" italics="true" pageId="29" pageNumber="522">C. flexuosus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and/or
|
|||
|
|
<taxonomicName id="53B84D69EB75ED58FE3AFE736148878F" authorityName="Kavanaugh & Cueva-Dabkoski" authorityYear="2023" box="[479,623,476,500]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="vespertilio">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FE3AFE736148878F" box="[479,623,476,500]" italics="true" pageId="29" pageNumber="522">C. vespertilio</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
cannot yet been confirmed. The range of
|
|||
|
|
<taxonomicName id="53B84D69EB75ED58FBD5FE7367B1878F" authorityName="Andrewes" authorityYear="1924" box="[1072,1174,476,500]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="inustis">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FBD5FE7367B1878F" box="[1072,1174,476,500]" italics="true" pageId="29" pageNumber="522">C. inustis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
approaches that of
|
|||
|
|
<taxonomicName id="53B84D69EB75ED58FEAFFE5262EF846F" authorityName="Wiedemann. A" authorityYear="1819" box="[330,456,508,532]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="522" phylum="Arthropoda" rank="species" species="suturalis">
|
|||
|
|
<emphasis id="A6CCEAF8EB75ED58FEAFFE5262EF846F" box="[330,456,508,532]" italics="true" pageId="29" pageNumber="522">C. suturalis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
at least in northern Indochina so there is at least potential for co-occurrence of these species, although the two have not yet been found near each other.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
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|||
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</document>
|